Trees provide numerous benefits crucial to urban environments, yet poor growing conditions often prevent trees from reaching their genetic potential for growth, longevity, and ecosystem function. To overcome these limitations, greater understanding of tree growth in the urban environment is needed. The goal of this research project was therefore to characterize a broad suite of soil characteristics associated with urban tree plantings and evaluate their suitability for modeling physical dimensions and growth rates of urban trees. A series of observational studies and experiments was conducted on urban soils inhabited by two tree species (Zelkova serrata (Thunb.) Mikano and Quercus phellos L.) in Washington, DC and one tree species (Quercus virginiana Mill.) in Jacksonville, FL – two major metropolitan areas of the eastern United States with contrasting climate and soils. Characterization of urban soil attributes within cities revealed low variability for some properties (soil texture, pH, and certain plant nutrients with coefficients of variation (CV) below 0.5), but high variability (CV>1.0) for others (nitrate, ammonium, copper, and zinc). This is dependent on the location. These findings suggest that tree planting site evaluations may not require measurements for all soil properties and that representative sampling may be sufficient to accurately characterize most soil properties within a city. Field assessment of urban tree soils also revealed that conventional measures of soil compaction are difficult to obtain due to obstructions by roots and other foreign objects. To address the critical need for efficient and reliable assessment of soil compaction around urban trees, an experiment was conducted to develop bulk density estimation models for four common soil texture classes using soil strength and soil moisture as predictor variables. These models provided medium (0.42) to high (0.85) coefficients of determination when volumetric water content (VWC) was log transformed, demonstrating that measurements of soil texture, strength, and moisture can provide rapid, reliable assessment of soil compaction. Tree growth modeling focused on three response variables: canopy projection (CP), canopy volume (CV), and peak-increment-area age (PIA). To calculate PIA, tree-ring analysis was used to determine the age at which maximal trunk diameter growth occurred between transplanting and time of sampling. Because Q. virginiana has difficult-to-distinguish growth rings, an intensive tree-ring analysis of cores collected from these trees was conducted. The analysis revealed interseries correlation coefficients of up to 0.66, demonstrating that Q. virginiana can be aged with fairly high confidence in an urban setting. Empirical models developed for all three tree species using the suite of soil and site variables explained 25% – 83% of the observed variability in tree physical dimensions and growth rates. Soil pH was found to be a significant predictor variable for the majority of growth models along with nutrients such as Fe, B, Mn, and Zn, which are also associated with soil alkalinity. Models for PIA possessed the highest coefficient of determination, suggesting that measurements of soil conditions can be used confidently to predict the age at which growth rate subsides in these species. CV and CP were not predicted as well by soil-related variables, presumably because above-ground constraints such as pruning and building encroachment can affect canopy size without necessarily affecting growth rate. Certain prediction models for all three species included predictor variables with counterintuitive influences on tree growth (e.g., negative influences of soil depth on Q. phellos and soil volume on Q. virginiana), suggesting that either these urban trees are responding to these variables in a novel manner or that variables unaccounted for in these models (perhaps related to urbanization or high vehicular traffic) are concomitantly influencing tree growth. / Ph. D.

The Tropical savannas of central Brazil (cerrado) and the Everglades wetland (Florida) ecosystems are ideal systems to study landscape spatial mosaics and their interactions. Both ecosystems show a variety of plant physiognomies distributed within small spatial scales and elevation gradients. Such variety of plant physiognomies provide an opportunity to investigate the roles of climate, topography, nutrient availability and water table dynamics as determinants of plant physiognomic distributions, and their role in shaping regional systems. South Florida Wetlands and the tropical savannas of central Brazil are examples of hydrologically-controlled ecosystems. In hydrologically-controlled ecosystems water sources, the availability of nutrients, and the patterns of water movement play important roles in determining vegetation structure and function. The main objective of this study was to understand ecosystem level processes that shape different physiognomies in two hydrologically-controlled ecosystems. I conducted field work at the IBGE ecological reserve, a field experimental station located in Brasilia, Brazil. I also worked at the Everglades National Park in an area located near the south entrance of the Park in Homestead, Florida. I carried out three interconnected studies investigating water and nutrient dynamics: (1) In a Brazilian savanna I manipulated levels of litter input and measured changes to soil properties, organic matter decomposition and tree growth. I found that changes in litter input affect soil physicochemical properties and soil biochemical processes. I also found that litter dynamics influence tree growth through their effects on soil physicochemical properties. (2) I also studied the effect of water table depth and its temporal variation on spatial patterns of vegetation distribution in the cerrado landscape. I monitored diurnal and seasonal changes in water table depth along two tree-density and topographic gradients. In addition, I measured woody species composition, growth rates of four tree species, litter production, soil nutrients, and nutrient resorption efficiency along those two gradients. I found that water table depth has an important role in determining the spatial distribution of cerrado physiognomies; it also affects tree growth, species composition and nutrient resorption efficiency. (3) In the Everglades I studied patterns of underground water uptake by two vegetation types. I monitored seasonal and diurnal changes in water table depth in a Hammock forest, in a stand dominated by the invasive woody species Schinus terebinthifolius, as well as the water level in an adjacent lake. I estimated stand level transpiration using two different approaches: with sap flow measurements and diurnal oscillations in water table levels. Then, I calculated the total quantity of groundwater withdrawn by evapotranspiration for the wet and dry seasons in the Hammocks and in the exotic invaded site and then compared the results. I found that water uptake by Everglades trees is well coupled to diurnal changes in water table depth and that the amount of water withdrawn from the groundwater was larger during the wet season than during the dry season. Finally, I detected hydrological feedbacks between different vegetation types and nearby bodies of water. Results of this study contributes to the current knowledge of ecosystem level processes in tropical and subtropical ecosystems where water circulation and water availability play a dominant role in shaping vegetation structure and function.

Summary In the Sudanian zone of West Africa like in the Borgou region of Benin, agricultural and livestock systems face more and more difficulties linked to climatic changes, the decrease in soil fertility and the degradation of natural grasslands. This leads to a decrease in agricultural yields and feeding resources for livestock. On another hand, the socio-economical conditions of the small-holders dont allow them to invest in mineral fertilisers to remedy to soil fertility decrease. The introduction of ley pastures in crop rotations may be an affordable solution to increase available feeds for livestock and to improve soil fertility. The first part of this work reviews the potentialities of various local and exotic grasses and legumes which could be cultivated in the Borgou region. The choice of these species take into consideration the pedo-climatic conditions of the region, their aptitude for production, their nutritive value and their contribution to soil fertility. In the second and third parts of the thesis, the results of experimental 3-year ley pastures using 4 grasses (Panicum maximum Jacq cv. C1, Andropogon gayanus Kunth with broad leaves, A. gayanus with narrow limbs and Pennisetum purpureum Schumach.) and 6 légumineuses (Aeschynomene histrix Poir, Stylosanthes fruticosa (Retz.) Alson, Centrosema pubescens Benth., Mucuna pruriens (L.) DC. var. utilis Wall. Ex Wight) Baker ex Burck, Cajanus cajan (L.) Millsp. and Leucaena leucocephala (Lam.) De Wit) and exploited without any supply in mineral fertilisers are described concerning their yields and nutritive values. The grasses productions varied between 2 200 and 4 600 forage units (UF, in French unites fourragères)/ha/year and between 170 and 450 kg digestible crude protein/ha/year according to species and years (P<0.05). The forage legumes produced between 1 400 and 4 800 kg dry matter (DM)/ha/year according to species and years (P<0.05) with crude protein contents ranging from 16 to 27 %. Except Na and Zn, the combination of these two forage groups can satisfy the ruminant needs in macro and micro-minerals. The fourth part of the manuscript presents a soil-fertility balance under the different ley pastures after 3 years of exploitation. At the end of this study, it appears that in the experimental conditions (cut-and-carry system without fertilisers) Andropogon gayanus with broad leaves and the ligneous legume species Cajanus cajan and Leucaena leucocephala ensure the highest DM yields and are the less exhausting for soil fertility due to deeper root systems enabling to bring up to the surface mineral elements from deep layers / Résumé Dans la région soudanienne de lAfrique de lOuest comme dans le Borgou au Bénin, les systèmes de production agricole et délevage connaissent de plus en plus de difficultés liées aux aléas climatiques, à lappauvrissement rapide des sols et à la dégradation des végétations naturelles. Il en résulte une baisse des rendements agricoles et des ressources alimentaires pour le bétail. Par ailleurs, les conditions socio-économiques des producteurs limitent leurs possibilités dachat d'engrais en vue de remédier à la baisse de fertilité des sols. Dans ce contexte lintroduction de jachères fourragères dans les rotations culturales constitue une solution pour augmenter les disponibilités alimentaires destinées au bétail et enrichir le sol. La première partie de ce travail présente un inventaire de graminées et de légumineuses locales ou exotiques qui peuvent être implantées dans le Borgou. Le choix de ces espèces tient compte des conditions pédo-climatiques de la région, de leur aptitude à la production, de leur valeur alimentaire et de leur contribution à la fertilité des sols. Dans les deuxième et troisième parties de la thèse, les résultats de jachères expérimentales de trois ans comprenant 4 graminées (Panicum maximum Jacq var. C1, Andropogon gayanus Kunth à larges feuilles, A. gayanus à fines feuilles et Pennisetum purpureum Schumach.) et 6 légumineuses (Aeschynomene histrix Poir, Stylosanthes fruticosa (Retz.) Alson, Centrosema pubescens Benth., Mucuna pruriens (L.) DC. var. utilis Wall. Ex Wight) Baker ex Burck, Cajanus cajan (L.) Millsp. et Leucaena leucocephala (Lam.) De Wit) et exploitées sans apports de fumure minérale sont décrits en ce qui concerne leurs rendements et leurs valeurs alimentaires. Les productions des graminées ont varié entre 2 200 et 4 600 unités fourragères/ha/an et 170 à 450 kg de protéines brutes digestibles/ha/an selon les espèces et les années (P<0,05). Les légumineuses ont produit entre 1 400 et 4 800 kg de matière sèche/ha/an selon les espèces et les années (P<0,05), avec des teneurs en protéines variant entre 16 et 27 %. A lexception du Na et du Zn, lutilisation combinée des fourrages de ces espèces peut satisfaire les besoins en minéraux et oligo-éléments des ruminants. La quatrième partie du manuscrit présente un bilan de fertilité des sols sous ces différentes cultures après 3 années de jachère. Au terme de cette étude, il apparaît que dans les conditions expérimentales (affouragement en vert sans fumure minérale) la graminée Andropogon gayanus à larges feuilles et les légumineuses arbustives Cajanus cajan et Leucaena leucocephala sont les plus productives et les moins épuisantes pour le sol, eu égard à leur enracinement profond, capable de ramener des éléments minéraux en surface.

This study reports forest structure, regeneration and the soil properties from unlogged and logged forest in the Bulungan Research Forest, Malinau District, East Kalimantan, Indonesia. Four sites were compared by using four 1-ha replicate plots in each of primary forest (PF), 5, 10 and 30-yr old logged forest (LF-5, LF-10, LF- 30). The tree species composition differ among forest types, as it was shown that the mean value of similarity indices for all pairs were 0.215 (for the Jaccard index) and 0.353 (for the Sorensen index). The low values for similarities among forest types were most probably caused by low numbers of species shared between each forest type. Both correlation values, r = 0.023 for Jaccard index and r = 0.031 for Sorensen index, showed no strong correlation between the similarity index (C) and the distance between forest types. This supports the use of a chronosequence approach. A total of 914 tree species with ³ 10 cm dbh were recorded from 223 genera and 65 families. There were no significant differences in mean species numbers (166 – 180/ha) among treatments. Mean density of species was lower in LF-5 and LF-10 (501/ha) than in PF or LF-30 (605/ha and 577/ha); similarly to mean basal area (LF-5, 28.5 m2/ha; LF-10, 32.6 m2/ha) vs. PF (45.8 m2/ha) and LF-30 (46.9 m2/ha). Dead wood on the forest floor was significantly higher in LF-10 (75 m3/ha) than in the other treatments. Seedlings (< 2 cm dbh) of 1,022 species were recorded from 408 genera and 111 families. The mean number of tree seedling species ranged between 170-206; the mean density of seedlings was about two-fold lower in LF-10 (2790/ha) than in the other treatments. Saplings (>2 – 9.9 cm dbh) of 802 species belonged to 241 genera and 65 families. There was a high variability in species richness across treatments (89 – 191/ha), but not in stem numbers. The Dipterocarpaceae family was dominant in all treatments, followed by the Euphorbiaceae. The soils were acidic, low in nutrients and had low to very low fertility. Both primary and logged forest areas are marginal or not suitable for sustained production of plantation crops. Logging caused soil compaction in LF-30. Although in terms of number of species and trees, amount of BA, number of saplings and seedlings LF-30 appeared to have satisfied prescriptions for a second harvest, ecologically the forest is far from mature. The Indonesian Selective Cutting and Replanting (TPTI) system may need to be revised to a 35 – 45 year cycle to ensure long-term forest productivity in terms of not only timber but other goods and ecosystem services, the value of which are never quantified in monetary terms, but can be higher than the timber revenue.

In den meisten Ländern Mitteleuropas gilt weniger als ein Prozent des verbleibenden Laubwaldes als ungestört und temperierte Wälder sind Herausforderungen wie Arteninvasion, Klimawandel und steigender Stickstoff(N)-Deposition ausgesetzt. In der Vergangenheit wurde gezeigt, dass hohe N-Einträge N-Limitierungen verringern, Phosphor(P)aufnahme behindern und P-Mängel in der Buche auslösen können. Die Artendiversität von Bäumen kann die Bestandsproduktivität durch die Prozesse Komplementarität und Facilitation (Wachstumserleichterung) erhöhen, wenn diese einen wachstumslimitierenden Nährstoff betreffen. Ein Schlüsselprozess im Nährstoffkreislauf ist der Weg von Nährstoffen durch die mikrobielle Biomasse während der Dekomposition. Es wurde gezeigt, dass die mikrobielle Biomasse um N bspw. mit Buchen und um P mit tropischen Moorpflanzen konkurriert. Die Buche ist eine sehr konkurrenzfähige Baumart in temperierten Waldökosystemen aber kann von der Eiche in trockenen Bereichen übertrumpft werden, während Hainbuche und Linde eine geringere Rolle spielen. Eichen erfahren jedoch in der jüngsten Vergangenheit in europäischen Wäldern einen Rückgang, der womöglich auf hohe N-Einträge zurückzuführen ist. Für diese Arbeit untersuchten wir die Nährstoff-, Konkurrenz- und strukturelle Dynamik eines unbewirtschafteten, sehr naturnahen Laubwaldes in Mitteldeutschland, der aus Buche (Fagus sylvatica), Eiche (Quercus petraea und Quercus robur), Hainbuche (Carpinus betulus) und Linde (Tilia cordata und Tilia platyphyllus) aufgebaut ist. Unsere Ziele waren (1) zu erforschen, ob Komplementarität und/oder Facilitation die Produktivität in diesem Waldökosystem erhöht, (2) festzustellen, ob es Konkurrenz um die Nährstoffe N, P und K zwischen Bäumen und mikrobieller Biomasse gibt und, (3) die Nachbarschaftsdynamik der genannten Baumarten zu untersuchen und herauszufinden, ob der Eichenrückgang mit hoher N-Deposition einhergeht. In Beständen einer Art sowie verschiedenen Mischbeständen aus je drei Arten ermittelten wir Biomasseproduktion und Nährstoffverfügbarkeit. Nährstoffnutzungseffizienzkurven (Nährstoffnutzungseffizienz = Biomasseproduktion pro verfügbare Nährstoffe) wurden genutzt um festzustellen, ob ein bestimmter Nährstoff das Baumwachstum limitiert. Die jährliche Netto-Nährstoffveränderung wurde in einer Laubbeutel-Studie als Differenz zwischen ursprünglichem und verbleibendem Nährstoffgehalt des sich zersetzenden Laubfalls nach einem Jahr kalkuliert. Die Nährstoffresorptionseffizienz berechneten wir über die Ermittlung der N-, P- und Kalium(K)-Konzentrationen in sonnenexponierten Blättern und im gefallenen Laub. Die Nachbarschaftsdynamik von Bäumen wurde über die Durchmesserverteilung, überirdische Holzbiomasse für jede Artenkombination sowie eine Polygon-Abschätzung von Wachstumsräumen erforscht. Zusätzlich wurde eine durchmesserbasierte nearest neighbor(nächster-Nachbar)-Analyse für Baumpaare durchgeführt. Ein Geographisches Informationssystem (GIS) wurde genutzt um Wachstumsraum-Polygone zu erstellen und nächste Nachbarn zu bestimmen. Auf Einzelbaum-Level, ermittelt durch einen Nachbarschaftsansatz, waren relative Wachstumsraten von Buchen im Einzelbestand geringer als in der Mischung mit Linde und Hainbuche während das Wachstum von Linde im Einzelbestand größer war als in Mischung mit Buche und Eiche. Die Nährstoffnutzungseffizienzkurve für Buche zeigte optimale P- und K-Nutzungseffizienz für die Art in Mischbeständen, während sie in Einzelbeständen  P- und K-limitiert war. Während die jährliche Netto-Nährstoffveränderung in sich zersetzendem Blattlaub die Verfügbarkeit von P und K im Boden beeinflusste, war dies für N nicht der Fall. Resorptionseffizienzen von N, P und K hingen negativ mit der jährlichen Netto-Nährstoffveränderung zusammen. In unserer Studie zur Nachbarschaftsdynamik von Bäumen fanden wir heraus, dass intraspezifische nearest neighbors gleiche Durchmesser aufwiesen und ihren Durchmesser gleichzeitig mit dem des Nachbarn vergrößerten. Im Gegensatz dazu waren die Durchmesser von interspezifischen nearest neighbors im Allgemeinen unterschiedlich und der Durchmesser des Nachbarn verringerte sich mit zunehmendem Durchmesser des Zielbaums. Eichen konnten ihren Wachstumsraum mit zunehmendem Durchmesser nicht vergrößern, aber dominierten ihre nearest neighbor über die Größe. Unsere Ergebnisse zeigten, dass im untersuchten Waldökosystem Nährstofflimitierungen artabhängig waren und dass die Nutzung von Nährstoffnutzungseffizienz und Nachbarschaftsansatz geeignete Mittel sind, den Einfluss einzelner Baumarten auf die Produktivität einer Art im Rein- und Mischbestand zu ermitteln – so wie die beobachtete Facilitation der Buche im Mischbestand. Diese Werkzeuge stellen eine wichtige Basis zur verbesserten Bewirtschaftung typischer temperierter Mischwälder dar. Wir schlussfolgerten weiterhin, dass Konkurrenz zwischen mikrobieller Biomasse und Bäumen für P und K hoch, aber für N weniger bedeutend war, was wahrscheinlich in hoher N-Deposition in diesem Waldökosystem begründet liegt, welche den internen N-Kreislauf entkoppelte. Die hohe N-Deposition trug wahrscheinlich auch zu geringer Verjüngung der Eiche bei, während ältere Eichen in unserem Untersuchungsgebiet im Wettbewerb um Licht erfolgreich waren. Die Bestandsstruktur war charakterisiert durch stärkere interspezifische verglichen mit intraspezifischer Konkurrenz. Daraus resultierend bildeten Reinbestände aus Buche, Eiche und Linde Klimaxbestände hoher Biomasse innerhalb eines sich verändernden, kleinskaligen Mosaiks verschiedener Artenzusammensetzungen. In Reaktion auf neue Bewirtschaftungsanforderungen des Globalen Wandels sind weiterführende Forschungen zu Nutzungseffizienz unterschiedlicher Ressourcen für Baumarten in verschiedenen Zusammensetzungen empfehlenswert.

The influence of tree species on forest soils has been the subject of study for at least a century. Of particular interest have been western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western redcedar (Thuja plicata Donn ex D. Don) – two of the most common tree species in coastal and southern British Columbia, but each with a different nutrient amplitude. It has generally been found that acid, mycogeneous Mor humus forms develop in hemlock stands, while less acid and more zoogenous Mormoder, Moder, or even Mull humus forms develop in redcedar stands. The objective of this study was to determine the influence of hemlock and redcedar, growing separately and together, on forest floor nutrient properties. The questions addressed were: (1) does each stand type have unique forest floor nutrient properties? and (2) can any forest floor nutrient property discriminate between stand types?

Les invasions biologiques sont une cause majeure de perte de biodiversité à l’échelle mondiale. L’impact direct des espèces exotiques envahissantes (EEE) sur la structure et la composition des communautés a été bien documenté. Par contre, leur impact sur le fonctionnement des écosystèmes a été comparativement moins étudié. Dans le présent travail, j’ai mesuré sur le terrain et en conditions expérimentales l’impact de 7 espèces de plantes exotiques particulièrement envahissantes en Europe sur les propriétés chimiques du sol, sur la productivité et sur le stock d’éléments minéraux dans la biomasse. Malgré la diversité des groupes fonctionnels considérés dans notre échantillon (des espèces annuelles aux arbres), des impacts récurrents ont pu être mis en évidence. Les EEE ont toujours une biomasse et une productivité supérieures à celles de la végétation qu’elles envahissent. Il en va de même pour le stock d’éléments minéraux dans la biomasse aérienne. En ce qui concerne le sol, les traits fonctionnels des espèces ne permettent pas de prédire la direction et l’amplitude des impacts. Par contre, ces impacts sur le sol se sont révélés partiellement prévisibles en fonction des conditions écologiques initiales (contexte pédologique). La disponibilité des cations et du P et la concentration en C et N organiques augmentent suite à l’invasion dans les sites initialement pauvres alors que ces mêmes paramètres baissent dans les sites plus eutrophes au départ. Les EEE entraînent, dans une certaine mesure, une homogénéisation des conditions écologiques dans les écosystèmes envahis. <p>Dans un deuxième temps, je me suis attaché à identifier les mécanismes de l’impact de Fallopia japonica sur le cycle de l’azote. Cette espèce, perenne rhizomateuse, adopte une gestion très conservatrice de l’azote en retransloquant jusqu’à 80 % de l’azote des tiges et des feuilles vers le système racinaire en automne avant l’abscission des feuilles. La litière restituée est, de ce fait, pauvre en azote. Ce paramètre, combiné à une teneur élevée en lignine, explique sa faible vitesse de décomposition par rapport à celle de la végétation non envahie. Une grande partie de l’azote de l’écosystème tourne donc en cycle quasi fermé entre les organes de réserve et les parties aériennes de la plante, tandis que l’azote restant est en grande partie bloqué dans la nécromasse de la plante et est donc non disponible pour les espèces concurrentes indigènes. Ce mécanisme explique, sans doute, en partie le succès invasif de l’espèce.<p><p>A l’avenir, les implications des impacts sur le sol en terme de restauration des communautés devront être étudiées. L’hypothèse d’une influence positive de ces impacts sur l’aptitude compétitive des espèces envahissantes devra être testée. Enfin, une étude plus fonctionnelle des impacts de ces espèces dans des sites contrastés devrait permettre d’identifier les mécanismes impliqués./Biological invasions are a major cause of biodiversity loss worldwide. The direct impact of alien invasive species (AIS) on community structure and composition has been well documented. On the other hand, their impact on ecosystem functioning has been comparatively less studied. In this work, I measured, on the field and in experimental conditions, the impact of 7 highly invasive alien plant species in Europe on soil chemical properties, biomass and aboveground nutrient stock. Despite the high diversity in considered functional groups (from annuals to trees), recurring impacts have been found. AIS had always higher biomass and aboveground nutrient stocks than invaded resident vegetation. Concerning soil, species functional traits did not allow us to predict impact direction and intensity. On the other hand, impacts on soil were partially predictable based on initial ecological conditions (pedologic context). Thus, cations and P availability and organic C and N concentrations increased in initially poor sites and decreased in eutrophic ones. AIS tend to homogenize soil properties across invaded landscapes.<p>In a second part, i tried to identify the mechanisms of the impacts of Fallopia japonica on N cycling. This perrennial rhizomatous species manage N in a very conservative way. About 80 % of aboveground N is translocated to rhizomes before leaves abscission. The litter is therefore poor in N. This parameter combined with a high lignin concentration explains its slow decomposition rate compared to that of uninvaded vegetation. A big part of the ecosystem N is engaged in a nearly closed cycle between above and belowground organs while the remaining N is blocked in the necromass and is thus not available for the indigenous competitors. This mechanism probably explains the invasive success of this species.<p><p>In the future, the implications of soil properties modifications in terms of community restoration should be studied. The hypothesis of a positive influence of these modifications on the competitive ability of AIS should be tested. At last, a functional study of these soil impacts in contrasted sites should allow us to identify the implied mechanisms.<p> / Doctorat en Sciences / info:eu-repo/semantics/nonPublished

The relations between environmental resources and biodiversity are crucial in the proper management and conservation of grasslands. Three nature reserves were chosen around the Gauteng Province, namely Roodeplaat Nature Reserve (RNR) in Tshwane, Suikerbosrand Nature Reserve (SNR) in Heidelberg, and Abe Bailey Nature Reserve (ANR) in Carletonville. We selected three sites within RNR, SNR and ANR, and paired these with adjacent sites in private farming/grazing areas adjacent to the nature reserves. At each site, species composition, species richness and plant diversity were determined with the use of 50m x 20m Modified-Whittaker plots (MWP), making a total of eighteen plots (MWP) at the nine paired sites. Two paired sites had high Shannon-Wiener Index (H’) average values at the adjacent grazing area as compared to the nature reserve area at ANR. Roodeplaat Nature Reserve (RNR) and Suikerbosrand Nature Reserve (SNR) had two paired sites with high H’ average values in the nature reserve as compared to the adjacent grazing area. The vegetation structure was similar for the three study locations, consisting of graminoids, herbs and isolated patches of shrubs. The species composition showed similarities between ANR and SNR sites, while RNR showed different species composition. SNR soils had the highest organic carbon (OC), total Carbon (C), total Nitrogen (N), Calcium (Ca), Potassium (K), Magnesium (Mg) and Sodium (Na) as compared to both ANR and RNR. Species richness had a significantly positive relationship with Organic Carbon and Total Nitrogen. Species diversity difference was detected between the nature reserves and adjacent grazing areas and the difference are likely due to a number of factors including soil properties, land disturbance and land use and management. More research is necessary to further understand the aspects impacting species richness, species diversity and species composition in grasslands. / School of Agriculture and Life Sciences / M. Sc.

Thesis (M.Sc. (Soil Science)) -- University of Limpopo, 2022 / In South Africa, land use planning has received limited attention in areas perceived as suitable for agricultural production. In the lack of reliable soil type and fertility status information, crop yields remain lower than the land’s potential, with subsequent land degradation. Despite this, studies that focused on land capability and soil suitability to date have not considered the spatial variability of the soil nutrients and factors influencing their variability. However, this information is key for site-specific soil management. Therefore, it is vital to link land capability and soi suitability with the spatial variability of soil nutrients as it opens opportunities for more rational management of the soil resources since soil nutrients directly affect crop growth and consequently yield. To address this issue, a study was conducted on a 12 ha banana plantation portion of the Makuleke farm. The main objectives of this study were to (1) survey, classify and characterise soils in order to derive and map land capability classes of Makuleke farm, (2) quantify the physical and chemical properties of the soils in order to derive and map the soil suitability of Makuleke farm for banana production, (3) assess the spatial variability and structure of soil nutrients across the Makuleke farm and (4) Identify the factors of control of the spatial variability of the soil nutrients across the Makuleke farm. To begin with, a field soil survey was conducted using transect walks complemented by auger observations to sub-divide the 12 ha banana plantation portion of the farm into varied soil mapping units. Thereafter, soil classification was done to group soils based on their morphological properties and pedological processes. During soil classification, a total of 12 representative profile pits (1.5 m × 1.5 m long × 2 m deep/limiting layer) were excavated, studied, described, and sampled. At each profile pit, three replicates samples were collected at 0 – 30 cm depth intervals giving rise to 36 bulk soil samples. From the gathered soil profile information, four soil units were thus delineated and identified across the 12 ha banana plantation. For soil fertility assessment, a grid sampling strategy at 50 × 50 m was adopted to collect the samples across the 12 ha banana plantation. A total of 27 composite samples were collected at the nodes of the grid, and thereafter bagged, labelled, and transported to the laboratory. In the laboratory, all collected samples were air-dried and sieved using a 2 mm sieve in preparation for soil physical and chemical properties analysis. The land capability assessment of Makuleke farm was done using the concepts and principles of the FAO framework for Land Evaluation (FAO, 1976), but adapted to South African conditions by Smith (2006). Soil suitability assessment was done using the FAO framework for Land Evaluation (FAO, 1976) coupled with the guidelines for rainfed agriculture (FAO, 1983) and the criteria proposed by Sys et al. (1993) and Naidu et al. (2006). To assess the spatial variability and structure of the soil nutrients across the farm, classical and geostatistical techniques were employed respectively. A correlation matrix was employed to identify key factors influencing the spatial variability of soil nutrients across the farm. For interpolation, ordinary kriging was used to generate soil nutrient spatial distribution maps. In this study, four soil forms were identified and classified as Hutton, Westleigh, Glenrosa, and Valsrivier, which are broadly distinguished as Lixisols, Plinthosols, Leptosols, and Cambisols. Land capability results revealed that 17% of the 12 ha portion of the farm has very high arable potential (I), 60% of the farm has medium arable potential (III), 6% has low arable potential (IV) and 17 % is non-arable (VI), which might explain the varied banana yields in the farm. Soil suitability analysis revealed that 12% of the 12 ha farm is highly suitable (S1), 34% is moderately suitable (S2), 38% is marginally suitable (S3) and 16% is permanently not suitable (N2) for banana production. The low arable and marginally suitable portion of the farm was under Valsrivier soils which were limited by its shallow depth, shallow rooting depth, acidic soil pH, low organic carbon (OC), and the fact that it was located on a steeper slope gradient. The non-arable and not suitable portion of the farm for banana production was under Glenrosa and it was limited by its location on a steep slope gradient and was characterised by shallow effective rooting depth, low OC, low clay content, and acidic soil pH. Classical statistical techniques revealed that phosphorus (P), potassium (K), calcium (Ca), zinc (Zn), manganese (Mn), and copper (Cu) content varied highly across the banana plantation, while magnesium (Mg) and total nitrogen (TN) varied moderately. In addition, the geostatistical analysis revealed that spatial dependency was weak (Ca, Cu, and TN), moderate (Mg and Zn), and strong (P, K, and Mn) for the different soil nutrients across the 12 ha banana plantation. Soil nutrients with strong spatial dependency have a good spatial structure and are easily manageable (in terms of fertilisation, liming, and irrigation) across the farm compared to the ones with weak spatial dependency which have a poor structure. This study also found that land attributes, which are soil type and topographic position were the main factors driving the spatial variability of the soil nutrients across the farm. In terms of soil type, soils such as Valsrivier and Glenrosa with 2:1 clay-type smectite were the ones that had nutrient content compared to soils with 1:1 clay-type kaolinite (e.g., Westleigh and Hutton). Higher nutrient contents were also observed in the footslope position compared to the middleslope of the farmland. Correlation analysis revealed that Mn was the key polyvalent cation influencing the spatial variability of P, K, and Zn. Soil pH and effective cation exchanges capacity (ECEC) were the key soil factors driving the spatial variability of Ca, while ECEC was the key factor affecting the spatial variability of Mg. Moreover, the spatial variability of soil Mn and Cu was driven by soil Cu and clay content, respectively. The kriged maps showed that P, Mg, Zn, and Mn were high in the northeast part and low in the northwest part of the farm. Similarly, K and Ca were low in the northwest part, but they were high in the south to the southwest part of the study area. Total nitrogen was high in the west part and low in the east-northeast part, while Cu was evenly distributed across the plantation. This study highlights the importance of prior land use planning (i.e., land capability and soil suitability) and fertility assessment for agricultural production. The research results obtained provide the actual reference state of the capability of the land for arable farming and soil suitability for banana production at Makuleke farm. Moreover, the research results provide the spatial variability and structure of the soil nutrients which have a greater impact on the growth and yield of bananas. The results obtained in this study will be useful for site-specific management of soil nutrients and other soil management practices (e.g., irrigation, fertilisation, liming, etc.), developing appropriate land use plans, and quantifying anthropogenic impacts on the soil system and thus improving land productivity. / National Research Foundation (NRF)

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