A high-resolution record of organic-walled dinoflagellate cyst production in Guaymas Basin, Gulf of California (Mexico) reveals a complex paleoceanographic history over the last ~40 ka. Guaymas Basin is an excellent location to perform high resolution studies of changes in Late Quaternary climate and paleo-productivity because it is characterized by high primary productivity, high sedimentation rates, and low oxygen bottom waters. These factors contribute to the deposition and preservation of laminated sediments throughout large portions of the core MD02-2515. In this study we document dinoflagellate cyst production at a centennial to millennial scale throughout the Late Quaternary. Based on the cyst assemblages three major dinoflagellate cyst zones, with seven subzones were established. The most dominant dinoflagellate cyst taxa found throughout the core were Brigantedinium spp. and Operculodinium centrocarpum. Dansgaard-Oeschger events 5-8 are inferred in the dinoflagellate cyst records on the basis of increases in warm taxa, such as Spiniferites pachydermus. Preceding and during the Last Glacial Maximum cysts of Polykrikos cf. kofoidii increase in abundance, responding to oceanographic changes in the Gulf of California perhaps caused by a regression in sea-level. Other intervals of interest are the Younger Dryas where cooler conditions are not recorded, and the Holocene which is characterized by the consistent presence of warm water species Stelladinium reidii, Tuberculodinidum vancampoae, Bitectatodinium spongium and an increase in Quinquecuspis concreta. Changes in cyst assemblages, concentrations and species diversity, along with geochemical data reflect major millennial scale climatic and oceanographic changes. / Graduate

The Gulf of California in western Mexico is a prime example of a young passive margin that is currently undergoing the transition from continental rifting to seafloor spreading. With less than ~25 km of the width of the original continental surface area submerged, the northern Gulf Extensional Province represents a key area to assess the history of strain localisation during the early stages of continental extension. Geological mapping revealed that the basins and ranges of the Sierra San Felipe, located in the hanging wall of the Main Gulf Escarpment, are bounded to the east by an en-echelon array of left-stepping moderate- to low-angle normal faults that represent the next dominant set of normal faults from the break-away fault in direction of transport. Structural displacement estimates suggest up to ~4.5–9 km of broadly east-directed extension on the Las Cuevitas, Santa Rosa and Huatamote detachments. Fault kinematics suggest a transtensional stress regime with NE- to SE-directed extension and permutating vertical and N–S subhorizontal shortening. Clockwise vertical-axis block rotations and constrictional folding of the detachments were an integral part of the late Miocene to Pleistocene deformation history of the San Felipe fault array. This overall constrictional strain regime is indistinguishable from the present-day deformation in the Gulf Extensional Province and indicates that the fault array formed during a single phase of integrated transtensional shearing since rifting began in the late Miocene. / Apatite fission track (AFT) and (U-Th)/He results of Cretaceous crystalline basement samples from the Sierra San Felipe record a three-stage Cenozoic cooling history. Moderate cooling (~4–7ºC/m.y.) during late Paleocene to Eocene times is attributed to progressive down-wearing and bevelling of the ancestral Peninsular Ranges. Beginning at ~45–35 Ma, a period of tectonic quiescence with cooling rates of ≤1ºC/m.y. marks final unroofing of the basement and the development of a regional Oligocene to Miocene peneplain. Thermal modelling of samples from the footwall of the Las Cuevitas and Santa Rosa fault systems indicates that accelerated cooling began at ~9–8 Ma. This cooling pulse is attributed to tectonic denudation of the footwall and implies that faulting initiated synchronously on both detachments at ~9–8 Ma. Late Miocene deformation occurred distributed throughout the Sierra San Felipe, but started waning after the Pacific-North America plate boundary had localised into the Gulf of California by ~4.7 Ma / During a late Pliocene structural reorganisation in the northern Gulf, the locus of extension shifted from the Tiburón to the Delfín basins, thereby initiating strike-slip faulting on the Ballenas fracture zone, a transform fault located approximately 1.5–4.5 km offshore in central Baja California. This is consistent with low-temperature thermochronometric data from two horizontal transects perpendicular to the strike of the transform, which document a pronounced late Pliocene to Pleistocene heating event that is related to the structural and/or magmatic evolution of the transform fault. During reheating, maximum paleotemperatures reached >100–120ºC near the coast, but did not exceed ~60ºC some 5–8 km further inland. Highly non-systematic overprinting patterns are best explained by circulating hydrothermal fluids, which are most likely associated with magmatic leaking along the transform fault. / AFT and (U-Th)/He ages from a vertical profile collected on the Libertad escarpment, which forms part of the Main Gulf Escarpment in central Baja, pre-date Neogene extension and indicate that rift-related denudation was insufficient to expose samples from temperatures higher than the sensitivity zones of the two systems. One sample from the base of the escarpment however, records a middle to late Miocene hydrothermal overprint and suggests that extension in central Baja California likely initiated before ~10–8 Ma.

xv, 95 p. : ill. (some col.), maps (some col.) Plate 1. Geologic Map of the Sierra Bacha, Coastal Sonora, Mexico (1:30,000 scale) attached as a separate file. / The Gulf of California is an active rift basin formed by late Cenozoic dextral-oblique extension along the Pacific-North America plate boundary. Well exposed volcanic and sedimentary rocks in the Sierra Bacha, coastal Sonora, Mexico, preserve a history of proto-Gulf (late Miocene) deformation and offer insight into the structures and kinematics responsible for localization of the plate boundary and inception of the Gulf at about 6 Ma. Geologic mapping, fault kinematic analysis, and paleomagnetic data suggest that proto-Gulf deformation in the Sierra Bacha occurred primarily by ENE-WSW extension and that vertical-axis rotation related to dextral strain was minor. Lack of significant dextral shear supports an emerging model for proto-Gulf deformation in which dextral strain was not ubiquitous across Sonora but instead became localized during latest Miocene time in a narrow coastal shear zone that mechanically weakened the lithosphere and helped facilitate continental rupture. This thesis includes the "Geologic Map of the Sierra Bacha, Coastal Sonora, Mexico" as supplemental material. / Committee in charge: Dr. Rebecca J. Dorsey, Chairperson; Dr. Marli B. Miller, Member; Dr. Ray J. Weldon II, Member

The fishery for octopus in Northwest Mexico has increased to over 2,000 tons annually, but to date the specific composition of the catch has been ignored. With at least three main species targeted by artisanal fisheries in the region with distinct life histories, the lack of basic biological information about the distribution, metapopulation size and structure of each species could impede effective fisheries management to avoid overexploitation. We tested if different life histories of three species of octopus could help predict observed patterns of genetic diversity, population dynamics, structure and connectivity and how this information could be relevant to the sustainable management of the fishery. We sequenced two mitochondrial genes and genotyped seven nuclear microsatellite loci to identify the distribution of each species in 20 locations from the Gulf of California and the west coast of the Baja California peninsula. We tested five hypotheses derived from population genetic theory based on differences in the fecundity and dispersal potential for each species. We discovered that Octopus bimaculoides with low fecundity and direct development (without a planktonic phase) had lower average effective population size and genetic diversity, but higher levels of kinship, population structure, and richness of private alleles, than the other two species. These features indicated limited dispersal and high local recruitment. In contrast, O. bimaculatus and O. hubbsorum with higher fecundity and planktonic phase as paralarvae had higher effective population size and genetic diversity, and overall lower kinship and population structure than O. bimaculoides. These observations supported higher levels of gene flow over a larger geographical scale. O. bimaculatus with the longest planktonic paralarval duration and therefore larger dispersal potential had differences in the calculated parameters possibly associated with increased connectivity. We propose O. bimaculoides is more susceptible to over exploitation of small, isolated populations and could have longer recovery times than the other two species. This species may benefit from distinct fishery management within each local population. O. bimaculatus and O. hubbsorum may benefit from fishery management that takes into account metapopulation structure over larger geographic scales and the directionality and magnitude of larval dispersal driven by ocean currents and population connectivity among individuals of each locality. The distribution of each species and variations in their reproductive phenology is also important to consider when establishing marine reserves or seasonal fishing closures.

One of the biggest questions surrounding common-pool natural resources (CPRs) lies in understanding the circumstances which increase the likelihood of sustainable use and those that lead to resource degradation. Small-scale fisheries are an example of a CPR that has proven difficult to manage sustainably. I use social network analysis methods to examine the social connectivity of small-scale fishing communities and the association of network structures with collaborative behavior of small-scale fisheries in the Northern Gulf of California, Mexico.I found considerable connectivity of communities via kinship ties of small-scale fishers, both within the region and to other areas in Mexico. Fisher kinship relationships are important mechanisms for information transfer. Identifying communities in the network that are most likely to share information with other communities allows managers to develop more effective and efficient education, outreach, and enforcement efforts.Communities are also connected by their use of the same fishing zones and Marine Protected Areas (MPAs). My results provide suggestions for dividing communities based on common use of fishing areas and MPAs. This may help fishers and managers to develop, implement, and enforce boundary rules that will facilitate regional management of small-scale fisheries. My results provided mixed evidence for the role of social structure in impacting positive outcomes for fisher' ability to collaborate and organize. A wide range of factors affect the emergence of institutions for CPR management. Similarly, finding a common network structure that can accurately predict sustainable use of CPRs is unlikely. Knowing how people are connected and the ways in which information about CPR resources moves through (or is hindered from moving through) a network can improve manager's ability to develop more effective strategies and actions. Adding social networks into the CPR management toolbox provides a mechanism by which those working in management and conservation can incorporate social structure into management activities.An understanding of the social networks that connect communities and the potential pathways for information transfer, combined with a system of enforceable rules and policies and effective outreach methods and materials, may help managers and resource users more effectively and sustainably manage CPRs in the long term.

abstract: Niche variation among sexes and life stages within a population has been documented in many species, yet few studies have investigated niche variation within demographic groups or across ecological contexts. We examined the extent to which pregnant California sea lions (Zalophus californianus) at each of three breeding colonies target alternative prey resources and habitats. The three colonies are distributed across distinct regions of the Gulf of California, Mexico and have divergent population dynamics. We compared the nature of niche variation among colonies and investigated the fitness consequences of different foraging strategies within each colony. We analyzed the δ13C and δ15N values from fur collected from 206 suckling pups to characterize relative maternal foraging locations (δ13C) and trophic levels (δ15N) during the metabolically demanding late stages of gestation and lactation that occur simultaneously in California sea lions. The δ13C and δ15N values were regressed against pup body condition index values to compare the relative individual-level fitness benefits of different maternal foraging strategies. We found that the nature and extent of niche variation differed among colonies. Niche variation was most pronounced at the two largest colonies that appear to experience the highest levels of intraspecific competition and the variation was consistent with habitat features. One colony (Granito) displayed two distinct foraging groups with indistinguishable median pup body condition values, whereas the second (San Jorge) exhibited continuous niche variation and pup body condition varied in relation to maternal foraging location and trophic level, suggesting disparities among alternative foraging strategies. For the smallest colony (Los Islotes), females occupy similar niches with a few outliers. Body condition values of pups at this colony were most variable, but did not vary with maternal foraging strategy. Our results provide evidence for intrapopulation niche variation among demographically similar individuals during a period of high metabolic stress and reproductive importance. This work suggests possible fitness benefits conferred by alternative foraging strategies, and calls into question the common assumption that members of a population are ecologically equivalent. Future research aimed at understanding animal foraging strategies should consider the nature and extent of niche variation in the context of local ecological conditions. / Dissertation/Thesis / M.S. Biology 2013

Le Golfe de Californie (GOC) est un exemple de rift très oblique, au stade d'accrétion depuis 3,6 Ma au sud. La déformation continentale débute au Miocène dans un contexte arrière-arc, en relation avec la subduction de la plaque Pacifique sous la Plaque Nord-Américaine. L'objectif de cette thèse est de déterminer I'histoire de la déformation à I'origine de la rupture continentale ainsi que les phénomènes post-rupture. Cette étude utilise des données tectoniques, sismologiques et géomorphologiques sur la marge Ouest du GOC en Basse Californie du Sud. Elles sont calées temporellement par des datations d'isotopes radiogéniques et cosmogéniques. Une histoire polyphasée de la déformation, essentiellement post-magmatique, est proposée en termes d'évolution des directions des failles et de déformations relativement à la cinématique des plaques. Je démontre dans cette zone que la déformation qui conduit à l'amincissement et à l'étirement lithosphérique est d'abord fortement oblique et devient transtensive et moins oblique relativement à la cinématique à partir de 7-8 Ma. La marge étudiée est intégrée de manière cohérente à I'ensemble des domaines déformés associés à la formation du GOC. On montre notamment que les marges du GOC sont diachrones et qu'elles se forment par migration vers I'Ouest de la déformation vers la zone de subduction, qui devient inactive vers 12 Ma. La dynamique post-breakup du GOC est interprétée en termes de mobilisation par fluage de la croûte inférieure en relation avec un évènement thermique lié à la rupture du slab. En conclusion, l'évolution du GOC ne peut s'expliquer par les modèles simples de rifting oblique et de formation de marges passives. / The Gulf of California (GOC) is an example of highly-oblique rift. Oceanic accretion started 3.6 Ma ago at its southern end. The earliest continental extension started during the Miocene, in a back-arc setting, in connection with the subduction of the Pacific plate (PAC) beneath North America (NAM). ln this study, I reconstitute the strain evolution along the proximal Baja California margin. For this, I used original tectonic and seismological data which I collected in Baja California Sur (BCS). Those data were time-constrained with absolute dating (radiogenic and cosmogenic isotopes). I outline that the main stretching and thinning of the Late Miocene-Pliocene Baja California margin was highly oblique regarding NAM-PAC kinematic vector, turning less oblique. By integrating the studied margin in the evolution of the GOC, it is proposed that the final break-up mechanism occurred within a broad semi-ductile right-lateral central shear-zone. lt is shown that the two GOC passive margins didn't form at the same time, crustal strain migrating westward during the Miocene in response to a probable retreat of the dying slab. The active post-breakup deformation in the proximal BCS is best interpreted in connection with an outward flow of the lower crust in a trend compatible with the margin shaping inherited from the major GOC-normal to GOC-oblique Miocene crustal deformation. This ductile flow would be enhanced by the heat input from the slab rupture. To conclude, the GOC evolution as an oblique rift system adjusts with no existing analogical or numerical model of strain field evolution of oblique rifting, a probable consequence to the complex dynamics in back-arc settings.