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Three kinds of processes are known to modify the geographical spatial arrangement of organisms: dispersal, extinction and vicariance. The Southern Hemisphere has an intriguing and complicated geological history that provides an ideal backdrop to study these processes. This thesis focuses on three historical events that illustrate these processes: the proposed marine inundation of New Zealand in the Oligocene, the asteroid impact at the K – Pg boundary, and the continental breakup of Gondwana. It investigates what impact these events had on species diversification by studying the phylogenetic relationships of two groups of taxa – the family Anostostomatidae (insects), and Neoaves (birds). Anostostomatidae were studied in relation to the Oligocene drowning and the break up of Gondwana as they have a wide southern distribution, found on all “Gondwanan” fragments with the exception of Antarctica, and are thought represent an ancient lineage that predates the Gondwanan breakup. Birds, in particular Neoaves, were studied in relation to the asteroid impact at the K – Pg boundary. Although birds are mobile and many circumnavigate the globe between seasons, they are suggested to have originated in the Southern Hemisphere in Gondwanan times, and subsequently undergone range expansion and diversification around the world. In order to address the relationship (if any) between modern biotic diversity and historical geological events, phylogenetic relationships were determined and where possible, molecular clock analysis carried out. Timing information provided by molecular clock analysis is important as it enables distinction between opposing hypotheses such as vicariance and dispersal. In Chapter Two, the phylogenetic relationships within the family Anostostomatidae are investigated. One of the most controversial times in New Zealand’s geological history is during the Oligocene. Some suggest that the lack of fossils and evidence for recent dispersal of numerous taxa support the notion that all modern biota reached the region during the last 25 million years. Anostostomatidae were chosen as they represent a group of insects that are thought to be ancient and there is little published data in the literature. Previous studies focused on the relationships within Hemideina and Deinacrida suggesting that these groups diversified in the early Miocene. The data presented here are from mitochondrial (COI and 12S) and nuclear (18S and 28S) sequences. Molecular dating using a relaxed clock as implemented in BEAST suggest that in fact some lineages were present at or shortly after continental breakup and could have survived throughout this turbulent time. As there were no definitive fossils to use for calibration points, geological events were used as calibration points for the molecular clock. Mutation rates obtained from the different analyses were compared to those published for other insects in an attempt to identify the most likely model. Both maximum likelihood and Bayesian analyses support the presence of three distinct ecological groups in New Zealand; Hemiandrus (ground weta), Anisoura/Motuweta (tusked weta) and Hemideina–Deinacrida (tree–giant weta). With regards to their Australasian relatives (taxa from Australia and New Caledonia) it appears that the family is divided with the most northern New Zealand taxa (tusked weta) more closely related to New Caledonian taxa while all other New Zealand taxa are more closely related to Australian taxa. There does not appear to be any link between the Australian and New Caledonian taxa studied here. Results should be viewed with caution however as an increased mutation rate was observed in the New Caledonian-tusked weta lineage, something future studies will have to address. Chapter Three presents new sequence data and phylogenetic analyses that go towards resolving the apparent basal polytomy of neoavian birds. This chapter includes analyses carried out on previously published data with the addition of nine new mitochondrial genomes. My contribution to this larger project was to perform the phylogenetic analysis and to sequence three of the nine mitochondrial genomes. The genomes I sequenced were the Southern Hemisphere species: dollar bird (Eurystomus orientalis), Owlet nightjar (Aegotheles cristatus cristatus) and great potoo (Nyctibius grandis). The inclusion of these nine new genomes allows assessment and comparison of the six hypothesised groups reported in Cracraft (2001). First an improved conditional down-weighting technique is described reducing noise relative to signal, which is important for resolving deeper divergences. Second, a formula is presented for calculating probabilities of finding predefined groupings in the optimal tree. Maximum likelihood and Bayesian based phylogenetic analyses were carried out and in addition, dating using a relaxed molecular clock was performed in BEAST. Results suggested that the six groups suggested by Cracraft (2001) represent robust lineages. The results suggested that one group, the owls, are more closely related to other raptors, particularly accipitrids (buzzards/eagles) and the osprey rather than the Caprimulgiformes, which could indicate morphological convergent evolution. In addition, a group termed shorebirds appears to be distinct from the large group referred to as ‘Conglomerati’ to which previous publications have suggested they belong. The ‘Conglomerati’ is the least well studied group and may actually comprise of at least three subgroups (as suggested by Cracraft). Within the three suggested groups, Cracraft grouped shorebirds with pigeons and sandgrouse, neither of which (pigeons or sandgrouse) were analysed here. So although the shorebirds are at least close to the ‘Conglomerati’ and may be within that group, their exact position is still not clear. The molecular dating reported here utilised two fossil calibrations (Vegavis and Waimanu), for which there is relatively little dispute as to age or the lineage to which they belong. Calibrations resulting from BEAST analyses suggest that at least 12 distinct lineages were present prior to the K – Pg boundary, a finding supported by previous studies. Robust phylogenies will allow future studies to investigate not only the relationships within Neoaves, but look more closely at the biological and ecological evolution of the group. Chapter Four for the first time investigates whether the phylogenetic relationships within the family Anostostomatidae follow the conventionally accepted order and timing of Gondwanan breakup. Following the initial restults for taxa studied in Australasia (Chapter Two) an attempt to resolve family relationships in a wider spatial (geographic) context was carried out to determine if Australasian taxa are monophyletic when other members of the family are included. Again both maximum likelihood and Bayesian phylogenetic analyses were carried out on both mitochondrial (COI and 12S) and nuclear (18S and 28S) sequences. In this chapter, datasets included samples from across the geographic range of Anostostomatidae (South Africa, Madagascar, South America, Australia, New Caledonia and New Zealand), and two clades were observed, congruent with earlier findings. Sequence divergence within geographic regions was found to be relatively high in the mitochondrial genes (COI and 12S) while low in the nuclear ribosomal RNA genes (18S and 28S) as expected given their relative mutation rates. Under the vicariance paradigm, phylogenetic relationships should follow the order of continental breakup, but this was not found. Further, if dispersal and colonisation were continuous, no geographic substructure is expected, however distinct geographic substructure within clades was consistently observed. This interesting phylogenetic pattern may be a case of convergent evolution or paraphyletic sampling which highlights taxonomic issues of the group. Future studies need to include not only molecular data but information on morphology, ecology and behaviour along with the implementation of biogeographic programs that can test alternative hypotheses (such as dispersal and vicariance) directly. Also, the inclusion of the recently reported fossil from the subfamily Euclydesinae (Martins-Neto 2007) should allow for more accurate date estimates within the family. Taken as a whole the results presented in this thesis suggest that microevolutionary processes are sufficient to explain modern diversity without the need to invoke abiotic events. The three cases investigated here - marine inundation, asteroid impact and continental drift - all appear to have had only a limited effect on the diversity of taxa studied. To reach even stronger conclusions future studies should incorporate different data (for instance nuclear genes, intron position, and genome structure) and use biogeographic software capable of including ecological, morphological and habitat information.

L'histoire géodynamique du Sud-Ouest Pacifique est contrôlée depuis le Mésozoïque par l'évolution des zones de subduction péri-Pacifique qui ont successivement bordé la marge Est du Gondwana. Le recul de la fosse de subduction par effondrement du panneau plongeant dans le manteau est probablement le moteur de la fragmentation continentale qui a donné naissance à des rifts avortés et des bassins arrière-arcs associés à des arcs volcaniques rémanents. Dans ce contexte géodynamique, les bassins ayant atteint un stade d'océanisation avancé ont enregistré les inversions du champ magnétique terrestre et développé des morphologies typiques de la croûte océanique. Ces caractéristiques ont permis d'identifier l'âge et la nature de la croûte de ces bassins avec un degré de confiance élevé. C'est le cas de la plupart des bassins du Sud-Ouest Pacifique formés après 45 Ma. A l'inverse, les bassins de Nouvelle-Calédonie et de Fairway, plus étroits et recouverts d'épaisses séries sédimentaires ont une origine mal renseignée qui est longtemps restée controversée. Bien que morphologiquement et structuralement remarquables à l'échelle régionale, ces bassins n'interviennent pas dans les schémas actuels de reconstruction géodynamique régionaux. L'objet du présent travail de thèse est d'approfondir la connaissance de la structure et de l'histoire de ces bassins afin d'affiner le puzzle géodynamique du Sud-Ouest Pacifique. Une synthèse géologique régionale, accompagnée d'une carte structurale (planche hors texte A0), permet de replacer ces bassins dans le contexte géologique de l'évolution post-jurassique de la marge Est-Australienne. Les nouvelles données sismiques d'imagerie profonde des campagnes ZoNéCo-11, Noucaplac-2 et TL-1, couplées aux données récentes de forages sur la marge de Taranaki en Nouvelle-Zélande, constituent une base solide de données nouvelles autorisant une interprétation chronostratigraphique rénovée des bassins de Fairway et de Nouvelle-Calédonie ainsi que des rides qui leur sont associées. Ces interprétations complétées par une nouvelle compilation des données gravimétriques et magnétiques régionales permettent de dégager trois étapes dans l'évolution géodynamique de ces bassins : Etape 1 : Formation du Bassin de Fairway - Aotea au Crétacé moyen, en position intra- ou arrière-arc continental, dans un contexte de subduction. La formation de ce bassin, relativement peu profond, reflète les prémices de la déchirure continentale de la marge Est-Gondwanienne au Cénomanien, dont la cause est à rechercher dans un changement de la dynamique de la subduction. Une « verticalisation » de la plaque plongeante aurait entraîné de l'extension dans la plaque chevauchante. L'augmentation du pendage du slab pourrait être attribuée à : (i) un processus gravitaire lié au poids du slab, l'amenant à couler dans l'asthénosphère, engendrant alors un « hinge rollback », (ii) un flux asthénosphérique rétrograde, exerçant une pression horizontale sur le slab le faisant ainsi reculer, et (iii) la cinématique des plaques lithosphériques impliquées dans la subduction pouvant jouer un rôle sur le pendage du slab et pouvant créer des zones de faiblesse dans la plaque chevauchante menant à de l'extension arrière-arc. Un phénomène de « tectonic escape », lié à un blocage de la zone de subduction par le plateau d'Hikurangi à 105 Ma, pourrait aussi être la cause du recul de la subduction menant à de l'extension arrière-arc. Etape 2 : Déformation locale affectant la partie Nord du Bassin de Nouvelle-Calédonie (au large de la Grande Terre, strictement) à l'Eocène terminal, synchrone de l'obduction ophiolotique néo-calédonienne. Cette déformation asymétrique, d'une durée de quelques millions d'années et d'amplitude verticale de l'ordre de 10 km, est marquée par la surrection de la ride de Fairway et la subsidence de la marge Est du bassin, le long de la côte Ouest de la Nouvelle-Calédonie. Le Bassin de Nouvelle-Calédonie aurait subsidé sous l'effet de la surcharge engendrée par le charriage de la croûte océanique du Bassin de Sud-Loyauté sur la ride de Norfolk à 37 Ma et aurait réagi comme un bassin flexural d'avant-pays suivant un processus de sous-charriage (underthrusting), au fur et à mesure que la nappe progressait sur le bâti Calédonien. Etape 3 : Subsidence Eocène-Oligocène d'envergure régionale, affectant l'ensemble « Ride de Lord Howe, Bassin de Fairway – Aotea, Ride de Fairway, Bassin de Nouvelle-Calédonie, Ride de Norfolk », de la Nouvelle-Zélande à la Nouvelle-Calédonie. Les traits morphostructuraux associés à cette subsidence suggèrent que l'ablation d'une partie de la croûte inférieure du système est responsable de cette subsidence. Nous émettons ainsi une nouvelle hypothèse selon laquelle les bassins de Nouvelle-Calédonie et de Fairway-Aotea, initialement peu profonds pendant le Crétacé (phase de rifting Crétacé, étape 1), auraient subi une subsidence de grande ampleur à l'Eocène – Oligocène lors de la réactivation de la convergence Australie – Pacifique. La reprise de la convergence vers ~45 Ma aurait entraîné un épaississement crustal de l'ensemble de rides et bassins aboutissant à une instabilité gravitaire de sa racine, provoquant ainsi son détachement et son effondrement dans le manteau. Ces nouvelles interprétations, et en particulier l'âge Crétacé moyen des sédiments les plus anciens des bassins de Fairway et Aotea, ont des implications pour le potentiel pétrolier de la région.

The southern Patagonian Andes record the Late Cretaceous closure and inversion of the Late Jurassic – Early Cretaceous Rocas Verdes marginal basin, subsequent development of the Patagonian retroarc fold-thrust belt and the Neogene to present tectonic superposition of a left-lateral strike-slip plate margin defined by the Magallanes- Fagnano fault zone. In this dissertation, I present new geologic maps, cross sections and detailed macro- and microscopic structural analyses that describe the geometry and kinematic evolution of the fold-thrust belt and superposed strike-slip deformation over ~200 km along-strike between 53° and 55° S latitude. Results are discussed in the context of the regional tectonic development of the southernmost Andes and are relevant to the understanding of important tectonic processes including the development of a retroarc fold-thrust belt, the formation of a basal décollement below and toward the hinterland of a fold-thrust belt and the spatial distribution of deformation along a strike-slip plate margin. New maps and balanced cross-sections of the Patagonian fold-thrust belt show that it developed during two main phases of Late Cretaceous to Paleogene shortening that were partly controlled by the antecedent geology and mechanical stratigraphy of the Rocas Verdes basin. During the Late Cretaceous, a thin-skinned thrust belt developed above a décollement that formed first in relatively weak shale deposits of the Rocas Verdes basin and later deepened to <1 km below the basement-cover contact. Ramps that cut mechanically rigid volcanic rocks of the marginal basin link the two décollements. Basement-involved reverse faults that cut the early décollements and probably reactivate Jurassic normal faults reflect Paleogene shortening. Shortening estimates increase northwest to southeast from 26 to 37% over 100 km along-strike and are consistent with regional models of the fold-thrust belt. Structural data, kinematic analyses, and microstructural observations from the lower décollement show that it is defined by transposition of several generations of northeast-vergent noncylindrical folds, shear bands, and a quartz stretching lineation that are kinematically compatible with first-generation structures of the fold-thrust belt. Quartz microstructural data from the décollement are consistent with deformation temperatures that decrease from ~500-650° C to ~400-550° C over ~75 km in the transport direction, indicating that the décollement dipped shallowly (~6°) toward the hinterland. The décollement decoupled the underthrust continental margin from the fold- thrust belt and exemplifies the kinematic relationship between shortening that occurs coevally in a retroarc fold thrust-belt and its polydeformed metamorphic ‘basement’. Fault kinematic data and crosscutting relationships show kinematic and temporal relationships between populations of thrust, strike-slip and normal faults that occur in the study area. Thrust faults form an internally compatible population that shows subhorizontal northeast-trending shortening of the fold-thrust belt and is kinematically distinct from populations of normal and strike-slip faults. Both strike-slip and normal faults crosscut the fold-thrust belt, are localized near segments of the Magallanes- Fagnano fault zone, have mutually compatible kinematic axes and are interpreted to be coeval. Strike-slip faults form Riedel and P-shear geometries that are compatible with left-lateral slip on the Magallanes-Fagnano fault-zone. Strike-slip and normal faults occur in a releasing step-over between two overlapping left-lateral, left-stepping segments of the Magallanes fault zone and record a tectonic event defined by sinistral transtension that probably reflects changing plate dynamics associated with the opening of the Drake Passage during the Early Miocene. / text

The island of Sri Lanka’s exceptional biodiversity and enigmatic biogeography begs investigation, as the island is key in understanding the evolution of the Asian tropical flora. Since the Jurassic, Sri Lanka has been subjected to remarkable tectonic changes, thus its flora could have been influenced by that of a number of nearby landmasses, as well giving Sri Lanka the potential to have played a wider role in the assemblage of floras elsewhere. Firstly, as Sri Lanka originated as a fragment of the supercontinent Gondwana, part of its flora may contain Gondwanan relict lineages. There is also the potential for immigration from Laurasia after the Deccan Plate collided with it 45-50 Mya. Further, Sri Lanka may harbour floristic elements from nearby land masses such as Africa and Southeast Asia as a result of long distance dispersals, and in situ speciation has the potential to have played an important role in enhancing the endemic Sri Lankan flora. I tested the relative contributions of the above hypotheses for the possible origins of the Sri Lankan flora using three representative families, Begoniaceae, Sapotaceae and Zingiberaceae. These families represent both herbaceous and woody elements, and have high diversity across the tropics. Dated molecular phylogenies were constructed for each family. I used recent analytical developments in geographic range evolution modelling and ancestral area reconstruction, incorporating a parameter J to test for founder event speciation. A fine scale area coding was used in order to obtain a better picture of the biogeography of continental Asia. Amongst all the models compared, a dispersal-extinction cladogenesis model incorporating founder event speciation proved to be the best fit for the data for all three families. The dates of origin for Sri Lankan lineages considerably post-date the Gondwanan break up, instead suggesting a geologically more recent entry followed by diversification of endemics within the island. The majority of Sri Lankan lineages have an origin in the Sunda Shelf (53%). Persistence of warm temperate and perhumid climate conditions in southwestern Sri Lanka resembling those of Peninsular Malaysia and Sumatra could have facilitated suitable habitats for these massive dispersals from the Sunda Shelf region. Some trans-oceanic long distance dispersals from Africa (11%) are also evidenced, again these are too young to accept a hypothesis of dispersal during the Deccan Plate’s migration close to the African coast during the late Cretaceous, but occurred later during the Miocene. Further, some lineages of Laurasian origin (20%) are evidenced in the Zingiberaceae with ancestral areas of China and Indochina, which is congruent with a post collision invasion. Among the families tested, dispersals have occurred stochastically, one during the Eocene, six during the Oligocene, seven during the Miocene, two during the Pliocene and one during the Pleistocene. The highest number of dispersals occurred during the Miocene when a warm climate was prevailing during the Miocene thermal maximum. My results confirm that in situ speciation is an important contributor to the Sri Lankan flora. More rapid radiation of endemics has occurred during Pliocene-Pleistocene; two endemics in Begoniaceae, ten endemics in Sapotaceae and ten endemics in Zingiberaceae have evolved in situ during this period. Sri Lanka will have been subjected to expansion and contraction of climatic and vegetation zones within the island during glacial and interglacial periods, potentially resulting in allopatric speciation. As a conclusion, long distance dispersals have played a prominent role in the evolution of the Sri Lankan flora. The young ages challenge the vicariant paradigm for the origin and current disjunct distributions of the world’s tropical lineages and provide strong evidence for a youthful tropics at the species level. The thesis contains six chapters; first two are introductory chapters, then there are three analytical chapters, one for each family, and finally a summary chapter is provided. Each analytical chapter is written as a stand-alone scientific publication, thus there is some repetition of relevant content in each.

M.Sc. (Geology) / The Mzimvubu River, situated in the Eastern Cape Province of South Africa, drains essentially strata of the Late Carboniferous to Jurassic Karoo Supergroup with minor intersection of the underlying Devonian Msikaba Formation near the mouth of the river at Port St. Johns. Rock- and river sediment samples were collected at specific points from within the Mzimvubu River drainage basin, based on changes in the geology through which the rivers flow. Detrital zircon age population data was obtained by LA-ICP-MS for each sample in order to meet the two-fold objective of the study; firstly to investigate the reliability of using detrital zircon grains as indicators of sedimentary provenance and secondly to determine possible source areas for the Karoo strata and underlying Msikaba Formation. Through the comparison of detrital zircon age population data for the rock units of the Karoo Supergroup and Msikaba Formation to that of the river sediment, it is concluded that detrital zircon grains hold value in deciphering the geological history of a sedimentary basin. This interpretation is based on similar distributions and trends that are present in both the zircon age populations of the rock- and sediment samples. However, complexities associated with detrital zircon analysis pertaining to rock type and depositional settings are noted and therefore certain procedures that can be implemented during field sampling have been suggested in this study so as to ensure accurate results are obtained. This will further ensure that reliable interpretations of the geological history of a sedimentary basin are achieved. Additionally, by utilising the detrital zircon population data obtained in the first part of the study in conjunction with published scientific data, the provenance of the Karoo Supergroup in the southeastern part of the Main Karoo Basin has been determined. From this data it was determined that, especially the upper part of the Karoo Supergroup in the Eastern Cape Province of South Africa, was deposited much later than previously thought and that many of the stratigraphic layers in the Karoo Basin were deposited coevally in different parts of the basin with lithostratigraphic boundaries being time-transgressive. Ultimately the data allowed for the construction of a tectono-sedimentary model to explain the deposition of the upper Cape- and Karoo Supergroups that started with the deposition of the Msikaba Formation in a passive continental margin setting, to deposition of the lower part of the Karoo Supergroup in an Andean type of foreland basin, with rifting starting during the times of deposition of the Molteno Formation. The deposition of the Molteno-, Elliot- and Clarens Formations took place as Gondwanaland was breaking apart coeval with the formation of the Karoo Igneous Province.

L’étude de l’Ordovicien de l’Anti-Atlas oriental s’intègre dans un vaste projet celui de l’étude de la plate-forme nord-gondwanienne qui se caractérise par une sédimentation essentiellement silico-clastique avec une polarité globale qui est Sud-Nord. Cette étude, basée sur les principes de la stratigraphie séquentielle et de la sédimentologie, met en évidence l’existence de deux systèmes de dépôts sur la marge nord-ouest de la plate-forme nord-gondwanienne. A l’Ordovicien moyen et supérieur préglaciaire, à l’emplacement de l’Anti-Atlas oriental, coexistent un système deltaïque (lato sensu) et un système de plate-forme dominée par l’action de la houle et des tempêtes. Ces deux systèmes de dépôts ont une certaine répartition spatiale et temporelle. Le système deltaïque domine le nord-est de l’Anti-Atlas oriental où il prend naissance à l’Est du Tafilalt. Ce système deltaïque est à dominance fluviale, il est actif lors des périodes de régression. Pendant les maximums de régression, il incise le haut du plateau continental en créant d’importantes structures érosives dont le remplissage se fait par des faciès de haute énergie à composante unidirectionnelle. Le système de haute énergie influencé par l’action de la houle et des tempêtes domine le secteur sud de l’Anti-Atlas oriental et se répand au nord au cours des périodes transgressives. La sédimentation durant l’Ordovicien préglaciaire sur la marge nord-ouest du Gondwana a un contrôle tectonique et eustatique avec une polarité des systèmes, dans le Tafilalt-Maïder qui diffère de celle de la plate-forme nord-gondwanienne et est Nord Est-Sud-Ouest. Le profil de dépôt est quasi plat où les clinoformes sont de faible amplitudes et se dessinent à l’intérieur des séquences de 3ème ordre. Le Darriwillien se caractérise par des une subsidence différentielle entre le domaine nord et le domaine sud en raison d’une tectonique active. Alors que, le Sandbien-Katien montre un taux de subsidence faible (x 10m/Ma) en adéquation avec un contexte intracratonique. L’Hirnantien dans le Tafilalt, décrit comme étant discontinu voir absent, est très bien développé il se caractérise par la mise en place d’une sédimentation mixte gréso-carbonaté à bryozoaires de type littoral et ce à l’extrémité oriental de l’Anti-Atlas et d’une épaisse série conglomératique (conglomérat d’Amessoui et d’Imzizoui) et flyschoïde qui peut atteindre les 500m dans le Tafilalt. / The Ordovician Anti-Atlas study is part of a larger project of comprehension the sedimentary system of the north Gondwana platform. This sedimentary area is characterized by essentially terrigenous sediments that come from the south from de West African Craton. This study based on sequence stratigraphy and sedimentology principles highlights two systems juxtaposed during the preglacial Ordovician period (before hirnantian). These two deposits systems have a spatial and temporal distribution. One deltaic occurs on north-eastern Tafilalt (east Anti-Atlas) and widespread during regressive periods. The second is storm dominate system which is developed in the south-western part of eastern Anti-Atlas and transgressive periods. The deltaic system is river dominant, it incise the continental shelf by creating a significant erosive structure whose filling is done by high energy facies unidirectional component. During the Ordovician, the sedimentation was controlled by tectonic and eustatic variation. The polarity of the system in the Tafilalt-Maïder is different from that of the northern Gondwana platform and it is given North East- South West. The Darriwilian characterized by differential subsidence, between the north and the south of the oriental Anti-Atlas, due to active tectonics, while, Sandbian-Katian show a low subsidence rates (x10m/Ma) in the line of intracratonic basin. The Hirnantian, in Tafilalt, is largely well developed and characterized by huge conglomeratic series, flysch deposits and Bryozoan limestone at the northeastern.

The Arabian–Nubian Shield (ANS) represents a major site of juvenile Neoproterozoic crustal addition on Earth and documents Neoproterozoic tectonics bracketed by two supercontinent cycles, namely the fragmentation of Rodinia and the amalgamation of Gondwana. There is general consensus that the ANS formed by juvenile magmatic arc accretion and subsequent shield–wide post–tectonic magmatism. However, detailed understanding about the timing of events and the nature of magma sources in parts of the shield are lacking. To date, there are no isotopic data from the Paleozoic sedimentary sequences of the ANS, except those from the northern part. New zircon U–Pb, δ18O and whole–rock Nd isotopes are presented for plutonic rocks from the eastern Ethiopia, Yemen and southernmost Arabian Shield in Saudi Arabia. This thesis also presents the first combined in situ zircon U–Pb–O–Hf isotope data on the Cambrian–Ordovician sandstones of the Arabian Shield. The results are used to elucidate the crustal evolution of these parts of the ANS and to evaluate terrane correlations. Specifically, the nature of crustal growth, i.e., relative proportions of juvenile magmatic additions vs. crustal reworking, nature of the magma source and mechanism of crust formation (plume material vs. subduction zone enrichment) and understanding the provenance of the Cambrian–Ordovician sandstone sequences were important research questions addressed. The results from Paper I suggest that the eastern Ethiopian Precambrian basement is dominated by reworking of pre-Neoproterozoic supracrustal material unlike contemporaneous rocks in the remaining parts of Ethiopia— indicating the presence of two distinct lithospheric blocks of contrasting isotopic compositions in Ethiopia. Metamorphic age distributions suggest that the eastern Ethiopian block was amalgamated with the juvenile Western Ethiopian Shield during ca. 580–550 Ma. Importantly, the suture between them may represent the northern continuation of a major suture identified further south in Africa along which Gondwana amalgamated. Similarly, the Abas terrane in Yemen (Paper II) is dominated by reworking of pre–Neoproterozoic crust and shows age and isotopic compositions that are inconsistent with the Afif terrane of Saudi Arabia, precluding correlation between the two regions. The trace element systematics of plutonic rocks from the southernmost Arabian Shield (paper III) point to enrichment due to subduction component, bear no evidence of a plume component, and are consistent with the adakite-like chemistry of some of the subduction–related plutonic samples. This reinforces the notion that the shield grew through juvenile magmatic arc additions. The combined zircon U–Pb–O–Hf data of the Cambrian–Ordovician sandstones (Paper IV) indicate their derivation from both the adjacent juvenile ANS and the more southerly crustal blocks that are dominated by reworking of pre–Neoproterozoic crust. The remarkable similarity in age spectra and homogeneity of Cambrian sandstones deposited across the northern margin of Gondwana point to continental–scale sediment mixing and dispersal regulated by the supercontinent cycle. / <p>At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 3: Manuscript. Paper 4: Manuscript.</p>

The Aegean is a classic setting for studying exhumation of high-pressure (HP) metamorphic rocks. Two end-member models are proposed to explain the uplift of these rocks: core-complex style extension along low-angle normal faults and extrusion-wedge uplift. Extrusion-wedge underplating is the mechanism that exhumed HP rocks on Evia whereas Tinos hosts several detachments varying in age from 30-9 Ma. Andros, situated between them, may be the geological manifestation of the interplay of these processes and provides an opportunity to test these models. Detachments on NW Tinos and on Andros and the enigmatic low-angle Makrotantalon Unit contact on Andros were insufficiently dated prior to this study. Geo- and thermochronometrycombined with structural observations from sampling transects in the transport direction from (1) lower plate Cycladic Blueschist Unit on Andros and Tinos, (2) middle plate Makrotantalon Unit on Andros, and (3) hanging wall Upper Unit address these issues. Maximum depositional ages from detrital zircon U-Pb geochronometry and structures reveal Paleocene-Eocene syn-HP metamorphism thrusting resulted in an inversed-age relationship between the Permian Makrotantalon Unit and the underlying Triassic-Eocene Cycladic Blueschist Unit on Andros. The Makrotantalon Unit has an internal inversed stratigraphy whereas the Cycladic Blueschist Unit on Andros and Tinos appear stratigraphically intact. Structures and zircon and apatite (U-Th)/He ages in transects from NW Tinos (~12-8 Ma) and central Andros Cycladic Blueschist Unit (~13-7 Ma) indicate rapid cooling due to exhumation associated with the Livada Detachment. Older cooling ages (~16-10 Ma) and structures in the Makrotantalon Unit indicate later brittle strain localization on the Makrotantalon Thrust contact is accommodated by rheologically weaker serpentinites and calc-schists, resulting in slivering of the footwall under the Livada Detachment on Andros. Estimated mean cooling slip rates of the Livada Detachment on Andros of ~3.8 (+1.2/-1.3) km/Myr and 2.1 (+0.2/-0.2) km/Myr on NW Tinos resulted in minimum vertical exhumations of 15 km and 4 km, respectively. The NCDS here accommodated ~12-25% of 60 km of HP-rock exhumation from ~30-7 Ma. We present a tectonic model to elucidate the evolution of the Makrotantalon Unit and the magnitude, temporal, and spatial variability of exhumation via detachments on these islands. / text

La compréhension des processus orogéniques nécessite l'acquisition de données permettant de suivre les variations de pression (P), température (T) et déformation (D) au cours du temps (t). La construction de trajets P-T-D-t implique nécessairement une approche pluridisciplinaire, qui combine une analyse pétrologique et structurale à l'acquisition de données géochronologiques. Dans ce mémoire, une telle approche est appliquée à des granulites polymétamorphiques de Ultra-Hautes Températures de l'unité basique d'Andriamena (Centre-Nord Madagascar). Une attention particulière a été portée sur la corrélation entre les données pétrologiques et géochronologiques afin de discuter la signification des trajets P-T et des âges obtenus en contexte polymétamorphique. L'évolution thermomécanique de l'unité d'Andriamena est marquée par la superposition d'au moins quatre événements thermiques distincts: ~2.7 Ga, 2.52-2.54 Ga, 790-730 Ma et 530-500 Ma. Si la signification de l'événement Archéen à 2.7 Ga reste encore problématique, l'événement fini-Archéen à 2.5 Ga correspond sans ambiguïté au métamorphisme de UHT (~1050°C, 11.5 kbar). Le Néoprotérozoïque moyen (790-730 Ma) est marqué par la mise en place d'un important complexe basique-ultrabasique contemporain d'un épisode de fusion partielle et d'un métamorphisme granulitique (~850-900°C, 7 kbar). Cet événement thermique majeur, à l'échelle du Centre-Nord Madagascar, est interprété comme le témoin d'un contexte tectonique du type arc continental lié à la fermeture de l'océan Mozambique lors de la fragmentation du supercontinent Rodinia. Le dernier événement affectant l'unité d'Andriamena (530-500 Ma) est à l'origine du champ de déformation finie, qui résulte de la superposition de deux phases D1 et D2 synchrones d'un métamorphisme amphibolitique à granulitique de basse pression (650-700°C, 5-6 kbar). La déformation Cambrienne observée dans le Centre-Nord Madagascar est compatible avec un raccourcissement horizontal Est-Ouest qui résulterait de la convergence de cratons lors de la consolidation finale du Gondwana. Par leur caractère réfractaire, les Mg-granulites de UHT préservent de nombreuses textures minéralogiques permettant de retracer un trajet PT pétrographique complexe et apparemment continu. Néanmoins, les données géochronologiques obtenues par la méthode de datation in-situ sur monazite à la microsonde électronique montrent que le trajet pétrographique ne doit pas être considéré comme issu d'un seul et même événement thermique, mais plutôt comme un trajet discontinu résultant de la superposition de deux événements distincts à 2.5 Ga et 790-730 Ma. D'autre part, nous montrons qu'une partie du trajet pétrographique (décompression isotherme) correspond à un trajet apparent sans signification tectonique. Nous suggérons que cette décompression apparente résulte de l'équilibration des paragenèses réfractaires de UHT (2.5 Ga) à plus basses pressions, lors de l'événement Néoprotérozoïque moyen (790-730 Ma), sans que les conditions P-T des réactions minéralogiques observées n'aient été atteintes. La distinction qui existe entre trajet pétrographique et trajet P-T réel montre l'importance de déterminer l'âge absolu des différents assemblages et réactions métamorphiques, afin de dater différentes portions du trajet P-T. Cet objectif est atteint grâce aux méthodes de datations ponctuelles et in-situ qui permettent de dater des minéraux dans leur contexte textural et donc de corréler âge et assemblage métamorphique. Ainsi, nous avons développé une nouvelle méthode de datation in-situ U-Th-Pb sur monazite, qui utilise les méthodes chimiques (microsonde) et isotopiques conventionnelles (ID-TIMS). Par ces deux méthodes, on combine une haute résolution spatiale (~3µm - microsonde) à une haute précision analytique (ID-TIMS). La particularité de cette nouvelle approche est que la datation isotopique est réalisée sur des grains individuels extraits par micro-forage directement sur lame mince et qui ont été au préalable caractérisés à la microsonde électronique (imagerie, datation chimique...). La position texturale de chaque grain daté est ainsi retenue.