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The timing, duration, and pressure-temperature (P-T) conditions of metamorphism provide a direct record of the physical and chemical evolution of the crust and inform our knowledge and understanding of plate tectonics. The characteristic timescales and length-scales of metamorphism vary by orders of magnitude, depending on the driving tectonic process. Two fundamental problems with the retrieval of this information from the metamorphic rock record are insufficient temporal resolution and processes that overprint or obscure the full record of metamorphism. Understanding what processes are recorded, and why they are recorded, is critical for accurate models of tectonics. This dissertation examines these processes in the metamorphic rock record in two settings: the central Appalachian orogen and the Western Alps fossil subduction zone. Chapters 2 and 3 focus on poly-metamorphic migmatites from the Smith River Allochthon (SRA) in the central Appalachians. A combination of petrography, thermodynamic modeling, and geochemistry is used to document and quantify the metamorphic evolution of the SRA and determine the petrologic processes that control metamorphic re-equilibration in high-temperature metamorphic systems. Chapter 2 presents new constraints for Silurian high-temperature (~750℃, 0.5 GPa) contact metamorphism in response to mafic magmatism and a cryptic Alleghanian metamorphism (~600℃, 0.8 GPa). A combination of extensive and highly variable melt loss followed by H2O-flux melting during contact metamorphism is shown to produce a range of modified bulk rock compositions and domains with variable fertilities for metamorphic re-equilibration during the Alleghanian. In chapter 3, monazite, allanite, and zircon laser ablation split-stream petrochronology are used to constrain the timing of poly-metamorphism and develop a tectonic model for the SRA. The SRA preserves evidence for at least three orogenic events, each with a relatively short duration (< 10 Myr.), likely due to repeated magmatic heating. The full record of this punctuated heating is obscured by dissolution-reprecipitation reactions that variably recrystallize monazite and decouple trace element chemistry from isotopic age and significantly restrict equilibrium length-scales. Chapters 4 and 5 examine the dynamic interplay between transient fluid flow, episodic metamorphism, and deformation in subduction zones. In chapter 4, diffusional speedometry is applied to eclogite breccias from the Monviso ophiolite to quantify the periodicity of transient deformation and metamorphism at eclogite facies P-T conditions. The maximum timescale for repeated fracturing is constrained to ~1 Myr., likely caused by cyclic variations in fluid pressure and strain rate (not necessarily seismicity). While difficult to preserve and detect in the rock record, this periodic metamorphism may play an important role in detachment and exhumation processes in subduction zones worldwide. Finally, in chapter 4 a combination of thermodynamic modeling and Sm-Nd garnet geochronology are used to construct a model for subduction and exhumation of the Voltri ophiolite. Garnet growth occurs rapidly and close to peak P-T conditions (~520℃, 2.4 GPa) across the ophiolite, with large (>10 km2) areas preserving near-identical ages, suggesting that the Voltri ophiolite was exhumed as several large coherent units, aided by the presence of buoyant serpentinites. / Doctor of Philosophy / Metamorphism provides a direct record of the physical and chemical evolution of Earth's crust and informs our knowledge and understanding of how plate tectonics works on Earth. Differences in the physical conditions (e.g. pressure, temperature) and timescales of metamorphism can provide clues for the operation of unique tectonic processes, such as the intrusion and cooling of magma deep underground or the collision of two tectonic plates and formation of a mountain range. The key is to correctly "read" the metamorphic rock record. One inherent difficulty in reading and interpreting metamorphic rocks is that few current methods are able to resolve very short timescale events (much less that 1 million years (Myr.) in duration), such as earthquakes, in the rock record. Moreover, metamorphic rocks experience numerous distinct 'events', which partly overprint one another and produce a complicated and near impossible puzzle for geologists to unravel. Solving this puzzle is critical to fully understand how plate tectonics works on Earth. This dissertation addresses these problems and examines metamorphism in two locations: the core of the ancient supercontinent Pangea (central Appalachians) and a fossil subduction zone (the Western Alps). Chapters 2 and 3 focus on the central Appalachians. Chemical and textural analysis of metamorphic rocks are used to understand the major heat sources that operated in the crust during the formation of the Appalachians and determine the processes that control metamorphic re-crystallization at extremely high temperatures. Chapter 2 presents new constraints for high-temperature (~750℃) metamorphism in response to magmatic heating and provides evidence for a younger metamorphic event that is cryptically recorded. A combination of compositional changes caused by earlier high-temperature metamorphism and the later addition of water along reactive grain boundaries are shown to be important factors in the cryptic record of the younger metamorphic event. In chapter 3, U-Pb geochronology is used to the determine the timing of metamorphism and construct a tectonic model for the central Appalachians, which preserves evidence for at least three tectonic events over ~200 Myr, but with each occurring over a relatively short duration (< 10 Myr.). These events are interpreted to represent repeated magmatic heating 'pulses' during the formation of Pangea. However, the full record of this punctuated heating is partly obscured by subsequent fluid alteration. Chapters 4 and 5 examine the dynamic interplay between transient fluid flow, earthquakes, and metamorphism deep in subduction zones. In chapter 4, fracture sets within metamorphic garnet crystals from the French Alps (Monviso) are used to determine the timescale of repeated fracturing and recrystallization during subduction. The fracture timescales are estimated to be much less than 1 Myr. and are interpreted to record repeated fluid "pulses" and possibly deep earthquakes. While difficult to preserve and detect in the rock record, this process may play an important role in bringing metamorphic rocks back from deep in subduction zones to Earth's surface. In chapter 4, a combination of mineral chemistry and geochronology are used to construct a tectonic model for the subduction and exhumation of a portion of the Italian Alps (Voltri). Metamorphic reactions occur synchronously and immediately before exhumation across a wide area (> 10 km2). This suggest that large (> 10 km2) pieces of oceanic crust can metamorphose, detach, and exhume deep in subduction zones.

La communication entre les différentes organelles se fait par l’intermédiaire du trafic vésiculaire, un processus qui nécessite un remodelage continu des membranes. Les vésicules fortement courbées bourgeonnent d'un compartiment donneur et fusionnent avec un compartiment accepteur. Les protéines impliquées dans le bourgeonnement et fusion des vésicules ont été largement étudiées. Récemment, la découverte de détecteurs de courbure membranaire a révélé que le trafic membranaire pourrait être régulé à un niveau supplémentaire, par la détection de la forme de la membrane. Le premier détecteur de courbure membranaire identifié était le motif ALPS (Amphipathic Lipid Packing Sensor), qui a été trouvé dans un certain nombre de protéines de la voie sécrétoire précoce et l'enveloppe nucléaire. La protéine d’arrimage GMAP-210 localisé au niveau du cis-Golgi, est composée d’une longue superhélice (coiled-coil) et d’un motif ALPS à l'extrême N-terminale. Il a été démontré in vitro, que ce motif se replie et forme une hélice amphipathique capable de se fixer sur des petits liposomes. Toutefois, l'identité des vésicules, reconnues par ce détecteur de courbure dans la cellule, reste inconnue. α-Synucléine est une autre protéine qui se lie préférentiellement à des membranes très courbées. Cette protéine localisée sur les vésicules synaptiques, est impliquée dans la régulation du taux de vésicules au niveau des terminaisons nerveuses pré-synaptiques. Connue pour son rôle central dans le développement de la maladie de Parkinson, α-synucléine contient une région non structurée en solution, mais qui forme une hélice amphipathique au contact de petits liposomes in vitro. Les hélices amphipathiques formées par le motif ALPS et α-synucléine sont très différentes aussi bien sur le plan chimique que sur le plan conformationel. Le motif ALPS possède une face hydrophobe bien développée, mais un coté polair pauvre avec très peu de résidus chargés. α-Synucléine, en revanche, a un côté hydrophobe modéré, et une face polaire zwitterionique riche en résidus chargés. L'objectif principal du projet était de comparer les propriétés de liaison aux membranaires in vivo et in vitro de ces deux hélices amphipathiques de structure opposée. L’expression de ces deux sondes chez la levure, favorise l'accumulation de structures vésiculaires de propriétés différentes. L'extrémité N-terminale de la protéine GMAP-210 contenant son motif ALPS (GMAPN) co-localisé spécifiquement avec des marqueurs de la voie sécrétoire précoce, alors une sonde contenant une portion de l’hélice amphipathique d’α-synucléine co-localise avec des marqueurs endocytiques et post-Golgiens. La mutagenèse du motif ALPS et l'inversion de la séquence de ALPS dans GMAPN confirment que ce détecteur de courbure membranaire se fixe spécifiquement aux vésicules via des interactions directes protéines-lipides, plutôt que les interactions protéines-protéines. Notre analyse a montré que ces détecteurs de courbure mammifères, exprimés dans la levure préservent leur capacité à cibler des vésicules spécifiques, vésicules de la voie sécrétoire précoce pour les motifs ALPS, et vésicules d’endocytose/post-Golgi pour α-synucléine. La composition membranaire de ces vésicules correspond à la composition des liposomes fixés par le motif ALPS et α-synucléine in vitro. Les propriétés biochimiques opposées du motif ALPS et α-synucléine, sont parfaitement adaptés à chacun de ces deux environnements membranaires dans la cellule. Le programme HeliQuest est conçu pour identifier des hélices amphipathiques capables de se lier sur les membranes, y compris les motifs ALPS. Un nouveau module conçu pour identifier les hélices amphipathiques avec des propriétés similaires à α-synucléine a été récemment élaboré. Les recherches effectuées dans les bases de données de protéines de levure et humaines ont permis d’identifier des hélices amphipathiques candidats qui ont des propriétés similaires à α-synucléine, dans de nombreuses protéines. Nous avons préparé un ensemble de sondes, dans lequel ces hélices sont insérées à la fin de la superhélice de GMAPN. Une première étude de leur co-localisation dans les cellules de levure avec un ensemble de marqueurs démontre une localisation spécifique, ce qui suggère que ces hélices peuvent avoir la capacité de cibler des membranes de manière spécifique. D'autres travaux seraient nécessaires pour confirmer ou pas si ces hélices amphipathiques font partie d'une nouvelle classe de détecteurs de courbure ayant les mêmes propriétés que α-synucléine. / Communication between membrane-bound organelles is mediated by vesicular trafficking, a process which requires continual membrane remodeling. Highly curved vesicles bud from a donor compartment through functioning of different coat protein complexes, and fuse with an acceptor compartment thanks to proteins of the membrane fusion machinery. The proteins involved in vesicle budding and fusion have been extensively studied. Recently, the discovery of membrane curvature sensors revealed that membrane trafficking could be regulated at an additional level, through detection of the shape of a membrane. The first membrane curvature sensor identified was the ALPS (Amphipathic Lipid Packing Sensor) motif, which has been found in a number of proteins that function in the early secretory pathway and nuclear envelope. One example is GMAP-210, a long coiled-coil tether localizing to cis-Golgi membranes, which has an ALPS motif at its extreme N-terminus. This ALPS motif was found to fold into an amphipathic helix and bind to small liposomes in vitro. However, the identity of the vesicles that this curvature sensor binds to in cells is not known. Another protein - α-synuclein - has also been reported to bind preferentially to highly curved membranes. This neuronal protein localizes to synaptic vesicles and is involved in maintaining the reserve pool of vesicles in pre-synaptic nerve terminals. α-Synuclein, known for its central role in the development of Parkinson’s disease, contains a region that is unstructured in solution, but forms an amphipathic helix upon binding to small liposomes in vitro. The chemistry and geometry of the amphipathic helices formed by ALPS motifs and α-synuclein are very different. The ALPS motif has a well-developed hydrophobic face but a poor polar side with few charged residues. α-Synuclein, in contrast, has a restrained hydrophobic side, and a zwitterionic polar face rich in charged residues. The main goal of the project was to compare the in vivo and in vitro membrane binding properties of these two amphipathic helices of opposite structure. When expressed in yeast cells, these two curvature sensors promoted the accumulation of vesicular structures possessing different characteristics. The N-terminus of GMAP-210 containing its ALPS motif (GMAPN) co-localized specifically with early secretory pathway markers, whereas a probe containing a portion of the amphipathic membrane-binding helix of α-synuclein co-localized with endocytic and post-Golgi markers. Mutagenesis of the ALPS motif and the inversion of the ALPS sequence in GMAPN support the conclusion that this membrane curvature sensor is targeted to specific vesicles in cells through direct protein-lipid, rather than protein-protein interactions. Our analysis has shown, remarkably, that mammalian curvature sensors expressed in yeast cells preserve their capacity to target specific vesicles, those of the early secretory pathway for ALPS motifs, and endocytic/post-Golgi vesicles for α-synuclein. The membrane composition of these vesicles corresponds to the preferred in vitro liposome binding properties of these membrane curvature sensors. The contrasting chemistries of ALPS motifs and α-synuclein are well adapted to each of these two major membrane environments in the cell. The HeliQuest algorithm is designed to search databases for membrane-binding amphipathic helices, including ALPS motifs. A new module designed to identify amphipathic helices with properties similar to α-synuclein has recently been developed. Searches of both yeast and human protein databases has identified candidate α-synuclein-like amphipathic helices in numerous proteins. We prepared a set of probes, in which these helices are displayed at the end of the GMAPN coiled-coil. An initial study of their co-localization in yeast cells with a set of organelle markers demonstrates specific localization patterns, suggesting that these helices may have specific membrane targeting capacities. Further work will explore the question of whether these amphipathic helices are part of a novel class of α-synuclein-like curvature sensors.

This present study aimed to: 1) Update and expand the data pool of normal Korean-English speaker’s generative naming task from previous data in Food, Clothes, and Animal categories, 2) analyze the relationship between language proficiencies and total number of words and different categories, and 3) provide easier means of testing Korean-English bilinguals with aphasia through translated standardized tests such as Aphasia Language Performance Scale (ALPS) and Boston Naming Test (BNT). Five additional subjects were added to 25 participants from Kim (2010). The participants were asked to name as many different items as possible in 60 seconds in Food, Clothes, and Animal category in both English and Korean. The participants generated more items in Korean than in English. A significant negative correlation was observed between number of words generated in Korean and Korean proficiency and between number of category doublets produced and language proficiency difference scores. A significant positive correlation was observed between number of words generated in English and English proficiency. Large differences in the number of words generated were observed between the participants assessed and participants from Kim (2010), indicating education level and field of study impacts generative naming ability. / text

Mountain glaciers are expected to be the greatest contributor to sea level rise over the next century. Glaciers provide a good indicator of global climate and how to monitor their change is an increasingly important issue for climate science and for sea level rise forecasts. However, there has been little direct measurement of glacier volume change in New Zealand. This study explores the use of remotely sensed data for measuring glacier volume change from 1965 to 2006. Digital photogrammetric methods were used to extract topographic data of the Tasman Glacier from aerial photography and ASTER imagery for the years 1965, 1986, 2002 and 2006. SRTM C band data from 2000 were also analysed. Data were compared to an existing digital elvation model produced from the New Zealand Digital Topographic Database to test for their reliability. Using regression analysis, the data were filtered and points representing rock were used to correct points on the glacier ice for vertical bias. The quality of the data extracted from the aerial photography was good on rock and debris covered ice, but poor on snow. The data extracted from ASTER was much more reliable on snow in the upper glacier than the aerial photography, but was very poor in the lower debris covered region of the glacier. While the quality of the SRTM data is very high, there is a second order distortion present in the data that is evident over elevation differences. However, the overall mean difference of the SRTM rock from TOPODATA is close to zero. An overall trend could be seen in the data between dates. However, the 2006 ASTER data proved unreliable on the debris covered section of the glacier. Total volume change is therefore calculated for the period between 1965 and 2002. The data show a loss of 3:4km³ or 0:092km³ per year, an estimated 6% of the total ice in New Zealand. This is compared to estimates using the annual end of summer snowline survey between 1977 and 2005 of 1:78 km³, or 0:064km³ per year. The spatial resolution of ASTER makes high temporal resolution monitoring of volume change unlikely for the New Zealand glaciers. The infrequency of aerial photography, the high cost and vast time involved in extracting good quality elevation data from aerial photography makes it impractical for monitoring glacier volume change remotely. However, SRTM and other radar sensors may provide a better solution, as the data do not rely heavily on user processing.

The thesis explores the work of two disparate figures, John Ruskin (1819-1900) and Albert Smith (1816-1860) who, together, helped transform the way the Alps were perceived in the mid nineteenth century. Both esteemed the Alps in their own way, although Ruskin’s cultural aestheticism contrasting markedly to the popular showmanship of Smith. Nevertheless, both Ruskin’s five-volumed Modern Painters (1843-1860), and Smith’s theatrical shows describing his ascent of Mont Blanc (1852-1858), contributed significantly to the growing popularity of the landscape resulting in the Alpine Club (1857) and the birth of modern tourism in the region. This work examines in detail the work and interests of both characters. This includes Ruskin’s drawings, art theory (especially in relation to his admiration of Turner), geological interests, religious convictions, and poetry. These reveal his desire to centre ideas of the sublime around his scientific interest in the area and the legacy of his Evangelical upbringing. The thesis investigates the tension between these elements. Smith’s climb of Mont Blanc (1851) and his subsequent shows highlighted his desire to thrill and entertain. For him, presentation of the Alps was a matter of showmanship and the thesis investigates his success, tracing its roots in elements of Victorian popular entertainment. Both Smith’s shows, and works like Of Mountain Beauty (Volume IV of Modern Painters (1856)), inspired many to explore the landscape for themselves. For Ruskin, this led to a decline in his interest in the Alps following the development of the rail network and the expansion of popular tourist sites, including his beloved Chamonix. For Smith, the public’s increasing familiarity with the region, and the popularity of other stories of Alpine ascents by members of the Alpine Club, led to a decline in interest in his shows by the end of the 1850s. Due to their interest in the region, the Romantic appreciation of the Alps in the early nineteenth century associated with theories of the sublime became a much more diverse phenomenon illustrating a number of key features of Victorian culture, including: the relationship of ‘high’ and ‘popular’ culture; the increasing influence of mass tourism; and the ways in which major figures in Victorian Britain explored and utilised foreign destinations. The thesis will also, from time to time, examine the relationship between cultural and visual forms and key elements in Victorian intellectual controversy, including the relationship of religion and science.

Lorsque l’extension de la croûte sous les parties hautes des chaînes de montagnes est colinéaire à la direction de convergence, il est traditionnellement admis que le moteur est l’effondrement gravitaire. Pourtant, des études récentes remettent en cause ce paradigme en montrant que l’érosion induit un soulèvement et de l’extension dans la partie centrale des chaînes de montagne à faible taux de convergence. L’objectif de notre étude est d’étudier l’impact de la dénudation de la topographie sur le régime sismo-tectonique des chaînes de montagnes.La première partie de ce travail présente une compilation de données dans les chaînes de montagnes afin de dégager des relations entre régime sismo-tectonique et érosion. Sur la base de ces observations, un modèle cinématique simple permettant de prévoir le régime de la chaîne est proposé. Ainsi, pour les chaînes à faible taux de convergence et d’élévation moyenne, ce modèle prédit de l’extension lorsque le taux de dénudation est 15% plus élevé que le taux de convergence.La deuxième partie est consacrée au développement d’un modèle thermo-mécanique 2D en éléments finis pour étudier l’impact des processus de surface sur la déformation des Pyrénées. Les résultats montrent que la réponse isostatique à l’érosion permet de réactiver des structures pré-existantes. La cinématique d’un plan de faille hérité peut être prédite grâce au gradient du profil des vitesses de surface horizontales. Ainsi, un plan situé dans la zone d’érosion est réactivé en faille normale alors qu’en bordure de cette zone une faille est réactivée en régime inverse. Ces résultats suggèrent que la déformation actuelle des Pyrénées pourrait être la conséquence d’un processus d’érosion.Compte tenu du faible nombre d’études quantifiant les taux d’érosion dans les Pyrénées, les modèles développés dans la deuxième partie souffrent d’une forte incertitude. Pour y remédier, nous avons cherché à les quantifier dans les Pyrénées Centrales grâce à une étude qui combine deux types de données : taux de dénudation des bassins versants à partir des isotopes cosmogéniques, et vitesses d’incision à partir des sédiments piégés dans les karsts). Ces résultats sont présentés dans la troisième partie. Les profils de dénudation obtenus sont compatibles avec un rejeu en faille normale d’un plan situé dans la Zone Nord Pyrénéenne des Pyrénées Centrales. Dans les Alpes, une bonne corrélation apparaît entre la valeur du taux d’érosion et la vitesse verticale géodésique, ce qui pose la question de l’impact de la déglaciation tardi-Wurmienne dans les Alpes sur la déformation actuelle. Un modèle numérique détaillant cette relation est présenté dans le quatrième chapitre. Les résultats montrent que la déglaciation des Alpes occidentales est contrôlée par l’hétérogénéité rhéologique de la croute. Certains de nos modèles prédisent des vitesses de surrection compatibles avec celles mises en évidence par la géodésie. / When mountain ranges upper parts express crustal extension direction collinear to the convergence direction, it is traditionally accepted that the extensive motor is gravitational collapse. However, recent studies challenge this paradigm by showing that erosion induces uplift and extension in the central part of the low convergent mountain ranges. Our goal is to investigate the impact of the denudation on the seismotectonic regime of mountain ranges.In order to identify a relationship between seismotectonic regime and erosion, the first part of this work presents a compilation of data in the mountain ranges. Based on these observations, a simple kinematic model is proposed to predict the seismotectonic regime of the chain. Thus, for low convergence rate chains with a moderate mean elevation, this model predicts an extension regime when the denudation rate is 15% higher than the convergence rate.The second part is devoted to the development of thermomechanical 2D finite element model to study the impact of surface processes on the deformation of the Pyrenees. The results show that the isostatic response to erosion reactivates pre-existing structures. The kinematics of an inherited fault plane can be predicted due to the gradient of the horizontal surface velocities profile. Thus, a plane located in the eroded zone is reactivated in normal fault when in a border area of this same plane is reactivated in reverse fault. These results suggest that the current deformation in the North Pyrenean Zone could be the result of surface processes.Given the small number of studies quantifying erosion rates in the Pyrenees, the models developed in the second part suffer from high uncertainty. To remedy this, we sought to quantify it in the central Pyrenees through a study that combines two types of data: watershed denudation rates from cosmogenic isotopes concentration, and incision rates from sediments buried in the karst. These results are presented in chapter 3. Denudation profiles obtained are consistent with a replay of a normal fault plane located in the North Zone of the Central Pyrenees.In the Alps, a good correlation appears between the value of the rate of erosion and geodetic vertical velocities, which raises the question of the impact of the late-würmian deglaciation in the Alps on the present deformation. A numerical model detailing this relationship is presented in the fourth chapter. The results show that deglaciation of the western Alps is controlled by the rheological heterogeneity of the crust. Some of our models predict uplift rates consistent with those highlighted by geodesy.

Les régions alpines sont particulièrement sensibles aux changements climatiques en cours. Ainsi, l’ouest des Alpes s’est réchauffé deux fois plus vite que l’hémisphère Nord au cours du XXème siècle. Les rythmes saisonniers des arbres, comme beaucoup d’autres organismes, sont fortement modifiés par le réchauffement climatique. La phénologie et les variations temporelles fines du climat apparaissent comme des composantes incontournables à prendre en compte pour prédire la répartition des espèces. L’objectif principal de ce travail de thèse a été de comprendre la réponse de la phénologie des espèces arborées au réchauffement climatique dans les Alpes et de développer des outils pour évaluer cette réponse dans le futur. Pour atteindre cet objectif nous avons utilisé des données phénologiques (débourrement, floraison, senescence foliaire,) pour le noisetier, le frêne, le bouleau, le mélèze et l’épicéa, issues du programme de sciences participatives Phénoclim.Nos résultats montrent que le réchauffement de l’hiver retarde la levée de la dormance des bourgeons et par conséquent les dates de débourrement et de floraison le long du gradient d’altitude. Cet effet est plus important à basse altitude. La robustesse des projections des modèles de répartition basés sur les processus dépend fortement de la robustesse des modèles phénologiques qu’ils utilisent. En comparant des modèles phénologiques présentant différents niveaux de complexité nous avons montré que les modèles basés sur les processus étaient les plus robustes particulièrement lorsque l’estimation de leurs paramètres reposait sur une estimation directe à l'aide de mesures expérimentales. Les modèles prévoient une réduction des écarts entre les dates de débourrement le long du gradient d'altitude pour toutes les espèces d'ici la fin du 21e siècle. Ceci est dû d’une part à un avancement des dates de débourrement à haute altitude et d’autre part à un retard des dates de débourrement à basse altitude. Nous avons également testé de nouvelles hypothèses sur le déterminisme environnemental de la croissance cellulaire dans les bourgeons, mais aucune des hypothèses testées n’a significativement amélioré les performances des modèles. Nous avons ensuite intégré les modèles phénologiques les plus performants que nous ayons obtenus au modèle d’aire de répartition basé sur les processus PHENOFIT. Nous avons réalisé pour la première fois avec ce modèle des simulations à haute résolution spatiale. Les projections du modèle montrent que les espèces arborées devraient se déplacer vers le haut du gradient d’altitude. Cependant, des phénomènes d’extinction locale pourraient avoir lieu dans les fonds des vallées liés à des dates de floraison trop tardives qui diminuerait le succès reproducteur des individus. Selon les espèces, la limite altitudinale supérieure serait contrôlée par le risque d'exposition au gel tardif des fleurs ou par la longueur de la saison de croissance qui détermine le temps disponible pour la maturation des fruits.L’ensemble de ces résultats nous a permis d’apporter des éléments de réponse sur la dynamique future des écosystèmes forestiers altitudes face au réchauffement climatique. Ils nous ont également permis de montrer que les données du programme Phénoclim étaient de qualité suffisante pour être utilisées dans des travaux de recherche scientifique. / Mountainous regions are particularly exposed to the ongoing climate change. Indeed, in the Western Alps the temperature increased twice faster than in the northern hemisphere during the 20th century. Trees’ annual cycle, as in many other organisms, is largely affected by climate change. Phenology and the fine temporal variations of climate appear key to predict species distribution. The main objective of this PhD thesis work was to understand the response of tree phenology to climate change in the Alps and to develop tools to evaluate this response in future conditions. It has been carried out using the phenological observations (budburst, flowering, leaf senescence) of five tree species (hazel, ash, birch, larch and spruce) of the citizen science program Phenoclim.Our results show that warmer winters slow down bud dormancy break, and consequently the budburst and flowering dates along the elevation gradient. This effect is stronger at low elevation. The robustness of process-based species distribution models depends strongly on the robustness of their process-based phenology sub-model. By comparing different phenology models differing in their level of complexity and we showed that process-based models were the most robust especially when their parameter estimates relied on forward estimation using experimental data. Models project a reduction in the phenological cline along the elevation gradient by the end of the 21th century. This is due, on one hand, to an advancement of the budburst dates at high elevation and on the other hand, to a delay of the budburst dates at low elevation. We also tested several hypotheses on the environmental determinism of bud cell growth. However, none of the hypotheses improved significantly the models’ performance. We then implemented the best phenology models we obtained in the process-based species distribution model PHENOFIT. We carried out for the first time simulations at high spatial resolution. Projections showed that species are expected to move up along the elevation gradient in response to climate change. However, local extinction events may occur in the bottom of the valleys due to late flowering dates that would decrease the reproductive success. Depending on the species, the upper altitudinal limit would be controlled by the risk of flowers’ exposure to late spring frost or to the length of growing season, which determine fruit maturation success.All of these results, allowed us to provide some answers on the future dynamics of high altitude ecosystems in the face of global climate change. They also allowed us to show that the Phenoclim data were of sufficient quality to be used to address important scientific questions.

The rapid rate of global warming currently underway highlights the need for a deeper understanding of abrupt climate change. The Younger Dryas is a Late-Glacial climate event of widespread and unusually rapid change whose study can help us address this need for increased understanding. Reconstructions from the glacial record offer important contributions to our understanding of the Younger Dryas due to (among other things) the direct physical response of glaciers to even minor perturbations in climate. Because the glacier equilibrium line altitude (ELA) provides a more explicit comparison of climate than properties such as glacier length or area, ELA methods lend themselves well to paleoclimate applications and allow for more direct comparisons in space and time. Here we present a physically based ELA model for alpine paleoglacier climate reconstructions that accounts for differences in glacier width, glacier shape, bed topography and ice thickness, and includes error estimates using Monte Carlo simulations. We validate the ELA model with published mass balance measurements from 4 modern glaciers in the Swiss Alps. We then use the ELA model, combined with a temperature index model, to estimate the changes in temperature and precipitation between the Younger Dryas (constrained by 10Be surface exposure ages) and the present day for three glacier systems in the Graubünden Alps. Our results indicate an ELA depression in this area of 320 m ±51 m during the Younger Dryas relative to today. This ELA depression represents annual mean temperatures 2.29 °C ±1.32 °C cooler relative to today in the region, which corresponds to a decrease in mean summer temperatures of 1.47 °C ±0.73 °C. Our results indicate relatively small changes in summer temperature dominate over other climate changes for the Younger Dryas paleoglaciers in the Alps. This ELA-based paleoclimate reconstruction offers a simple, fast, and cost-effective alternative to many other paleoclimate reconstruction methods. Continued application of the ELA model to more regions will lead to an improved understanding of the Younger Dryas in the Alps, and by extension, of rapid climate events generally.

This thesis investigated marine isotope stage {MIS) 3-2 glacial sequences in the South Fork Hoh River Valley, Washington and the Lake Hawea Valley, New Zealand. Research objectives were to reconstruct the style and timing of ice advance in both areas and to assess the viability of luminescence dating of glacial sediments in various depositional facies and distances from the ice front. This thesis focused on the sedimentology and stratigraphy of surficial and older glacial sequences in the South Fork Hoh and Lake Hawea areas and used OSL and radiocarbon dating techniques to establish age control for the deposits. Specifically, this research identified, described, and dated the stratigraphy of glacial sequences in order to reconstruct ice dynamics. This work also presents updated geomorphic maps for both study areas as an additional way to show ice advance and retreat events recorded in deposited sediment and geomorphic surfaces. The glacial sequence expressed in the Lake Hawea moraine exposure shows four distinct depositional events that represent retreat from an ice position down -valley, re-advance to the Hawea moraine position, and subsequent retreat and deglaciation broadly spanning -32-18 ka. These results document the terminal glacial advance and subsequent retreat in the Lake Hawea Valley and contribute to the wider swath of research studying the last phase of glacial retreat and its connections to climate on the South Island of New Zealand. The Hawea chronology corresponds to other glacial records and paleoclimate reconstructions from the South Island that collectively suggest the commencement of deglaciation at -13 ka. Three late Pleistocene ice positions are preserved in the South Fork Hoh River Valley, here referred to as South Fork 1-3 (SF 1-3). One of these positions has not previously been recognized in this valley or in the mainstem Hoh River Valley. Optically stimulated luminescence (OSL) and radiocarbon (14C) ages are generally consistent throughout the valley. These finding s advocate for a detailed sedimentologic and stratigraphic approach to glacial depos its and questions whether a similar advance or still -stand occurred in other valleys in the region. If so, this may reveal information regarding climate influences on MIS 2 glaciers in the Olympic Mountains. This research also assesses the applicability of OSL dating to glacial deposits in both field areas. Quartz OSL dating was used in the South Fork Hoh study area; however, quartz produced unreliable results in the Hawea study area, so samples were therefore assessed using feldspar methods. The results advocate for a facies-based sampling approach in glacial settings, where better sorted sandy facies and more distal deposits produce better bleached and more reliable age results than other deposits.