Sclerochronology affirms the well-established fact that banded growth increments in marine molluscs accurately record oscillations in climate and environment for the past millennia or more. This study considers how such records can enhance understanding of environmental shifts across the 18th to 20th centuries. Specifically, it investigates whether sclerochronological data are fundamentally associated with stable carbon and oxygen isotope values and if the climate phenomenon of the Little Ice Age impacts the marine radiocarbon reservoir effect. Furthermore, this study compares the North Sea and the Irish Sea to discover whether observed environmental changes are equivocal or not. Three shells of Arctica islandica were retrieved from the Dutch natural history museum "Naturalis" in Leiden, the Netherlands, to construct a chronology dating back to the 18th century from growth increments. Collection dates for the three shells vary between 1882 and 1954 in the North Sea and the Irish Sea. Shells were prepared, photographed and milled for calcium carbonate material. Digital images allowed counting of increments while the milled material was measured for stable carbon and oxygen isotopes and was radiocarbon dated. The ontogenetic bias was removed from sclerochronological data using MATLAB software to produce a purely environmental growth signal. The results agreed with the proposed hypothesis that there is a link between growth increments and the stable isotope data. However, further analysis is required to validate the presence of the Little Ice Age in the radiocarbon reservoir effect. These results indicate that the stable carbon and oxygen isotopes and growth increments are a good indicator of favourable growth conditions for Arctica islandica. Therefore, this study highlights that Arctica islandica is a suitable proxy for the North Sea and Irish Sea palaeoenvironment reconstructions. On this basis, future climate research can accurately depend on sclerochronological data to aid in understanding the patterns of anthropogenic climate change.

A 147cm sediment core from Lake DV09, northern Devon Island, Nunavut, Canada (75° 34’34”N, 89° 18’55”W) contains annually-laminated (varved) sediments, providing a 1600-year record of climate variability. A minerogenic lamina deposited during the annual thaw period and a thin deposit of organic matter deposited during the summer and through the winter, together form a clastic-organic couplet each year. The thinnest varves occur from AD800-1050, and the thickest from AD1100-1300, during the Medieval Warm Period. The relative sediment density is also highest during this period suggesting increased sediment transport energy. The coldest period of the Little Ice Age appears to be during the AD1600s. Varve widths over the past century indicate climate warming in the region. / This research program was funded by the Natural Sciences and Engineering Research Council of Canada (NSERC), and the Canadian Foundation for Climate and Atmospheric Sciences (CFCAS). A tuition bursary from Ultramar Inc. also helped in making this research possible. Logistical support was provided by the Polar Continental Shelf Project (PCSP Contribution number 04508).

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No / There is a series of common assumptions about prehistoric sex, associated with the prejudice that it must have been more natural because it happened closer to our evolutionary origins. The development of primate studies reveals a high degree of social variation between and within primate species, along with evidence for the practice of non-reproductive sex both recreationally and for expressing dominance relations. Yet, hypotheses about the behavior of human ancestors and early modern humans have been hampered by a lack of an integrated methodology. Although there is no single trajectory for either the elaboration or restriction of sexual behaviors after the emergence of culture, I argue here that it is possible to identify key turning points with more or less universal validity. These points include the reasons for and implications of brain size increase at the time of the emergence of genus Homo, the crystallization of impersonal gender by mid-Upper Paleolithic Ice Age societies, the early development of systems of control over both fertility and the projection and alteration of sexual identity, and the inferred emergence of homonegativity in early, reproduction-oriented farming societies. Further, archaeological data allows naturalist assumptions to be effectively refuted.

Symbiosis brought important evolutionary novelties to life on Earth. Lichens, the symbiotic entities formed by fungi, photosynthetic organisms and bacteria, represent an example of a successful adaptation in surviving hostile environments. Yet many aspects of the lichen symbiosis remain unexplored. This thesis aims at bringing insights into lichen biology and the importance of symbiosis in adaptation. I am using as model system a successful colonizer of tundra and alpine environments, the worm lichens Thamnolia, which seem to only reproduce vegetatively through symbiotic propagules. When the genetic architecture of the mating locus of the symbiotic fungal partner was analyzed with genomic and transcriptomic data, a sexual self-incompatible life style was revealed. However, a screen of the mating types ratios across natural populations detected only one of the mating types, suggesting that Thamnolia has no potential for sexual reproduction because of lack of mating partners. Genetic data based on molecular markers revealed the existence of three morphologically cryptic Thamnolia lineages. One lineage had a clear recombination structure and was found in the tundra region of Siberia, shorelines of Scandinavia, and Aleutian Islands. The other lineage was allopatric with the previous, and was highly clonal; only two haplotypes were found across the alpine region of central and southeastern Europe. However, the third lineage was sympatric with the other two, had a worldwide distribution, and although highly clonal, showed a recombinant population structure. Our data could not reveal whether the signs of recombination resulted from rare recombination events due to the extreme low frequency of the other mating type or ancestral variation before the loss of sexual reproduction. However, investigation of Thamnolia’s green algal population showed that in different localities, different algal genotypes were associated with the same fungal genotype. Furthermore, data suggest that Thamnolia carried several algal genotypes within its thalli and shared them with other distantly related but ecologically similar fungal species.

This thesis investigates how variations in the oxygen isotopic composition of lake waters in northern Fennoscandia are recorded in lake sediment archives, especially diatoms, and how these variations can be used to infer past changes in climate and atmospheric circulation. Results from analyses of the oxygen isotopic composition of lake water samples (δ18Olakew) collected between 2001 and 2006 show that δ18O of northern Fennoscandian lakes is mainly controlled by the isotopic composition of the precipitation (δ18Op). Changes in local δ18Op depend on variations in ambient air temperature and changes in atmospheric circulation that lead to changes in moisture source, vapour transport efficiency, or winter to summer precipitation distribution. This study demonstrates that the amount of isotopic variation in lake water δ18O is determined by a combination of the original δ18Olakew, the amount and timing of the snowmelt, the amount of seasonally specific precipitation and groundwater, any evaporation effects, and lake water residence time. The fact that the same isotope shifts have been detected in various δ18Olakew proxies, derived from hydrologically different lakes, suggests that these records reflect regional atmospheric circulation changes. The results indicate that diatom biogenic silica isotope (δ18Odiatom) records can provide important information about changes in atmospheric circulation that can help explain temperature and precipitation changes during the Holocene. The reconstructed long-term Holocene decreasing δ18Op trend was likely forced by a shift from strong zonal westerly airflow (relatively high δ18Op) in the early Holocene to a more meridional flow pattern (relatively low δ18Op). The large δ18Olakew depletion recorded in the δ18O records around ca. 500 cal yr BP (AD 1450) may be due to a shift to more intense meridional airflow over northern Fennoscandia resulting in an increasing proportion of winter precipitation from the north or southeast. This climate shift probably marks the onset of the so-called Little Ice Age in this region. / At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 1: In press. Paper 2: Submitted. Paper 5: In progress.

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As the world faces the effects of rapid climate change it is essential that we understand how the Earth’s climate has evolved in recent geological history. The island of Jersey harbours loess deposits which hold clues to recent climate change which have not been extensively studied. In this study, grain size analysis was employed on loess from the site of La Motte in order to shed light on climate change in the region during the last glacial. Several metrics such as the ratio of coarse to fine silt, the rate of deposition and the formation of ancient soils was used to assess the climate conditions during the time of deposition. The results show that the loess of La Motte is coarse compared to typical European loess but similar in grain size distribution to other sites along the English Channel, with relatively high amounts of sand and low amounts of clay. The relative coarseness of the deposits suggests a cold and dry depositional environment as well as proximity to the source of sediment and high wind intensities during the time of deposition. The fraction of coarse silt to fine silt, along with the rate of deposition and the occurrence of soil formation across the stratigraphy indicates that the climate slowly cooled from around 55 000 to 24 000 years ago when a sudden cooling event took place, then warmed until about 16 800 years ago where the climate suddenly cooled again. The timing of these rapid climatic shifts appears to correspond with Heinrich events 1 and 2 when large masses of ice were discharged from continental ice sheets. / Medan världen står inför effekterna av snabba klimatförändringar är det väsentligt att vi förstår hur jordens klimat har utvecklats under den senaste geologiska tiden. Ön Jersey hyser lössavlagringar som håller ledtrådar om den senaste tidens klimatförändringar och som inte studerats har studerats omfattande. I denna studie användes kornstorleksanalys på löss från lokaliteten La Motte med avsikt att utöka kunskapen om klimatförändringar i regionen under den senaste nedisningen. Ett flertal mått så som förhållandet mellan grov och fin silt, depositionshastigheten och bildningen av gamla jordar användes för att bedöma klimatförhållandena under avsättningstiden. Resultatet visar att La Mottes löss är grov jämfört med europeiska lössjordar, men i kornstorleksdistribution lik med andra lokaliteter längs med Engelska kanalen med relativt höga mängder sand och låga mängder lera. Avlagringarnas relativa grovhet tyder på en kall och torr avsättningsmiljö såväl som närhet till sedimentkällan och höga vindstyrkor under avsättningstiden. Förhållandet mellan grov och fin silt, tillsammans med depositionshastigheten och förekomsten av jordbildning genom stratigrafin tyder på att klimatet långsamt kyldes ner från omkring 55 000 år sedan till 24 000 år sedan då en plötslig nedkylning skedde. Klimatet värmdes sedan upp fram till omkring 16 800 år sedan då klimatet plötsligt kyldes ner igen. Tidpunkterna då dessa snabba avkylningar skedde verkar korrespondera med Heinrich-händelserna 1 och 2 då stora ismassor lossades från kontinentala istäcken ut i Nordatlanten.