Semi-natural grasslands are one of the most threatened habitats in Scandinavia. The grassland in this study is situated in the northern part of the nature preserve of Haväng och Vitemölla strandbackar in the province of Skåne and consists of grey dunes and a pasture which partly consists of calcareous grassland and the vegetation type sand steppe which is very rare in Sweden. There are several threatened plants in the area and during the summer of 2017 their distribution was registered to be able to protect them during conservation management. The results showed that most of the area contained important species and the most important areas to protect were the sand steppe areas. This is probably due to the dry, sandy, calcareous and nutrient poor conditions in these areas which many of the threatened plants are connected to. Most of the threatened plants were found in abundance in the area but have a limited distribution in Sweden. The population of Eryngium maritimum however was very small and is threatened by extinction. The plants in the area are threatened by a long-term process of nutrient enrichment and continuous leakage of calcium which may lead to lower pH and a more nutrient rich soil which leads to a higher availability of phosphorous. This may lead to a change in species composition and loss of species richness. Continuous grazing, soil disturbance to expose lime-rich sand and also removal of litter to minimize the nutrient levels are needed to maintain favorable conditions in the area. However, the number of important species requires attention and careful planning, especially in the sand steppe areas. / Ängs-och hagmarker är några av de mest hotade och viktiga habitattyperna i Skandinavien. Gräsmarken som studerades i detta projekt ligger i norra delarna av naturreservatet Haväng och Vitemölla strandbackar i Skåne och består av gräsbevuxna sanddyner och en hage med inslag av kalkgräsmark och marktypen sandstäpp som är väldigt sällsynt i Sverige. I reservatet finns flera skyddsvärda kärlväxter och under sommaren 2017 kartlades deras utbredning för att bättre kunna skydda dem vid skötsel- och restaureringsåtgärder. Resultatet visade att större delen av området innehåller skyddsvärda arter och att de mest skyddsvärda områdena var de delar av området där det fanns inslag av sandstäpp. Detta beror med stor sannolikhet på att de flesta skyddsvärda arterna i området var knutna till de torra, sandiga, kalkhaltiga och näringsfattiga förhållanden som rådde där. De flesta skyddsvärda arter fanns i stora antal i det undersökta området men har en begränsad utbredning i Sverige. Populationen av Eryngium maritimum var dock så litet att det hotas av utrotning. I övrigt hotas växterna i området på längre sikt av ökad tillsättning av näringsämnen och långvarig urlakning av kalk. Detta kan leda till lägre pH och mer näringsrik mark och gör att den nuvarande artsammansättningen kan försvinna och ersättas. För att området ska fortsätta ha en gynnsam status kommer det krävas fortsatt bete, skapande av sandblottor för att röra upp kalkrik sand samt åtgärder så som avlägsnande av förna och ytskikt för att hålla nere näringshalten. Det stora antalet rödlistade och skyddsvärda arter i området gör dock att det krävs varsamhet och noggrann planering vid en åtgärd, framför allt i sandstäppsområdena.

The main theme of this thesis is the existential anxiety and the motive of death in the early prosaic texts of the Italian author Dino Buzzati. The first chapter is dedicated to the author's biography where mainly the early years of his literary career are emphasized. After this introductory session follows an overview of the main features of existentialism as well as its place in the context of 20th century European literature and philosophy. In the last section of the theoretical part of the thesis we comment on the possible analogies between the literary work of Dino Buzzati and those of Franz Kafka, which were noticed by the contemporary literary criticism. The analytical part of the thesis is dedicated to the literary analysis of the primary texts: the novel The Tartar Steppe and selected short stories of the collection The Seven Messengers. In the last section we draw conclusions on the literary style of Dino Buzzati based on the previous analysis of the literary texts.

Die semiaride Steppe der Inneren Mongolei (China) ist ein gefährdetes Ökosystem. Der Wandel vom traditionellen nomadischen Lebensstil hin zur konventionellen Landwirtschaft überlastet die Steppe und führt zu Degradierung und Desertifikation. Besonders die intensive Beweidung belastet die weiten Grasflächen und mindert deren natürliches Potential Kohlen-stoff (C) im Boden zu speichern. Um den Einfluss unterschiedlicher Beweidungsintensitäten auf die semiaride Steppe zu untersuchen, wurden im Rahmen des Projektes Matter fluxes in grasslands of Inner Mongolia as influenced by stocking rate (MAGIM) das Einzugsgebiet des Xilin Flusses in der Inneren Mongolei von 2004 bis 2009 untersucht. Dafür wurden u. a. meteorologische und Eddykovarianz-Messungen an definierten Standorten durchgeführt. Ziel dieser Messungen war es, die Unterschiede im Energiehaushalt und den Kohlendioxid- und Wasserflüssen (CO2- und H2O-Flüsse) für die dominanten Steppenarten und unter verschiedenen Beweidungsintensitäten zu erfassen. Die Schließung der Energiebilanz ergab eine Schließungslücke von 10 – 30% in Abhängigkeit der meteorologischen Bedingungen, wobei die Lücke unter feuchten Bedingungen kleiner ist. Die gemessenen CO2- und H2O-Flüsse sind klein im Vergleich zu Grasländern in den gemäßigten Zonen und reagieren sensitiv auf Veränderungen der Einflussfaktoren. Dabei ist die Evapotranspiration (ET) eng an den eingehenden Niederschlag (P) gekoppelt und über längere Zeiträume wie ein Jahr entspricht ET dem P (ET: 185,7 mm a-1 bis 242 mm a-1; P: 138 mm a-1 bis 332 mm a-1). Die Jahressummen für den Nettoökosystemaustausch (NEE) reichen von -10,7 g C m-2 a-1 (2005) bis -67,5 g C m-2 a-1 (2007) für die unbeweidete Steppe und charakterisieren diese als eine leichte Nettosenke für atmosphärisches CO2. Grundsätzlich zeigt die unbeweidete Steppe eine höherer C-Sequestrierung (maximale C-Sequestrierung im Mittel -0,06 g C m-2 s-1) als die beweidete Steppe (maximale C-Sequestrierung im Mittel -0,02 g C m-2 s-1). Die Messergebnisse zeigen, dass die Steppe unter trockenen Verhältnissen zur CO2-Quelle wird, unter erhöhten Niederschlagsbedingungen zur CO2-Senke und die Beweidung die C-Sequestrierung des Ökosystems unter beiden Bedingungen einschränkt. Im Vergleich der beiden Steppenarten (Leymus chinensis und Stipa grandis) konnte für Leymus chinensis eine höhere Trockentoleranz beobachtet werden. Diese führt zu einer höheren C-Sequestrierung unter trockeneren Verhältnissen. Unabhängig von der Steppenart sind die wichtigsten Einflüsse auf das Ökosystem die Bodenfeuchte, die vom eingehenden P abhängt, die Temperatur (T) und die Beweidung. Diese Faktoren können dabei nicht unabhängig voneinander betrachtet werden. Der Einfluss durch die Beweidung beeinflusst das Ökosystem nachhaltig, wobei die Intensität und die Dauer (Jahre) der Beweidung entscheidend sind, da nicht nur die oberirdische Biomasse reduziert wird, sondern gleichzeitig die Bodeneigenschaften. Um die Sensitivität auf den CO2- und H2O-Austausch der semiariden Steppe über die Messungen hinaus abzuschätzen, wurden Simulationen mit den Modellen BROOK90 und DAILYDAYCENT (DDC) durchgeführt. Beide Modelle konnten gut an die Bedingungen der semiariden Steppe angepasst werden, wobei die Übereinstimmung zwischen der gemessenen und modellierten ET für BROOK90 besser war (r2 = 0,7) als für DDC (r2= 0,34). Beide Modelle konnten gut die Dynamik der ET-Messungen wiedergeben. Die Sensitivitätsanalyse hat gezeigt, dass die Beziehung zwischen P und ET entscheidend für das Ökosystem ist und sich Änderungen in der T nur zum Ende und Beginn der Vegetationsperiode auf den Wasseraustausch auswirken. DDC konnte sehr gut den gemessenen CO2-Austausch simulieren. Die Ergebnisse zeigen die Sensitivität gegenüber den klimatischen Faktoren T, P und der Beweidung. Die CO2-Flüsse werden durch hohe Beweidungsintensitäten so stark minimiert, dass andere Einflussfaktoren dahinter zurücktreten. Bei leichten Beweidungsintensitäten wirkt sich dagegen besonders der P auf die Austauschprozesse aus. Die DDC-Ergebnisse zeigen, dass unter den derzeitigen Bedingungen der bodenorganische Kohlenstoff (SOC) verringert wird, also C aus dem Boden freigesetzt wird. Auch unter unbeweideten Verhältnissen steigt der SOC nicht wieder auf das Ausgangsniveau (von vor der Beweidung) an. Die Ergebnisse zeigten, dass die C-Sequestrierung der Steppe nur erhöht werden kann, wenn der P steigt, die T in einem Optimumbereich (+/- 2°C) bleibt und die Beweidung minimiert wird. Die Messungen und Modellergebnisse zeigen, dass der Niederschlag der limitierende Faktor der semiariden Steppe ist. P bestimmt die Bodenfeuchte, diese wiederum beeinflusst das Pflanzenwachstum und somit den CO2- und H2O-Austausch der Pflanzen. Die Beweidung strapaziert das Ökosystem und reduziert dadurch die CO2- und H2O-Flüsse und verändert die Bodeneigenschaften nachhaltig. Unabhängig von der klimatischen Entwicklung, ist die derzeitige überwiegend hohe Beweidungsintensität der Steppe eine Belastung für das Ökosystem und schränkt das Pflanzenwachstum langfristig ein, was u. a. die Desertifikation begünstigt. / Semiarid grasslands in Inner Mongolia (China) are degrading. The change from the traditional Nomadic lifestyle to conventional agriculture stresses the semiarid grasslands and increases desertification. In particular, intense grazing of the semiarid grasslands reduces their potential of storing carbon (C) in the soil. In the project Matter fluxes in grasslands of Inner Mongolia as influenced by stocking rate (MAGIM) a team of scientists researched the catchment area of the Xilin River to investigate impacts of different grazing intensities on semiarid grasslands. Meteorological and eddy covariance measurements took place from 2004 to 2009. The aim of the measurements was to examine the energy balance and the exchange of the carbon dioxide (CO2) and water (H2O) fluxes of the dominant grasslands in Inner Mongolia under different grazing intensities. The energy balance could be closed by 70 – 90% depending on the driving factors. The energy balance shows a smaller gap for moist conditions. The CO2 und H2O fluxes in the study area are much smaller than in temperate grasslands and show a high sensitivity towards the driving factors. Evapotranspiration (ET) is closely connected to the precipitation (P) and over longer periods of a year or more, ET nearly matches P (ET: 185.7 mm a-1 to 242 mm a-1; P: 138 mm a-1 to 332 mm a-1). The annual net ecosystem exchange (NEE) of ungrazed grassland ranges from -10.7 g C m-2 a-1 (2005) to -67.5 g C m-2 a-1 (2007), which makes the grassland a small CO2-sink. Overall, ungrazed grassland shows higher C sequestration (averaged maximum -0.06 g C m-2 s-1) than grazed grassland (averaged maximum -0.02 g C m-2 s-1). The measurements show the semiarid grassland as a CO2-source under dry conditions and as a CO2-sink under moist conditions, while grazing decreases the C sequestration for both climatic conditions. A comparison of the two dominant steppe types (Leymus chinensis and Stipa grandis) showed a higher tolerance for Leymus chinensis under dry conditions, which resulted in higher C sequestration for this vegetation. Besides the steppe type, the main driving factors are P, temperature (T) and grazing. These factors can not only be considered in isolation, but cross correlation needs to get considered as well. Grazing affects the sustainability of the ecosystem, with an increasing impact due to grazing intensity and duration (years). The impact of grazing influences the vegetation directly and shows indirect impacts for the soil properties. Simulations with the models BROOK90 and DAILYDAYCENT (DDC) enable a sensitivity analysis of CO2 and H2O fluxes of the semiarid grassland. Both models performed well, but BROOK90 showed a better fit to observed ET (r2 = 0.7) than DDC (r2= 0.34). Both models simulated the dynamics of the measured ET well. The sensitivity analysis showed a close relationship between P und ET and a smaller impact on ET due to a change in T. DDC performs well in the simulation of CO2 exchange dynamics of the semiarid grassland. The results show for high grazing intensities a decreasing influence of the other driving factors. A change in P has an influence on CO2 and H2O fluxes under low grazing intensities. The results also show a decrease in soil organic carbon (SOC) as grazing intensity increases (under current climatic conditions). An increase in SOC could only be achieved under an increase in P, an optimum T and low grazing intensities. The measurements and results of the simulations indentify P as the main driving factor controlling the CO2 and H2O fluxes in the semiarid grassland. P influences the soil moisture and this influences plant growth, which governs the CO2 and H2O exchange of the vegetation. Grazing decreases the CO2 and H2O exchange and affects the soil properties in the long term. Besides climate change, the current high grazing intensities of the semiarid grassland have a negative impact on the ecosystem, decreasing plant growth and increasing desertification.

No description available.

Land-use legacies can affect landscapes for decades to millennia. A long history of shrub management exists in the sagebrush steppe of the Intermountain West where shrub-removal treatments, a type of managed disturbance, have been implemented for over 50 years to reduce sagebrush cover. The assumption behind managed disturbances is that they will increase forage for domestic livestock and improve wildlife habitat. However, the long-term effects of managed disturbance on plant community composition and diversity are not well understood. We investigated the legacy effects of three common types of managed disturbance (chemical, fire, and mechanical treatments) on plant community diversity and composition. We also examined sagebrush steppe resilience to managed disturbance. Based on management assumptions and resilience theory, we expected within-state phase shifts characterized by an initial reduction in biodiversity followed by a return to prior state conditions. We also expected changes in species proportions, characteristic of within-state shifts in state-and-transition models. We also expected an increase in non-native contribution to overall diversity. We found that plant communities experienced a fundamental shift in composition following disturbance, and responded in a flat linear fashion, giving no indication of return to prior community composition or diversity. As expected, we found post-disturbance increases in the number of non-native grass species present. However, native forb species made the largest contribution to altered diversity. Disturbance modified functional group composition, so contrary to our expectations, within-state changes did not occur as a result of disturbance. Our results indicated that sagebrush steppe plant communities are not resilient to chemical, fire, and mechanical treatments, and subsequent to managed disturbance, community composition tips over a threshold into an alternate stable state.

Seed germination during unhospitable environmental conditions can be a major barrier to direct seeding efforts in dryland systems. In the sagebrush steppe, Artemisia tridentata Nutt. ssp. wyomingensis and Artemisia arbuscula are important shrub species that are being used in restoration, but seeding success is highly sporadic due to inter-annual and intra-seasonal weather variability. Altering and expanding the period of germination, as a form of bet-hedging, may improve plant establishment. Our objective was to determine if we could expand the period of germination using plant growth regulators (PGRs) applied in a conglomerated seed coating treatment. In a laboratory study, the seed was either left untreated, conglomerated separately with two concentrations of a germination inhibitor, abscisic acid (ABA), or with two different germination promoters, gibberellic acid (GA3) and 1-Aminocyclopropane carboxylic acid (ACC), a precursor to ethylene. Seeds were incubated in a loam soil at five constant temperatures (5-25 C) for approximately three months. Results indicate that seed treatments with PGRs can delay or speed germination. The greatest response to the seed treatments was observed at 5 C. For example, at this temperature PGRs delayed the time for 25% of the seeds to germinate by a maximum of 35 and 21 d and decreased this time by 5 and 25 d for A. t. ssp. Wyomingensis and A. arbuscula, respectively. Field studies are needed to determine if the bet-hedging strategy developed in this study will increase the likelihood that some seeds will germinate during periods that are more favorable for plant establishment.

<p>This paper deals with archery in the Viking Age settlement of Birka and in particular the presence of Euro Asiatic, steppe nomadic archery equipment at the Birka Garrison and one Birka grave. The equipment contains for example closed quivers and a bow case. This paper also contains a discussion of archery battle techniques and tactics in Viking Age Birka and the implications of the above mentioned equipment to this discussion. The analysis insinuates the importance and status of archery in 10th century Birka.</p>

Durant des siècles, la steppe algérienne a été exploitée par des tribus nomades qui vivaient de l'élevage pastoral transhumant des petits ruminants. Fractionnée et déstabilisée par des politiques et des projets inappropriés pendant et après la colonisation française, cette exploitation collective et régulée des parcours a cédé la place à un mode d'exploitation familial concurrentiel. Pour répondre à une demande croissante de viande ovine, consécutive à un fort accroissement démographique, les éleveurs ont accru leurs troupeaux, étendu la céréaliculture fourragère motorisée et multiplié les achats de céréales fourragères. Surchargés, les parcours qui fournissaient l'essentiel de l'alimentation des animaux, ont été dégradés, la transhumance réduite et les petits éleveurs appauvris. Aujourd'hui, l'interdiction de la céréaliculture sur les terres fragiles et la mise en défens des pâturages dégradés sont plus que nécessaires. Elles ne seront cependant ni assez respectées pour protéger ou restaurer la fertilité de la steppe, ni suffisantes pour améliorer les conditions d'existence des agropasteurs, et elles le resteront tant que les ressources inexploitées en terres cultivables et en pâturages de la région n'auront pas été durablement mobilisées pour accroitre et réguler les disponibilités fourragères. Ainsi rendu possible, le développement durable de l'élevage agropastoral dans la steppe serait grandement facilité par l'élaboration et la réalisation participatives d'un ensemble coordonné de programmes et de projets, intégré aux stratégies de développement durable de la région et du pays.

Western juniper has rapidly expanded into sagebrush steppe communities in the Intermountain West during the past 120 years. This expansion has occurred across a wide range of soil types and topographic positions. These plant communities, however, are typically treated in current peer-reviewed literature generically. The focus of this research is to investigate watershed level response to Western juniper encroachment at multiple topographic positions. Data collected from plots used to measure vegetation, soil moisture, and infiltration rates show that intercanopy sites within encroached Western juniper communities generally exhibit a significant decrease in intercanopy plant density and cover, decreased infiltration rates, increased water sediment content, and lower soil moisture content. High-resolution remotely sensed imagery and Geographic Information Systems were used with these plot level measurements to characterize and model the landscape-scale response for both biotic and abiotic components of a Western juniper encroached ecosystem. These data and their analyses included an inventory of plant density, plant cover, bare ground, gap distance and cover, a plant community classification of intercanopy patches and juniper canopy cover, soil moisture estimation, solar insulation prediction, slope and aspect. From these data, models were built that accurately predicted shrub density and shrub cover throughout the watershed study area, differentiated by aspect. We propose a new model of process-based plant community dynamics associated with current state-and-transition theory. This model is developed from field measurements and spatially explicit information that characterize the relationship between the matrix mountain big sagebrush plant community and intercanopy plant community patterns occurring within a Western juniper dominated woodland at a landscape scale. Model parameters (states, transitions, and thresholds) are developed based on differences in shrub density and cover, steady-state infiltration rates, water sediment content, and percent bare ground in response to juniper competition and topographic position. Results from both analysis of variance and multivariate hierarchical cluster analysis indicate that states, transitions, and thresholds can be accurately predicted for intercanopy areas occurring within the study area. In theory, this model and the GIS-based layers produced from this research can be used together to predict states, transitions, and thresholds for any location within the extent of the study area. This is a valuable tool for assessing sites at risk and those that have exceeded the ability to self-repair. / Graduation date: 2005

This paper deals with archery in the Viking Age settlement of Birka and in particular the presence of Euro Asiatic, steppe nomadic archery equipment at the Birka Garrison and one Birka grave. The equipment contains for example closed quivers and a bow case. This paper also contains a discussion of archery battle techniques and tactics in Viking Age Birka and the implications of the above mentioned equipment to this discussion. The analysis insinuates the importance and status of archery in 10th century Birka.