Changes in species abundance and richness, stand structure, and species responses to habitat characteristics including disturbance intensity, were investigated in 28 permanent sample plots (PSP) covering a total area of about 6.2 hectares of subtropical rainforests in north-east New South Wales, Australia. The disturbance that occurred over 36 years previously varied from unlogged (controls), through single-tree selection (light), moderate selection and repeated single-tree selection to intensive (heavy) logging. Multivariate and univariate approaches were used to investigate changes in various plant groups, including all vascular plants, regenerating species (trees and understorey species > 1.3 m in height and < 10 cm diameter at 1.3 m above the ground level (dbh)), juvenile trees (regenerating species capable of attaining at least 10 cm dbh), and trees ¡Ý 10 cm dbh. In addition, dynamic parameters (recruitment, growth and mortality) were estimated using hierarchical multilevel modelling and parameter estimates used to simulate behaviour of the subtropical rainforest tree species in unlogged and logged stands, both within and beyond the range of available data.Chronological post-disturbance responses and changes in species abundance and richness as well as stand structure for trees ¡Ý 10 cm dbh were investigated using 20 PSP in a bid to better understand rainforest post-disturbance regeneration. The results portrayed minor and gradual ecological changes in the undisturbed controls analogous with natural forest dynamics where the changes were not significant. In logged sites, initial gradual changes were followed with more rapid and significant changes.In both unlogged and logged stands, the changes proceeded through three clearly identifiable stages. The first two stages lasted between 5 and 15 years each. In the initial stage, the number of individuals decreased in species with high frequency mainly in the shade tolerant species, with little or no change in species richness. In the second stage, species abundance and richness changed due to localised species turnover, replacement and losses with very little recruitment, and the numbers of both speciesand individuals declined to minima. In the third stage, due to recruitment beyond the 10 cm dbh, the net loss of species and trees ¡Ý 10 cm dbh were halted and reversed, and species diversity and abundance began to return to pre-disturbance levels.Species abundance and diversity for both shade tolerant and intolerant species in less intensively logged sites (single-tree selection logging) have recovered to levels comparable with that observed in the controls, but more intensively logged sites have not recovered to similar levels. Increased logging intensity was associated with increased and decreased densities in the shade intolerant and ¨Ctolerant species respectively. It was evident that stand structure in logged sites had yet to recover to levels comparable with that of unlogged controls. It appears the restoration of floristic diversity to levels similar to that of intact primary forest takes considerably less time than structural recovery. Changes in regeneration patterns in a subtropical rainforest in north-east New South Wales were investigated for a 13-year period during the 3rd and 4th decades following repeated single-tree selection logging. Multivariate and univariate analyses results showed that there were no significant differences in floristic assemblages within and between censuses; however, two contrasting trends of changes in plant groups were detected. In trees ¡Ý 10 cm dbh, the stem density and species richness increased in shade tolerant group, and while stem density increased in the shade intolerant group, species richness decreased during the study period. Amongst smaller sized species including trees (< 10 cm dbh), a general decrease in species richness was observed along with significant changes in stem densities (P = 0.03) with the number of individuals in the shade tolerant species increasing, while that of both shade intolerant and vine species decreased. Excluding the vines and understorey species from the broader regenerating species group, revealed a decrease in species richness in juvenile canopy tree, and a significant change in densities (P = 0.004) with the number of individuals in shade tolerant increasing, while that of shade intolerant trees decreased. A comparison between the canopy trees ¡Ý 10 cm dbh and juvenile canopy trees group showed that these groups were tending towards similar floristic assemblages. These results suggest gradual replacement of shade intolerant by shade tolerant species as stands were tending toward later stages of recovery within the regeneration. The results also show that the inclusion of regenerating species in long-term studies is both complementary to the larger plant component and more revealing of both trends and changes.Species-specific estimates of shade tolerance and size structure at maturity derived from observations on 23 plots were used to intuitively group 277 vascular plant species into 3 main groups including full floristic (all together), shade tolerant and intolerant groups. The shade tolerant and intolerant groups were further grouped into smaller plants groups depending on the development stage and maximum size at maturity. These groups included regenerating species, juvenile trees, and trees ¡Ý 10 cm dbh. Multivariate analyses of these nominated groups revealed that floristic assemblages were significantly associated with environmental gradients based on simple site characteristics.At the landscape (larger) scale, the floristic assemblages were significantly different between low (200-400), mid (400-700) and high altitude (over 700 meters above sea level). In addition, abundance of shade tolerant species including Doryphora sassafras Endl, Orites excelsa R.Br and Caldcluvia paniculosa (F.Muell) Hoogland was positively correlated with the altitudinal gradient. At the smaller (local) scale, intensively logged sites where large gaps were created had recovered their species richness, but effects of past logging were evident in trees ¡Ý 10 cm dbh, especially trees greater than 50 cm dbh where species abundance in shade tolerant decreased significantly. Less intensively logged sites at mid altitude where a few stems were removed had recovered their species richness in respect to trees ¡Ý 10 cm dbh, but the small gaps created may have healed quickly for appreciable regeneration response to occur, as juvenile trees component was significantly different to that of the (unlogged) controls.The abundance of shade intolerant and juvenile shade tolerant tree species increased in concert with levels of disturbance and abundance of shade intolerant species such as Duboisia myoporoides R.Br, Acacia melanoxylon R.Br, Rubus moorei F.Muell and R. hilli F.Muell was positively correlated with the disturbance gradient. Increase in number of individuals of certain tree species was positively correlated with both disturbance intensity and topography (mid slope through lower slope to creek/gully), but negatively correlated with aspect (NE - NW). We concluded that logged sites are yet to recover their pre-logging stand structure and rainforest tree species were found in sites where favourable recruitment and growth conditions were the encountered. These results show that logged sites had not yet regenerated to the pre-logging stand structure. They also show that habitat characteristics including history of disturbance and topography can influence the floristic assemblages in the sub-tropical rainforests at both the small (local) and large (geographic) scales. This supports the environmental control model that states ¡°species are found at sites where they encounter favourable living conditions¡±. Site characteristics as surrogate for some of these favourable living conditions were identified as useful potential variables to investigate the rainforest dynamic parameters (growth, recruitment and mortality) in both logged and unlogged stands.A quantitative model was developed using over 3 decades of data to describe and simulate the dynamics including recruitment, growth and mortality in unlogged stands and others subjected to different silvicultural regimes in uneven-aged mixed-species subtropical rainforests of north-eastern New South Wales. Hierarchical multilevel regression analyses including Poisson, Binomial and multinomial logit regression were used to estimate the rainforest dynamic parameters based on the assumption that trees perform differently in space and time, thus there are variations at both the plot and tree by measurement levels.Variations at the tree level required the botanical identity of trees to species level, and then the species-specific size at maturity and shade tolerance were used to classify species into 5 groups, each consisting of species with similar ecological characteristics. These groups were labelled as emergent and shade tolerant main canopy, shade tolerant mid canopy, shade tolerant understoreys, moderate shade tolerant and persistent, and shade intolerant pioneer tree species. Significant variables at the plot level including site characteristics such as topography (elevation, slope and aspect), and past disturbance were used as explanatory variables in species group models. The final model is as a classical matrix management-oriented model with an ecological touch and maximum size-dependent parameters of ingrowth and outgrowth. The model provides a tool to simulate stand performance after logging and to assess silvicultural prescriptions before they are applied in these types of forests.The simulations indicate that full recovery following a logging intensity where 47% of the overstorey basal area is removed with a checkerboard of logged and unlogged patches (group selection) on a 120-year cycle could enable sustainable timber production without compromising the ecological integrity in these forests. Following single-tree selection (33%), recovery takes about 150 years, and more intensive harvesting practices where 50-78% overstorey basal area is removed may take 180-220 years to recover. Pre-harvest climber cutting coupled with poisoning of non-timber species followed by intensive logging of merchantable trees would allow logging on a 300-year cycle. Shorter logging cycles may lead to changes in the forest structure and floristic composition where the overall species density is low with higher density of shade intolerant species.

No description available.

Changes in species abundance and richness, stand structure, and species responses to habitat characteristics including disturbance intensity, were investigated in 28 permanent sample plots (PSP) covering a total area of about 6.2 hectares of subtropical rainforests in north-east New South Wales, Australia. The disturbance that occurred over 36 years previously varied from unlogged (controls), through single-tree selection (light), moderate selection and repeated single-tree selection to intensive (heavy) logging. Multivariate and univariate approaches were used to investigate changes in various plant groups, including all vascular plants, regenerating species (trees and understorey species > 1.3 m in height and < 10 cm diameter at 1.3 m above the ground level (dbh)), juvenile trees (regenerating species capable of attaining at least 10 cm dbh), and trees ¡Ý 10 cm dbh. In addition, dynamic parameters (recruitment, growth and mortality) were estimated using hierarchical multilevel modelling and parameter estimates used to simulate behaviour of the subtropical rainforest tree species in unlogged and logged stands, both within and beyond the range of available data.Chronological post-disturbance responses and changes in species abundance and richness as well as stand structure for trees ¡Ý 10 cm dbh were investigated using 20 PSP in a bid to better understand rainforest post-disturbance regeneration. The results portrayed minor and gradual ecological changes in the undisturbed controls analogous with natural forest dynamics where the changes were not significant. In logged sites, initial gradual changes were followed with more rapid and significant changes.In both unlogged and logged stands, the changes proceeded through three clearly identifiable stages. The first two stages lasted between 5 and 15 years each. In the initial stage, the number of individuals decreased in species with high frequency mainly in the shade tolerant species, with little or no change in species richness. In the second stage, species abundance and richness changed due to localised species turnover, replacement and losses with very little recruitment, and the numbers of both speciesand individuals declined to minima. In the third stage, due to recruitment beyond the 10 cm dbh, the net loss of species and trees ¡Ý 10 cm dbh were halted and reversed, and species diversity and abundance began to return to pre-disturbance levels.Species abundance and diversity for both shade tolerant and intolerant species in less intensively logged sites (single-tree selection logging) have recovered to levels comparable with that observed in the controls, but more intensively logged sites have not recovered to similar levels. Increased logging intensity was associated with increased and decreased densities in the shade intolerant and ¨Ctolerant species respectively. It was evident that stand structure in logged sites had yet to recover to levels comparable with that of unlogged controls. It appears the restoration of floristic diversity to levels similar to that of intact primary forest takes considerably less time than structural recovery. Changes in regeneration patterns in a subtropical rainforest in north-east New South Wales were investigated for a 13-year period during the 3rd and 4th decades following repeated single-tree selection logging. Multivariate and univariate analyses results showed that there were no significant differences in floristic assemblages within and between censuses; however, two contrasting trends of changes in plant groups were detected. In trees ¡Ý 10 cm dbh, the stem density and species richness increased in shade tolerant group, and while stem density increased in the shade intolerant group, species richness decreased during the study period. Amongst smaller sized species including trees (< 10 cm dbh), a general decrease in species richness was observed along with significant changes in stem densities (P = 0.03) with the number of individuals in the shade tolerant species increasing, while that of both shade intolerant and vine species decreased. Excluding the vines and understorey species from the broader regenerating species group, revealed a decrease in species richness in juvenile canopy tree, and a significant change in densities (P = 0.004) with the number of individuals in shade tolerant increasing, while that of shade intolerant trees decreased. A comparison between the canopy trees ¡Ý 10 cm dbh and juvenile canopy trees group showed that these groups were tending towards similar floristic assemblages. These results suggest gradual replacement of shade intolerant by shade tolerant species as stands were tending toward later stages of recovery within the regeneration. The results also show that the inclusion of regenerating species in long-term studies is both complementary to the larger plant component and more revealing of both trends and changes.Species-specific estimates of shade tolerance and size structure at maturity derived from observations on 23 plots were used to intuitively group 277 vascular plant species into 3 main groups including full floristic (all together), shade tolerant and intolerant groups. The shade tolerant and intolerant groups were further grouped into smaller plants groups depending on the development stage and maximum size at maturity. These groups included regenerating species, juvenile trees, and trees ¡Ý 10 cm dbh. Multivariate analyses of these nominated groups revealed that floristic assemblages were significantly associated with environmental gradients based on simple site characteristics.At the landscape (larger) scale, the floristic assemblages were significantly different between low (200-400), mid (400-700) and high altitude (over 700 meters above sea level). In addition, abundance of shade tolerant species including Doryphora sassafras Endl, Orites excelsa R.Br and Caldcluvia paniculosa (F.Muell) Hoogland was positively correlated with the altitudinal gradient. At the smaller (local) scale, intensively logged sites where large gaps were created had recovered their species richness, but effects of past logging were evident in trees ¡Ý 10 cm dbh, especially trees greater than 50 cm dbh where species abundance in shade tolerant decreased significantly. Less intensively logged sites at mid altitude where a few stems were removed had recovered their species richness in respect to trees ¡Ý 10 cm dbh, but the small gaps created may have healed quickly for appreciable regeneration response to occur, as juvenile trees component was significantly different to that of the (unlogged) controls.The abundance of shade intolerant and juvenile shade tolerant tree species increased in concert with levels of disturbance and abundance of shade intolerant species such as Duboisia myoporoides R.Br, Acacia melanoxylon R.Br, Rubus moorei F.Muell and R. hilli F.Muell was positively correlated with the disturbance gradient. Increase in number of individuals of certain tree species was positively correlated with both disturbance intensity and topography (mid slope through lower slope to creek/gully), but negatively correlated with aspect (NE - NW). We concluded that logged sites are yet to recover their pre-logging stand structure and rainforest tree species were found in sites where favourable recruitment and growth conditions were the encountered. These results show that logged sites had not yet regenerated to the pre-logging stand structure. They also show that habitat characteristics including history of disturbance and topography can influence the floristic assemblages in the sub-tropical rainforests at both the small (local) and large (geographic) scales. This supports the environmental control model that states ¡°species are found at sites where they encounter favourable living conditions¡±. Site characteristics as surrogate for some of these favourable living conditions were identified as useful potential variables to investigate the rainforest dynamic parameters (growth, recruitment and mortality) in both logged and unlogged stands.A quantitative model was developed using over 3 decades of data to describe and simulate the dynamics including recruitment, growth and mortality in unlogged stands and others subjected to different silvicultural regimes in uneven-aged mixed-species subtropical rainforests of north-eastern New South Wales. Hierarchical multilevel regression analyses including Poisson, Binomial and multinomial logit regression were used to estimate the rainforest dynamic parameters based on the assumption that trees perform differently in space and time, thus there are variations at both the plot and tree by measurement levels.Variations at the tree level required the botanical identity of trees to species level, and then the species-specific size at maturity and shade tolerance were used to classify species into 5 groups, each consisting of species with similar ecological characteristics. These groups were labelled as emergent and shade tolerant main canopy, shade tolerant mid canopy, shade tolerant understoreys, moderate shade tolerant and persistent, and shade intolerant pioneer tree species. Significant variables at the plot level including site characteristics such as topography (elevation, slope and aspect), and past disturbance were used as explanatory variables in species group models. The final model is as a classical matrix management-oriented model with an ecological touch and maximum size-dependent parameters of ingrowth and outgrowth. The model provides a tool to simulate stand performance after logging and to assess silvicultural prescriptions before they are applied in these types of forests.The simulations indicate that full recovery following a logging intensity where 47% of the overstorey basal area is removed with a checkerboard of logged and unlogged patches (group selection) on a 120-year cycle could enable sustainable timber production without compromising the ecological integrity in these forests. Following single-tree selection (33%), recovery takes about 150 years, and more intensive harvesting practices where 50-78% overstorey basal area is removed may take 180-220 years to recover. Pre-harvest climber cutting coupled with poisoning of non-timber species followed by intensive logging of merchantable trees would allow logging on a 300-year cycle. Shorter logging cycles may lead to changes in the forest structure and floristic composition where the overall species density is low with higher density of shade intolerant species.

京都大学 / 0048 / 新制・課程博士 / 博士(人間・環境学) / 甲第19070号 / 人博第723号 / 新制||人||173(附属図書館) / 26||人博||723(吉田南総合図書館) / 32021 / 京都大学大学院人間・環境学研究科相関環境学専攻 / (主査)教授 市岡 孝朗, 教授 加藤 眞, 教授 瀬戸口 浩彰 / 学位規則第4条第1項該当 / Doctor of Human and Environmental Studies / Kyoto University / DGAM

No description available.

Tropical forests, which provide important ecosystem functions and services, are increasingly threatened by anthropogenic pressures. This has resulted in an urgent need to understand tree species diversity of those forests. Where knowledge of that diversity is largely from the botanical surveys and local ecological studies, data must inevitably be up-scaled from point observations to the landscape and regional level if a holistic perspective is required. This thesis explores aspects of the spatio-temporal heterogeneity of canopy reflectance patterns over the forests of French Guiana, in order to assess whether this information could help defining an ecologically consistent forest typology. To gain insight into both the spatial and temporal heterogeneity of French Guiana’s forests, instrumental artefacts affecting the satellite data first had to be addressed. Data used in this study represent the spectral response of forest canopies, and the way in which such data are captured makes them susceptible to the ‘bi-directional reflectance distribution function’ (BRDF). BRDF indicates that objects do not reflect light in equal proportions in all directions (isotropically). Thus, forest canopies will reflect light anisotropically depending on factors including canopy roughness, leaf optical properties and inclination, and the position of the sun relative to the sensor. The second chapter of this thesis examines how BRDF affects the canopy reflectance of forests in French Guiana, and how not correcting for BRDF affects spectral classifications of those forests. When monthly reflectance data corrected for the artefact are examined, these suggest seasonally-occurring changes in forest structure or spectral properties of French Guiana’s forests. The third chapter of this thesis thus examines temporal effects of BRDF, and used cross-regional comparisons and plot-level radiative transfer modelling to seek to understand the drivers of the monthly variation of the forests’ canopy reflectance. For the latter, the Discrete Anisotropic Radiative Transfer (DART) model was used along with aerial laser scanning (ALS) observations over different forest structures, indicating that the observed variation in reflectance (and derivatives known as vegetation indices) could not be explained by monthly variations in solar direction. At the regional scale, it was also demonstrated that forests in the Guiana Shield possess temporal variation distinct from forests in central Africa or northern Borneo, forests also lying just above the Equator. Had the observed temporal variation in vegetation indices been the result of BRDF, it would have been expected that the forests in the three zones would have similar patterns of variation, which they did not. Central African forests appear to have their greening synchronized with rainfall, whereas forests in the Guianas appear synchronized with the availability of solar radiation. Further analysis of the vegetation index time-series of observations also indicated that different types of forests in French Guiana possess distinct patterns of temporal variation, suggesting that tropical forest types can be discriminated on the basis of their respective “temporal signatures.” That was exploited in the fourth chapter of the thesis, which maps forests in French Guiana based on their combined spatio-temporal canopy reflectance patterns and by so doing presents a novel way of addressing forest typology, based on ecologically meaningful information. The thesis presented demonstrates that it is possible to adequately address remote sensing data artefacts to examine patterns of spatial and temporal variation in tropical forests. It has shown that phenological patterns of tropical rainforests can be deduced from remote sensing data, and that forest types can be mapped based on spatio-temporal canopy reflectance patterns. It is thus an important contribution to understand the ecology of tropical forests in French Guiana and to improve the toolbox of scientists dealing with the identification of spatio-temporal patterns observable in forests at the landscape level.

Among the most compelling and least well-understood tree species growing in the temperate forests of southern South America is Fitzroya cupressoides, a high biomass species and the second longest-lived tree species in the world. This thesis quantified the main components of the carbon cycle in Fitzroya forests (i.e. net primary productivity (NPP) and soil respiration) and evaluated the environmental variables that are most related to them. The study was focused on medium-age and old-growth forests growing in the Coastal Range (Alerce Costero National Park, AC) and the Andean Cordillera (Alerce Andino National Park, AA) of southern Chile, respectively. The specific objectives of this thesis were to: 1) assess the forest structure, species composition and characterise the environmental conditions of these forests; 2) assess biomass, aboveground NPP, carbon allocation and mean wood residence time in these forests; 3) assess soil respiration and relate it to soil environmental conditions. Additionally, to use a mass balance approach to estimate fine root productivity; 4) estimate total NPP using biometric and indirect estimates of productivity; 5) evaluate the climatic factors mainly related to Fitzroya stem radial change on an intra-annual basis; and 6) evaluate changes in Fitzroya's tree growth and carbon isotopes during recent decades, and determine which environmental factors are more related to them. The last two objectives focus on Fitzroya as the dominant species and the subject of this study. Two 0.6 ha plots were installed within each national park; NPP was estimated for a year and soil respiration and high resolution stem growth measurements were monitored over almost two years. Aboveground biomass estimates for the Andean site are among the most massive reported in the world and carbon fluxes in Fitzroya forests are among the lowest reported for temperate wet forests worldwide. The longevity as well as the particularly rainy and nutrient poor soil conditions where these ecosystems grow may influence their exceptionally slow carbon dynamics. Differences in carbon fluxes between sites seem most probably driven by different environmental conditions rather than by developmental stage. Moreover, carbon fluxes were more sensitive to interannual climate variability in AC than AA. Warmer and drier summer conditions, likely to become more common under future climate change, more significantly affected stem growth and soil respiration in the Coastal Range than in the Andes. Regarding long-term changes, tree growth has been decreasing in the coastal site in the last 40 years and increasing in the Andes since the 1900s. These trends have been accompanied by an increase in intrinsic water use efficiency which is likely caused by rises in CO₂ and changes in climate conditions in both sites. Although Fitzroya grows in particularly wet and cool areas, projected drier and warmer conditions may have a negative effect on Fitzroya stem growth and carbon sequestration in both study sites. This effect would be more critical in the Coastal Range though, because of its more Mediterranean climate influence and more restrictive soil conditions in this area. Adequate resources are needed for the monitoring and conservation of these slow growth and massive forests especially in the Coastal Range, in order to avoid ongoing illegal cuttings and threatening forest fires.

Este estudo objetivou estudar os carrapatos e a infecção por patógenos, e sua correlação com os vertebrados desta região entre 10/2007 e 09/2008 foram realizadas mensalmente doze visitas a uma área de floresta ombrófila densa da Amazônica na Estação Ecológica de Samuel. Foram coletados, 60 aves, 36 mamíferos, 14 répteis e 11 anfíbios. Em vida livre foram realizadas 80 coletas de carrapatos em um total de 39.600m em cerca de 2.640min (44h) e coletados 265 carrapatos adultos de quatro espécies, sendo elas: A. scalpturatum (81), A. latepunctatum (84), A. oblongogutattum (18), A. naponense (69) e 597 ninfas de Amblyomma sp. Dos 104 adultos (11,8% do total de carrapatos) foram testados para Anaplasmataceae, obtive-se 27 (25,9%) positivos e mais 36 adultos (total de 140) e 11 carrapatos foram positivos (1) A. scalpturatum, (4) A. latepunctatum, (3) A. naponense. É notável o impacto da hidrelétrica sobre a fauna de pequenos mamíferos. / This study was aimed to analyze the ticks and the pathogenic infections, and their correlation with the vertebrates of such a region from October 2007 to September 2008, 12 surveys were monthly carried out at the Samuel Ecological Station in an Amazonian dense ombrophilous forest. They were collected from 60 birds, 36 mammals, 14 reptiles, and 11 amphibians. Eighty(80) collections of free-living ticks were carried out within an area of about 39.000 meters, during 2.640 minutes (44 hours), and 265 adult ticks or four species were picked up as following: 81 A.scalpturatum, 84 A. latepunctatum, 18 A. oblongogutattum, 69 A. naponense , and 597 Amblyomma sp. nymphs. 104 the adults, which accounted for 11.8% of total of ticks collected, were tested for Anaplasmataceae, and 27 (25.9%) were positive. In the Rickettsia spp. sample, additionally to the above mentioned 104 ticks tested, 36 adult ticks (out of a total of 140) were positive. It is noteworthy the impact caused by the hydropower plant on the small mammal fauna.

En 1988, après deux décennies de régime militaire, le Brésil rédige une nouvelle constitution afin de bâtir une nation fondée sur la démocratie, le multiculturalisme et l’intégration. C’est dans ce cadre que (re)naissent les quilombos, communautés afro-descendantes auxquelles est accordée, en guise de réparation historique, la propriété collective de leurs terres. Sont ainsi démarqués des territoires différenciés pour des populations spécifiques au nom de la diversité. Mais une telle mesure peut-elle suffire à assurer la durabilité de ces communautés noires rurales ? Pour répondre à cette question, nous recourons à trois études de cas, trois communautés quilombolas (Amazonie, Etat du Pará pour deux d’entre elles et vallée du Ribeira, Etat de São Paulo) dont nous croisons les caractéristiques et les trajectoires, les structures et les stratégies. Il s’agit, de cette façon, d’observer les mécanismes de leur durabilité, à partir d’un double point de vue. Le point de vue social d’abord, que nous évaluons en nous appuyant sur les travaux d’Elinor Ostrom et sa théorie des biens communs. Mais aussi un point de vue territorial qui, nous le supposons, permet d’enrichir l’observation des systèmes sociaux et de leurs mécanisme d’adaptation et de démontrer les enjeux territoriaux de la durabilité elle-même. Finalement, l’étude de nos trois cas permet de mettre en évidence que la « quilombolisation » des communautés semble remplir partiellement ses objectifs, en assurant des garanties territoriales et sociales certes, mais en rencontrant quelques difficultés à les inscrire pleinement dans le temps long. / In 1988, after two decades of military regime, Brazil writes a new constitution in order to build a nation founded on democracy, multiculturalism and integration. In this framework, Quilombos are reborn. They are afro-descendant communities to whom is given, as a historical redemption, the collective tenure of their lands. Differentiated territories are tenured for specific populations on behalf of diversity. But can such a measure be sufficient to ensure the sustainability of these black rural communities? To answer this question, we focus on three case studies, three Quilombola communities (two in the Amazon, State of Pará and one in the Ribeira Valley, State of São Paulo) and we cross their features and trajectories, their structures and strategies. In this way, the aim is to observe the mechanisms of their sustainability, from a double point of view. A social point of view, first, that we assess by relying on Elinor Ostrom’s studies and her theory of Commons. But a territorial point of view too, that, we assume, completes the observation of social systems and their adaptation mechanisms and demonstrate the territorial stakes of the sustainability. Finally, the three case studies highlight that the “quilombolization” of the communities seems to partially fulfill its objectives, ensuring territorial and social guarantees, but still having some difficulties to last. / Em 1988, após duas décadas de governo militar, o Brasil escreve uma nova constituição para construir uma nação fundada sobre a democracia, o multiculturalismo e a integração. É nesse marco que (re)nascem os quilombos, comunidades afrodescendentes as quais é concedida, como reparação histórica, a propriedade coletiva de suas terras. Assim são demarcados territórios diferenciados par populações específicas em nome da diversidade. Mas, será que tal medida poda ser suficiente para garantir a sustentabilidade dessas comunidades negras rurais? Para responder a esse pergunta, mobilizamos três estudos de caso, três comunidades quilombolas (da Amazônia, Estado do Pará e do vale do Ribeira, Estado de São Paulo) das quais cruzamos as características e as trajetórias, as estruturas e as estratégias. Trata-se, dessa forma, de observar os mecanismos de sua sustentabilidade, a partir de um duplo ponto de vista. O ponto de vista social primeiro, que avaliamos nos apoiando nos trabalhos de Elinor Ostrom e sua teoria dos bens comuns. Mas também um ponto de vista territorial que, assumimos, permite enriquecer a observação dos sistemas sociais e seus mecanismos de adaptação e de demonstrar os desafios territoriais da própria sustentabilidade. Finalmente, o estudo de nossos três casos permite evidenciar que a “quilombolização” das comunidades parece cumprir parcialmente seus objetivos, assegurando garantias territoriais e sociais, mas encontrando algumas dificuldades para inseri-las plenamente no tempo longo.

A tese versa sobre a detecção e avaliação de padrões de coexistência de espécies arbóreas, buscando esclarecer as relações entre a diversidade fito-estrutural e os fatores condicionantes em escalas local e regional em uma zona de contato entre duas regiões fitoecológicas (Florestas Ombrófilas Mista e Densa). O trabalho está dividido em duas abordagens; a primeira trata de um inventário fitossociológico de comunidades definidas como “sítios”, onde o grau de singularidade local é avaliado através da identidade das espécies com maior valor de importância (VI) estrutural e de suas possíveis conexões fitogeográficas, bem como através de indicadores de diversidade alfa e beta. Na segunda abordagem, comunidades são definidas estatisticamente (Multivariada, Análise do Valor Indicador das Espécies) com base na composição e abundância de espécies (área basal dos troncos), testando-se a hipótese de que comunidades distintas devem diferir pelo menos na diversidade específica, fertilidade do solo e ou produtividade primária (produção de serrapilheira PS). O estudo foi realizado no Centro de Pesquisas e Conservação da Natureza Pró-Mata, localizado na borda sudeste do Planalto Meridional Brasileiro (29o30’S; 50o11’W). Cinco sítios de 100m x 100m (dois sítios no planalto, dois na encosta e um na borda planalto-encosta) foram selecionados, onde em cada sítio as árvores (DAP>10 cm) foram medidas e determinadas em vinte e uma unidades amostrais de 100m2 (regularmente distribuídas). Sessenta e seis espécies foram levantadas, no conjunto Cabralea canjerana e Myrcia retorta tiveram os maiores valores de importância. A primeira caracteriza a encosta e a segunda o planalto, sendo ambas co-dominantes no sítio da borda do planalto-encosta. Riqueza e diversidade foram maiores nessa borda, menor no planalto e intermediária na encosta. A beta-diversidade apresenta-se maior com medidas de abundância do que apenas com a composição de espécie, em função da diferenciação das dominantes, caracterizando comunidades discretas, indicando ao mesmo tempo um contínuo gradiente florístico. As análises de ordenação e agrupamento foram utilizadas com dois tamanhos de unidade amostral (100 m2 e 500 m2), das quais somente a última apresentou padrões interpretáveis, similares aos obtidos fitossociologicamente nos sítios, onde dois principais tipos de vegetação apresentam afinidades fitogeográficas e riqueza de espécies peculiares. Os subgrupos refletiram processos histórico-ecológicos mais recentes, como a perturbação antrópica e condições locais especiais. A determinação de espécies com valores indicadores significativos é útil para a diferenciação ecológica dos tipos de comunidades. Apenas uma comunidade diferiu significativamente das demais na fertilidade do solo, devido à condição especial topográfica e litológica. A produção de serrapilheira destas florestas foi similar aos valores médios das florestas brasileiras tropicais e subtropicais. Conclui-se que, na zona de transição estudada, tende a ter um ecótono da vegetação associado à transição geomorfológica planalto-encosta. Algumas considerações sobre o alcance metodológico de detecção de biodiversidade e das perspectivas de pesquisas integradas sobre os processos ecossistêmicos são apresentadas. / This thesis searches to detect patterns of the arboreal species coexistence on a transition zone of two phyto-ecological regions, Mixed and Dense Rainforests, contributing to the knowledge of ecological processes on the local to regional scale. The work is divided in two approaches; the first is a phytosociological inventories aiming to answer if in this zone exists a set of local communities with similar floristic-structural patterns, or not, and what are their phytogeographical affinities; the second tests the hypothesis that the communities types statistically defined have indicator species and differs in specific diversity attributes, soil fertility and the primary productivity (litterfall). The study was carried out at CPCN Pró-Mata, a research center located on the southeastern border of the Meridional Brazilian Plateau (29o30’S; 50o11’W). Five sites of 100 x 100m (two plateau sites, two slope sites, and one plateau-slope site) were selected; the trees (DBH>10cm) were measured and determined and twenty-one sampling units of 100m2 (regularly distributed), in each site. Sixty-six species were surveyed. Cabralea canjerana and Myrcia retorta had the highest importance value indexes. The first species characterizes the slope; the second species characterizes the plateau; and both are co-dominant at the slope-plateau site. Richness and diversity (Shannon index) were higher at slope-plateau transition, lower at the plateau and intermediate at the slopes. The Beta diversity shows that the sharpness between slope and plateau communities is more associated to species abundance than to species composition only. The Ordination and Cluster analyses were used with two sized sampling units (100 m2 and 500m2), which only the latter showed interpretable patterns, similar to those phytosociological results, where emerges the two main vegetation types indicated the broader regional floristic affinities and its influence on species richness. The sub-groups reflected more recent local historical-ecological processes, mainly identified as anthropic disturbance regime and special local conditions. The Indicator Species Analysis was very responsive to the present objective helping to define the best indicative species and ecological context. Soils differ significantly only for one group, differentiated by special topographic condition and lithologic origin. The annual litterfall of these forests were equivalent among sites and they are similar to register for average the tropical and subtropical Brazilian forests. Conclude that in this transition zone trends to have a vegetation ecotone is associated to the geomorphologic transition slope-plateau. Then, communities types statistically defined reflect hierarchically phytoecological regional affinities and particular ecological conditions. Some considerations about methodological biodiversity detection integrated to researches about ecosystem process are shown.