As pesquisas apresentadas nesta dissertação integram o projeto CNPq - PROANTAR 550352/02-3 \"Mudanças paleoclimáticas na Antártica durante o Cenozóico: o registro geológico terrestre\", que estuda os depósitos cenozóicos da ilha Rei George em busca elucidação do histórico ambiental e climático desta região antártica. A evolução dos padrões de circulação marinha e atmosférica no Hemisfério Sul ocorreu em resposta ao isolamento geográfico e térmico da Antártica, resultado de sua separação da Austrália, no limite Eoceno/Oligoceno, e da América do Sul, no final do Oligoceno. Sob este aspecto, o estudo de organismos fósseis registrados nos depósitos cenozóicos da Antártica contribui para o entendimento das evoluções biológicas e ambientais ocorridas concomitantemente às mudanças paleogeográficas, oceanográficas e climáticas na região ao longo do Cenozóico. Frente à dificuldade de acesso, demanda logística e extensa cobertura de gelo, apenas uma pequena porção do registro geológico da Antártica está acessível para pesquisa. Afloramentos da ilha Rei George registram as mudanças climáticas e ambientais ocorridas do Oligoceno ao Mioceno, incluindo evidências do primeiro evento de glaciação perene no oeste do continente (Oligoceno). A despeito da abundância de fósseis nos estratos cenozóicos da ilha, são poucos os trabalhos taxonômicos com descrição sistemática detalhada de bivalves fósseis. O primeiro módulo do presente estudo apresenta a descrição taxonômica de invertebrados de depósitos cenozóicos aflorantes em duas localidades da ilha Rei George, Antártica ocidental. Da Formação Cape Melville (Mioceno), península Melville, foram descritos sete táxons de bivalves, incluindo seis espécies novas. Da Formação Polonez Cove (Oligoceno), Pico Vauréal, uma região previamente inexplorada paleontologicamente, foram descritos sete táxons de invertebrados (bivalves, braquiópodes, tubos de serpulídeos, briozoários e fragmentos de equinodermes), incluindo duas espécies novas. O segundo módulo corresponde à reunião dos gêneros de bivalves registrados em depósitos cenozóicos da Antártica. A análise do registro apontou para o conhecimento bastante incipiente sobre a diversidade de bivalves antárticos ao longo do Cenozóico. Além disso, a comparação entre gêneros de bivalves cenozóicos registrados na Antártica e Nova Zelândia revelou que a maior parte dos gêneros compartilhados está registrada em depósitos eocênicos, o que suporta o isolamento geográfico da Antártica e a redução do intercâmbio faunístico entre a Antártica e regiões periféricas após o Oligoceno. A análise do registro sugeriu um evento de dispersão intenso durante o Eoceno, e pequenos pulsos de dispersão após o Oligoceno. O padrão de distribuição dos táxons concorda parcialmente com as reconstituições de paleocorrentes disponíveis na literatura. A dispersão durante o Eoceno teria ocorrido da Antártica para a Nova Zelândia na direção do Atlântico para o Pacífico. Este evento de dispersão concorda com a hipótese de existência de conexões marinhas de plataforma rasa entre o oeste e o leste da Antártica (\"Passagem de Shackleton\") e da província Weddeliana do final do Cretáceo ao Eoceno. Os eventos de dispersão pósoligocênicos teriam ocorrido durante e após o estabelecimento da Corrente Circum-Antártica, não mais pela \"Passagem de Shackleton\", mas margeando a Antártica pelas bordas ocidental atlântica e oriental em direção à Nova Zelândia. A análise do registro dos bivalves cenozóicos da Antártica também concorda com a hipótese de glaciação perene a partir do início do Oligoceno na região leste do continente, e na metade do Oligoceno na região oeste, com temperaturas mais amenas que as observadas atualmente. / The research presented in this dissertation comprised part of the CNPq - PROANTAR Project 550352/02-3 \"Mudanças paleoclimáticas na Antártica durante o Cenozóico: o registro geológico terrestre\", which studies Cenozoic deposits from King George Island in order to elucidate the environmental and climatic Cenozoic histories of this Antarctic region. Cenozoic evolution of marine and atmospheric circulation in the Southern Hemisphere occurred in response to the geographic and thermal isolation of Antarctica, which resulted from the separation of Antarctica from Australia, around Eocene/Oligocene boundary, and from South America, during the late Oligocene. Thus, study of fossil organisms from Antarctic Cenozoic deposits contributes to the understanding of biological and environmental evolutions that accompanied paleogeographic, oceanographic and climatic changes during the Cenozoic. As a result of the difficult access, logistic demand and extensive ice cover, only a small part of the Cenozoic Antarctic record is available for study. King George Island records climatic and environmental changes from the Oligocene to the Miocene, including evidence of the first full-scale glaciation (Oligocene) of West Antarctica. Despite the abundance of fossils in Cenozoic deposits of the island, taxonomic studies with detailed systematic descriptions of bivalves are very rare. The first section of this work consists of taxonomic descriptions of invertebrates from Cenozoic deposits cropping out in two localities of King George Island, West Antarctica. Seven taxa of bivalves, including six new species were described from the Cape Melville Formation (Miocene), at Melville Peninsula. Seven taxa of invertebrates (bivalves, brachiopods, serpulid tubes, bryozoans, and echinoderm fragments) were described from the Polonez Cove Formation (Oligocene), at Vauréal Peak, a site previously unexplored paleontologically. The second section presents the results of a survey of the Cenozoic fossil record of Antarctic bivalves. The analysis of the fossil record confirmed that the current knowledge about the Cenozoic diversity of the group is very scarce. Moreover, comparison of Cenozoic bivalve genera from Antarctica and New Zealand showed that the greatest number of shared taxa is recorded in Eocene deposits. This finding supports the geographic isolation of Antarctic and the drop in faunal interchange between Antarctica and periphery after the Oligocene. Analysis of the fossil record suggested an intensive dispersal event during the Eocene, and restricted pulses of dispersal from the Oligocene onwards. The distribution pattern of taxa provides partial support for available reconstructions of marine currents. Eocene dispersal would have occurred from Antarctica to New Zealand in Atlantic-Pacific direction. This dispersal event is consistent with the hypothesis of shallow marine connections between West and East Antarctica (\"Shackleton Seaway\"), as well of the existence of the Weddellian Province from the Late Cretaceous to the Eocene. Dispersal events following the Oligocene would have occurred during and after the establishment of the Circum-Antarctic Current, along the West-Atlantic and East margins of Antarctica towards New Zealand, and no longer through \"Shackleton Seaway\". These analyses also support the hypothesis of full-scale glaciation in West Antarctica from the early Oligocene onwards, and in East Antarctica since the mid-Oligocene, with warmer temperatures than today.

Apesar de negligenciados por muito tempo, os Pleurodira, uma das duas linhagens de Testudines (cágados, jabutis e tartarugas-marinhas), vem atraindo crescente atenção dos pesquisadores, com trabalhos que geralmente tratam da descrição de novas espécies (viventes ou extintas), de comportamentos ou hábitos alimentares, ou de análises filogenéticas dos sub-grupos de Pleurodira, como Chelidae ou Pelomedusoides, esta sub-dividida em duas linhagens com representates viventes: Pelomedusidae e Podocnemidae. Análises filogenéticas com táxons terminais de todos os principais grupos de Pleurodira existem apenas com dados moleculares. Assim, uma análise baseada em dados morfológicos que englobe táxons de todos os grandes clados do grupo, incluindo fósseis, faz-se necessária para melhor compreensão da história evolutiva do grupo. Nesta dissertação, descrevemos uma nova espécie de Bairdemys (Podocnemidae) do Mioceno médio da Venezuela e a incluímos em uma análise filogenética com ampla amostragem taxonômica, incluindo táxons de todas as linhagens de Pleurodira. Esta análise é baseada em caracteres morfológicos (novos ou previamente propostos) e a maior análise filogenética exclusiva do grupo já realizada, permitindo inferências sobre padrões gerais de diversificação e morfologia de Pleurodira. Ao mesmo tempo, o novo táxon descrito apresenta características morfológicas que, juntamente com aspectos geológicos de sua localidade tipo, permitem a inferência de que houve, durante o Cenozóico, uma irradiação marinha de Podocnemidae, que alcançou grande sucesso, principalmente no final do Oligoceno e começo do Mioceno / Though neglected for a long time, one of the two lineages of Testudines (turtles and tortoises) the Pleurodira has been attracting the researchers, whose articles usually deal with descriptions of new species (extant and extinct), behavior or feeding habits, or phylogenetic analyzes of one of the subgroups of Pleurodira, Chelidae or Pelomedusoides, the latter subdivided in two lineages with extant taxa: Pelomedusidae and Podocnemidae. There some phylogenetic analyzes with terminal taxa from the main groups of Pleurodira, but only with molecular data. Therefore, a morphological data analysis containing taxa from all the clades of this group, including fossils, is necessary to better understand the evolutionary history of this lineage. In this dissertation, I describe a new species of Bairdemys (Podocnemidae) from the middle Miocene of Venezuela, and I include it in a phylogenetic analysis with extensive taxonomic sampling, including taxa from all the lineages of Pleurodira. This analysis is based on morphological characters (new and previously proposed ones) and it is the largest exclusive phylogenetic analysis ever made for Pleurodira, allowing inferences about the general patterns of diversification and morphology of the group. At the same time, the new described táxon shows morphological characters that, along with geological aspects of its type locality, allow inferences about a marine wave of diversification of Podocnemidae during the Cenozoic, which achieved great success, specially by the end of Oligocene and beginning of Miocene

As pesquisas apresentadas nesta dissertação integram o projeto CNPq - PROANTAR 550352/02-3 \"Mudanças paleoclimáticas na Antártica durante o Cenozóico: o registro geológico terrestre\", que estuda os depósitos cenozóicos da ilha Rei George em busca elucidação do histórico ambiental e climático desta região antártica. A evolução dos padrões de circulação marinha e atmosférica no Hemisfério Sul ocorreu em resposta ao isolamento geográfico e térmico da Antártica, resultado de sua separação da Austrália, no limite Eoceno/Oligoceno, e da América do Sul, no final do Oligoceno. Sob este aspecto, o estudo de organismos fósseis registrados nos depósitos cenozóicos da Antártica contribui para o entendimento das evoluções biológicas e ambientais ocorridas concomitantemente às mudanças paleogeográficas, oceanográficas e climáticas na região ao longo do Cenozóico. Frente à dificuldade de acesso, demanda logística e extensa cobertura de gelo, apenas uma pequena porção do registro geológico da Antártica está acessível para pesquisa. Afloramentos da ilha Rei George registram as mudanças climáticas e ambientais ocorridas do Oligoceno ao Mioceno, incluindo evidências do primeiro evento de glaciação perene no oeste do continente (Oligoceno). A despeito da abundância de fósseis nos estratos cenozóicos da ilha, são poucos os trabalhos taxonômicos com descrição sistemática detalhada de bivalves fósseis. O primeiro módulo do presente estudo apresenta a descrição taxonômica de invertebrados de depósitos cenozóicos aflorantes em duas localidades da ilha Rei George, Antártica ocidental. Da Formação Cape Melville (Mioceno), península Melville, foram descritos sete táxons de bivalves, incluindo seis espécies novas. Da Formação Polonez Cove (Oligoceno), Pico Vauréal, uma região previamente inexplorada paleontologicamente, foram descritos sete táxons de invertebrados (bivalves, braquiópodes, tubos de serpulídeos, briozoários e fragmentos de equinodermes), incluindo duas espécies novas. O segundo módulo corresponde à reunião dos gêneros de bivalves registrados em depósitos cenozóicos da Antártica. A análise do registro apontou para o conhecimento bastante incipiente sobre a diversidade de bivalves antárticos ao longo do Cenozóico. Além disso, a comparação entre gêneros de bivalves cenozóicos registrados na Antártica e Nova Zelândia revelou que a maior parte dos gêneros compartilhados está registrada em depósitos eocênicos, o que suporta o isolamento geográfico da Antártica e a redução do intercâmbio faunístico entre a Antártica e regiões periféricas após o Oligoceno. A análise do registro sugeriu um evento de dispersão intenso durante o Eoceno, e pequenos pulsos de dispersão após o Oligoceno. O padrão de distribuição dos táxons concorda parcialmente com as reconstituições de paleocorrentes disponíveis na literatura. A dispersão durante o Eoceno teria ocorrido da Antártica para a Nova Zelândia na direção do Atlântico para o Pacífico. Este evento de dispersão concorda com a hipótese de existência de conexões marinhas de plataforma rasa entre o oeste e o leste da Antártica (\"Passagem de Shackleton\") e da província Weddeliana do final do Cretáceo ao Eoceno. Os eventos de dispersão pósoligocênicos teriam ocorrido durante e após o estabelecimento da Corrente Circum-Antártica, não mais pela \"Passagem de Shackleton\", mas margeando a Antártica pelas bordas ocidental atlântica e oriental em direção à Nova Zelândia. A análise do registro dos bivalves cenozóicos da Antártica também concorda com a hipótese de glaciação perene a partir do início do Oligoceno na região leste do continente, e na metade do Oligoceno na região oeste, com temperaturas mais amenas que as observadas atualmente. / The research presented in this dissertation comprised part of the CNPq - PROANTAR Project 550352/02-3 \"Mudanças paleoclimáticas na Antártica durante o Cenozóico: o registro geológico terrestre\", which studies Cenozoic deposits from King George Island in order to elucidate the environmental and climatic Cenozoic histories of this Antarctic region. Cenozoic evolution of marine and atmospheric circulation in the Southern Hemisphere occurred in response to the geographic and thermal isolation of Antarctica, which resulted from the separation of Antarctica from Australia, around Eocene/Oligocene boundary, and from South America, during the late Oligocene. Thus, study of fossil organisms from Antarctic Cenozoic deposits contributes to the understanding of biological and environmental evolutions that accompanied paleogeographic, oceanographic and climatic changes during the Cenozoic. As a result of the difficult access, logistic demand and extensive ice cover, only a small part of the Cenozoic Antarctic record is available for study. King George Island records climatic and environmental changes from the Oligocene to the Miocene, including evidence of the first full-scale glaciation (Oligocene) of West Antarctica. Despite the abundance of fossils in Cenozoic deposits of the island, taxonomic studies with detailed systematic descriptions of bivalves are very rare. The first section of this work consists of taxonomic descriptions of invertebrates from Cenozoic deposits cropping out in two localities of King George Island, West Antarctica. Seven taxa of bivalves, including six new species were described from the Cape Melville Formation (Miocene), at Melville Peninsula. Seven taxa of invertebrates (bivalves, brachiopods, serpulid tubes, bryozoans, and echinoderm fragments) were described from the Polonez Cove Formation (Oligocene), at Vauréal Peak, a site previously unexplored paleontologically. The second section presents the results of a survey of the Cenozoic fossil record of Antarctic bivalves. The analysis of the fossil record confirmed that the current knowledge about the Cenozoic diversity of the group is very scarce. Moreover, comparison of Cenozoic bivalve genera from Antarctica and New Zealand showed that the greatest number of shared taxa is recorded in Eocene deposits. This finding supports the geographic isolation of Antarctic and the drop in faunal interchange between Antarctica and periphery after the Oligocene. Analysis of the fossil record suggested an intensive dispersal event during the Eocene, and restricted pulses of dispersal from the Oligocene onwards. The distribution pattern of taxa provides partial support for available reconstructions of marine currents. Eocene dispersal would have occurred from Antarctica to New Zealand in Atlantic-Pacific direction. This dispersal event is consistent with the hypothesis of shallow marine connections between West and East Antarctica (\"Shackleton Seaway\"), as well of the existence of the Weddellian Province from the Late Cretaceous to the Eocene. Dispersal events following the Oligocene would have occurred during and after the establishment of the Circum-Antarctic Current, along the West-Atlantic and East margins of Antarctica towards New Zealand, and no longer through \"Shackleton Seaway\". These analyses also support the hypothesis of full-scale glaciation in West Antarctica from the early Oligocene onwards, and in East Antarctica since the mid-Oligocene, with warmer temperatures than today.

Thesis (M.S.)--Ohio University, August, 2009. / Title from PDF t.p. Includes bibliographical references.

Wood mice of the genus Apodemus rank among the most common extant mammals throughout Europe. A history of its species and their distributional ranges is to be looked upon as a serious topic relevant also for general concepts on current setting of European biotic diversity. Nevertheless, despite several phylogeographic studies on that subject and a rich fossil record of the genus, many details of that issue remains only very poorly comprehended. This state is essentially caused by enormous complications with phenotype discrimination of individual species, their extensive variation and broad between-species overlaps in all phenotype traits. The present project was intended to establish the techniques enabling reliable species identification applicable to fragmentary fossil materials and to use them for retrieving information on range dynamics and phylogenetic history of individual species during the Quaternary past. It begun with detailed morphometric analyses of extant populations, using the characters available in fossils, and with establishing a concept of alternative parataxonomic classification covering the complete phenotype span of the genus. The respective techniques were applied onto an extensive set of well-dated records from the Late Pleistocene-Holocene sedimentary series from diverse...

No description available.

No description available.

The southwest monsoon is responsible for a summer precipitation maximum for much of the southwest U.S. Biostratigraphies of pollen, plant macrofossils, and aquatic fossils in lakes from near modern monsoon boundaries in conjunction with climate modelling suggests variations in strength of the monsoon system during the late Quaternary. At Montezuma Well, Arizona, high percentages of Pinus and Juniperus pollen as well as maximum influxes of Quercus and Gramineae pollen suggest a shift from dominantly winter to summer precipitation between ca. 12,000 and 9000 yr BP. Maximum aridity occurred 7000-4000 yr BP, coincident with lowest lake levels. In the High Plateaus region of the Colorado Plateau, high Artemisia to Chenopodiaceae-Amaranthus pollen ratios suggests precipitation maxima until ca. 6000 yr BP at Fryingpan Lake and 5000 yr BP at Posy Lake. Pollen records suggest that Pinus edulis, P. ponderosa, and Quercus gambelii, were present on the western Colorado Plateau throughout the Holocene. Expansion of shadscale steppe vegetation at low elevations and upslope movement of ecotones for Pinus edulis, P. ponderosa, and Q. gambelii after ca. 6000 yr BP and low lake levels ca. 5000-3700 yr BP, suggest a period of maximum aridity from decreased summer precipitation. In the San Luis Valley, Colorado, pollen records from Head Lake on the basin floor suggest an expansion of oaks and junipers at the Pleistocene/Holocene boundary that may indicate increased summer precipitation. Lake levels of Head Lake fell sharply after ca. 9500 yr BP. Pollen records from Como Lake in the Sangre de Cristo Mountains suggest that Pinus ponderosa was established in the area by ca. 12,000 yr BP and Pinus edulis by ca. 9500 yr BP. Highland regions such as the High Plateaus (until ca. 6000-5000 yr BP) and central Colorado (until ca. 4000 yr BP) may have experienced Holocene summer precipitation maxima later into the Holocene than sites in lower elevation regions. Regional orographic uplift as a catalyst for convective summer precipitation may be responsible for the duration of summer precipitation maxima in these regions. On a longitudinal gradient, sites to the west such as in the southern Great Basin and Mohave Desert may have recorded enhanced summer precipitation earlier, reflecting different histories of the low-level jets in the southwest. The paleoecologic record generally confirm predictions of general circulation models (GCMs) that southwest monsoon circulation was enhanced from 12,000-6000 yr BP in response to peaks in annual (11,500-11,000 yr BP) and summer insolation (10,000-9000 yr BP) during the late Quaternary.

Les érymides sont des crustacés décapodes marins ayant une morphologie comparable à celle des homards actuels. Ils sont regroupés au sein d’une unique famille, les Erymidae Van Straelen, 1925, caractérisée par la présence d’une plaque intercalaire dorsale. Ces crustacés sont présents dès la fin du Permien. Ils se sont diversifiés et répandus dans le monde entier au Jurassique et ont perduré jusqu’au Paléocène. Ils sont particulièrement abondants au Jurassique, fossilisés dans des dépôts issus d’environnements variés : de faible profondeur – comme les calcaires lithographiques de Solnhofen (Allemagne) –, très profonds – comme La Voulte (France) –, ou encore dans différents milieux de plate-forme – comme le Terrain à Chailles (France). Depuis les premières descriptions d’érymides, au début du XIXe siècle, de nombreux auteurs se sont attachés à décrire de nouvelles formes et à tenter d’élucider les affinités phylogénétiques de ces crustacés éteints. Ces nombreux travaux ont abouti à l’installation et à la propagation de confusions rendant douteuse la systématique des érymides tant au niveau des genres que des espèces. Ces problèmes taxinomiques particulièrement marqués chez les érymides — on parle d’ailleurs de « problème érymidien » — sont accompagnés d’un débat sur leur classification au sein des Pleocyemata. Jusqu’au début du XXIe siècle, la majorité des auteurs les classaient dans l’infraordre des Astacidea mais de récentes analyses phylogénétiques suggèrent l’intégration des érymides au sein des Glypheidea. Une autre étude a même abouti à la remise en cause du statut éteint des érymides. En effet, Schram & Dixon (2004) ont observé la plaque intercalaire sur l’actuel Enoplometopus A. Milne Edwards, 1862. Leur analyse a ensuite conduit au regroupement de cette forme actuelle avec les érymides au sein d’un même clade nommé Erymida. Les objectifs de cette thèse sont donc de remédier aux problèmes taxinomiques des érymides, d’élucider leurs affinités phylogénétiques et d’apporter des éléments permettant de mieux comprendre leur mode de vie et leur succès évolutif. Pour ce faire, une révision systématique aussi exhaustive que possible, appuyée sur l’étude de plus d’un millier de spécimens, a été réalisée. Elle a permis d’homogénéiser la description des 6 genres et des 75 espèces reconnues et d’identifier les caractères nécessaires à l’étude phylogénétique. L’arbre obtenu montre clairement que les érymides constituent un groupe particulier d’Astacidea auquel Enoplometopus n’appartient pas. De plus, la topologie de l’arbre de strict consensus soutient une refondation complète de la systématique du groupe. D’une unique famille, les érymides se retrouvent désormais répartis dans deux familles, distinguées par la présence/absence de la zone post-orbitaire : les Enoploclytidae n. fam. et les Erymidae. Cette dernière est d’ailleurs elle-même divisée en sous-familles, Eryminae Van Straelen, 1925 et Tethysastacinae n. s.–fam., en raison de l’architecture très simple des sillons de la carapace de Tethysastacus Devillez et al., 2016 comparée à celle des autres genres. Cette étude a aussi été l’occasion de formuler des hypothèses paléobiogéographiques qui demeurent, hélas, en grande partie spéculatives et incomplètes du fait des importantes discontinuités géographiques et stratigraphiques du registre fossile. Les observations de stades larvaires, des yeux, de la morphologie des pinces, de pores cuticulaires et de la variabilité intraspécifique chez certains spécimens ont également permis, en s’appuyant sur les formes actuelles, d’émettre des hypothèses sur le mode de vie de ces crustacés disparus. Enfin, la grande tolérance environnementale déduite des différentes formations géologiques ayant livré des érymides fossiles est probablement une des clés de leur succès au Mésozoïque et soulève la question des raisons de leur extinction. / Erymids are marine decapod crustaceans with a morphology close to that of extant lobsters. They are grouped within an unique family, Erymidae Van Straelen, 1925, based on the presence of a characteristic intercalated plate. These crustaceans were already present in the Permian, have become diversified and widespread during the Jurassic and have lasted until the Paleocene. The erymids are abundant during the Jurassic. They fossilized in deposits from various paleoenvironments: shallow water environments – like lithographic limestones from Solnhofen (Germany) –, from deep environments – like in La Voulte (France) –, and also from different platform environments – like the Terrain à Chailles (France). Since the first descriptions of erymids in the first part of the XIXth century, numerous authors have described new species and have attempted to establish the phylogenetic affinities of these extinct crustaceans. This high number of studies resulted with the apparition and propagation of confusions. So, the systematics of the erymids has become doubtful at both generic and specific levels. These taxonomic problems strongly affecting the erymids — the so called « erymidian problem » — are increased by their uncertain phylogenetic relationships among the Pleocyemata. Until the XXIst century, most of the authors classified the erymids within the infraorder Astacidea but recent phylogenetic analyses suggest an integration within Glypheidea. Moreover, a study has led to question the extinct status of the erymids. Indeed, Schram & Dixon (2004) have observed an intercalated plate in the extant Enoplometopus A. Milne Edwards, 1862. Their analysis has resulted with the clustering of this extant lobster together with the erymids within a same clade named Erymida. So, the purposes of this thesis are to rectify the taxonomic problems of the erymids, to elucidate their phylogenetic affinities and to provide observations which enable a better comprehension of their lifestyles and their evolutionary success. To reach these goals, a systematic review, supported by the examination of more than a thousand specimens, has been done. It has resulted in a homogenisation of the descriptions of the 6 genera and 75 species herein recognized and in the identification of useful characters for the phylogenetic study. The phylogenetic tree obtained clearly shows that erymids form a particular group of Astacidea and that Enoplometopus does not belong to this group. Moreover, the topology of the strict consensus tree supports a new systematic building of the group. From a unique family, the erymids are now spread into two families supported by the absence/presence of a post-orbital area: Enoploclytidae n. fam. and Erymidae. The last is also divided in subfamilies, Eryminae Van Straelen, 1925 and Tethysastacinae n. s.-fam., based on the very simple carapace groove pattern of Tethysastacus Devillez et al., 2016. This new study on the erymids was also an occasion to provide some paleobiogeographic hypotheses. But, unfortunately, they remain speculative and incomplete due to geographic and stratigraphic discontinuities of the fossil record. Observations of larval stages, of eyes, of P1 chela morphologies, of cuticular pores, and of intraspecific variability on some specimens have also enabled comparisons with extant forms. These observations led to provide hypotheses on the lifestyle of these extinct lobsters. Finally, the strong environmental tolerance was probably one of the keys of the success of the erymids during the Mesozoic but raised interrogations about the reasons of their extinction.

O conhecimento acerca dos invertebrados devonianos no Brasil teve início em fins do século XIX e a primeira metade do século XX, período no qual foram feitas expedições pioneiras às principais bacias paleozoicas do país - as bacias do Paraná, Parnaíba e Amazonas. Desta fase também resultaram importantes trabalhos científicos (e.g., Clarke, 1913), usados ainda hoje como referência em estudos. Dentre os trilobitas, os grupos mais representativos são os Homalonotidae e os Calmoniidae, cuja importância como ferramentas em questões paleoambientais e paleobiogeográficas vem sendo constatada a partir de recentes revisões sistemáticas em fósseis da Bacia do Paraná. Estes estudos também têm demonstrado que muitas das variações nos fósseis, interpretadas por pesquisadores anteriores como taxonômicas, decorrem de alterações tafonômicas, que podem levar a classificações errôneas. Neste contexto, o presente trabalho teve como objetivos revisar a sistemática dos trilobitas Phacopida (Homalonotidae e Calmoniidae) nas bacias do Parnaíba e Amazonas, incluindo, na primeira localidade, a nova ocorrência de São João Vermelho, no município de João Costa (PI), investigar, sempre que pertinente, a implicação das alterações tafonômicas na sistemática desses fósseis, identificando fatores diagenéticos/intempéricos que podem interferir nos caracteres morfológicos e discutir as implicações paleobiogeográficas dos Homalonotidae e Calmoniidae do Devoniano das bacias do Amazonas e Parnaíba, em relação a outras localidades adjacentes à estas bacias. Resultados das pesquisas mostram que a localidade de São João Vermelho é bastante promissora do ponto de vista paleontológico, pois consiste em uma nova localidade para o táxon M. tuberculatus, o qual também é documentado nas duas subbacias da Bacia do Paraná - Alto Garças e Apucarana, indicando que esta espécie apresenta alto grau de cosmopolitanismo. A análise de exemplares de Burmeisteria (Homalonotidae) desta localidade sugere que as diferenças nas espécies deste gênero no Brasil são mais sutis do que imaginado. Na Bacia do Amazonas, os Phacopida estão representados especialmente por Calmoniidae do gênero Malvinella, também presentes na Bolívia. As relações paleobiogeográficas entre as bacias paleozoicas brasileiras e a Bolívia, durante o Eo e Mesodevoniano, teriam sido mais estreitas do que se imaginava. A principal rota de migração para a Bacia do Parnaíba teria sido através da Bacia do Paraná, conforme constatado pela presença de M. tuberculatus. Já a migração para a Bacia do Amazonas teria se dado a sudoeste desta bacia, já que Malvinella ocorre apenas nesta bacia e na Bolívia / The knowledge on Brazilian Devonian invertebrates started by the end of the 19th century and the first half of the 20th century, period which pioneering expeditions were made to the main Paleozoic basins of the country - Paraná, Parnaíba and Amazon basins. Important scientific contributions also resulted from this fase (e.g. Clarke, 1913), still used today as reference in studies. Among trilobites, the most representative groups are the Homalonotidae and Calmoniidae, whose importance in paleoenvironmental and paleobiogeographical questions have been noted from recent systematic revisions in Paraná Basin fossils. These studies also have demonstrated that many variations in fossils, interpreted by previous researchers as taxonomic, are due to taphonomic alterations, which can lead to erroneous classifications. In this context, the current study aimed to review the systematics of Phacopida trilobites (Homalonotidae and Calmoniidae) from the Parnaíba and Amazon basins, including, in the former locality, the new occurrence of São João Vermelho, in the João Costa municipality (PI); to investigate, whenever relevant, .the implication of the taphonomic alterations in the systematics of these fossils, identifying diagenetic/weathering factors which can interfere in the morphological features; and to discuss the paleobiogeographic implications of the Homalonotidae and Calmoniidae from the Devonian of the Parnaíba and Amazon basins, in relation to other localities adjacent to these basins. Study results show that the locality of São João Vermelho is paleontologically promising, as it consists of a new place for the M. tuberculatus taxon, which is also documented in the both Paraná Basin subbasins - Alto Garças and Apucarana, indicating that this species is highly cosmopolitan. The analysis of Burmeisteria (Homalonotidae) exemplars from this locality suggests that differences between the species of this genus in Brazil are slighter than thought. In the Amazon Basin, Phacopida trilobites are especially represented by the Calmoniidae genus Malvinella, also present in Bolivia. The paleobiogeographic relations between the Paleozoic Brazilian basins and Bolivia, during Early and Middle Devonian, would have been closer than thought. The main migration route to the Parnaíba Basin would have been through the Paraná Basin, as found by the presence of M. tuberculatus. The migration to the Amazon Basin, in turn, would have occurred southwestward this basin, as Malvinella occurs only in this locality and in Bolivia.