O conhecimento acerca dos invertebrados devonianos no Brasil teve início em fins do século XIX e a primeira metade do século XX, período no qual foram feitas expedições pioneiras às principais bacias paleozoicas do país - as bacias do Paraná, Parnaíba e Amazonas. Desta fase também resultaram importantes trabalhos científicos (e.g., Clarke, 1913), usados ainda hoje como referência em estudos. Dentre os trilobitas, os grupos mais representativos são os Homalonotidae e os Calmoniidae, cuja importância como ferramentas em questões paleoambientais e paleobiogeográficas vem sendo constatada a partir de recentes revisões sistemáticas em fósseis da Bacia do Paraná. Estes estudos também têm demonstrado que muitas das variações nos fósseis, interpretadas por pesquisadores anteriores como taxonômicas, decorrem de alterações tafonômicas, que podem levar a classificações errôneas. Neste contexto, o presente trabalho teve como objetivos revisar a sistemática dos trilobitas Phacopida (Homalonotidae e Calmoniidae) nas bacias do Parnaíba e Amazonas, incluindo, na primeira localidade, a nova ocorrência de São João Vermelho, no município de João Costa (PI), investigar, sempre que pertinente, a implicação das alterações tafonômicas na sistemática desses fósseis, identificando fatores diagenéticos/intempéricos que podem interferir nos caracteres morfológicos e discutir as implicações paleobiogeográficas dos Homalonotidae e Calmoniidae do Devoniano das bacias do Amazonas e Parnaíba, em relação a outras localidades adjacentes à estas bacias. Resultados das pesquisas mostram que a localidade de São João Vermelho é bastante promissora do ponto de vista paleontológico, pois consiste em uma nova localidade para o táxon M. tuberculatus, o qual também é documentado nas duas subbacias da Bacia do Paraná - Alto Garças e Apucarana, indicando que esta espécie apresenta alto grau de cosmopolitanismo. A análise de exemplares de Burmeisteria (Homalonotidae) desta localidade sugere que as diferenças nas espécies deste gênero no Brasil são mais sutis do que imaginado. Na Bacia do Amazonas, os Phacopida estão representados especialmente por Calmoniidae do gênero Malvinella, também presentes na Bolívia. As relações paleobiogeográficas entre as bacias paleozoicas brasileiras e a Bolívia, durante o Eo e Mesodevoniano, teriam sido mais estreitas do que se imaginava. A principal rota de migração para a Bacia do Parnaíba teria sido através da Bacia do Paraná, conforme constatado pela presença de M. tuberculatus. Já a migração para a Bacia do Amazonas teria se dado a sudoeste desta bacia, já que Malvinella ocorre apenas nesta bacia e na Bolívia / The knowledge on Brazilian Devonian invertebrates started by the end of the 19th century and the first half of the 20th century, period which pioneering expeditions were made to the main Paleozoic basins of the country - Paraná, Parnaíba and Amazon basins. Important scientific contributions also resulted from this fase (e.g. Clarke, 1913), still used today as reference in studies. Among trilobites, the most representative groups are the Homalonotidae and Calmoniidae, whose importance in paleoenvironmental and paleobiogeographical questions have been noted from recent systematic revisions in Paraná Basin fossils. These studies also have demonstrated that many variations in fossils, interpreted by previous researchers as taxonomic, are due to taphonomic alterations, which can lead to erroneous classifications. In this context, the current study aimed to review the systematics of Phacopida trilobites (Homalonotidae and Calmoniidae) from the Parnaíba and Amazon basins, including, in the former locality, the new occurrence of São João Vermelho, in the João Costa municipality (PI); to investigate, whenever relevant, .the implication of the taphonomic alterations in the systematics of these fossils, identifying diagenetic/weathering factors which can interfere in the morphological features; and to discuss the paleobiogeographic implications of the Homalonotidae and Calmoniidae from the Devonian of the Parnaíba and Amazon basins, in relation to other localities adjacent to these basins. Study results show that the locality of São João Vermelho is paleontologically promising, as it consists of a new place for the M. tuberculatus taxon, which is also documented in the both Paraná Basin subbasins - Alto Garças and Apucarana, indicating that this species is highly cosmopolitan. The analysis of Burmeisteria (Homalonotidae) exemplars from this locality suggests that differences between the species of this genus in Brazil are slighter than thought. In the Amazon Basin, Phacopida trilobites are especially represented by the Calmoniidae genus Malvinella, also present in Bolivia. The paleobiogeographic relations between the Paleozoic Brazilian basins and Bolivia, during Early and Middle Devonian, would have been closer than thought. The main migration route to the Parnaíba Basin would have been through the Paraná Basin, as found by the presence of M. tuberculatus. The migration to the Amazon Basin, in turn, would have occurred southwestward this basin, as Malvinella occurs only in this locality and in Bolivia.

It is widely believed that Australian Aboriginals utilised fire to manage various landscapes however to what extent this impacted on Australia???s ecosystems remains uncertain. The late Pleistocene/Holocene fire history from three sites within the Sydney Basin, Gooches Swamp, Lake Baraba and Kings Waterhole, were compared with archaeological and palaeoclimatic data using a novel method of quantifying macroscopic charcoal, which is presented in this study. The palynology and other palaeoecological proxies were also investigated at the three sites. The Gooches Swamp fire record appeared to be most influenced by climate and there was an abrupt increase in fire activity from the mid-Holocene perhaps associated with the onset of modern El Ni??o dominated conditions. The Kings Waterhole site also displayed an abrupt increase in charcoal at this time however there was a marked decrease in charcoal from ~3 ka. Lake Baraba similarly had displayed low levels of charcoal in the late Holocene. At both Kings Waterhole and Lake Baraba archaeological evidence suggests intensified human activity in the late Holocene during this period of lower and less variable charcoal. It is hence likely that at these sites Aboriginal people controlled fire activity in the late Holocene perhaps in response to the increased risk of large intense fires under an ENSO-dominated climate. The fire history of the Sydney Basin varies temporally and spatially and therefore it is not possible to make generalisations about pre-historic fire regimes. It is also not possible to use ideas about Aboriginal fire regimes or pre-historic activity as a management objective. The study demonstrates that increased fire activity is related to climatic variation and this is likely to be of significance under various enhanced Greenhouse scenarios. There were no major changes in the composition of the flora at all sites throughout late Pleistocene/Holocene although there were some changes in the relative abundance of different taxa. It is suggested that the Sydney Sandstone flora, which surrounds the sites, is relatively resistant to environmental changes. Casuarinaceae was present at Lake Baraba during the Last Glacial Maximum and therefore the site may have acted as a potential refugium for more mesic communities. There was a notable decline in Casuarinaceae during the Holocene at Lake Baraba and Kings Waterhole, a trend that has been found at a number of sites from southeastern Australia.

It is widely believed that Australian Aboriginals utilised fire to manage various landscapes however to what extent this impacted on Australia???s ecosystems remains uncertain. The late Pleistocene/Holocene fire history from three sites within the Sydney Basin, Gooches Swamp, Lake Baraba and Kings Waterhole, were compared with archaeological and palaeoclimatic data using a novel method of quantifying macroscopic charcoal, which is presented in this study. The palynology and other palaeoecological proxies were also investigated at the three sites. The Gooches Swamp fire record appeared to be most influenced by climate and there was an abrupt increase in fire activity from the mid-Holocene perhaps associated with the onset of modern El Ni??o dominated conditions. The Kings Waterhole site also displayed an abrupt increase in charcoal at this time however there was a marked decrease in charcoal from ~3 ka. Lake Baraba similarly had displayed low levels of charcoal in the late Holocene. At both Kings Waterhole and Lake Baraba archaeological evidence suggests intensified human activity in the late Holocene during this period of lower and less variable charcoal. It is hence likely that at these sites Aboriginal people controlled fire activity in the late Holocene perhaps in response to the increased risk of large intense fires under an ENSO-dominated climate. The fire history of the Sydney Basin varies temporally and spatially and therefore it is not possible to make generalisations about pre-historic fire regimes. It is also not possible to use ideas about Aboriginal fire regimes or pre-historic activity as a management objective. The study demonstrates that increased fire activity is related to climatic variation and this is likely to be of significance under various enhanced Greenhouse scenarios. There were no major changes in the composition of the flora at all sites throughout late Pleistocene/Holocene although there were some changes in the relative abundance of different taxa. It is suggested that the Sydney Sandstone flora, which surrounds the sites, is relatively resistant to environmental changes. Casuarinaceae was present at Lake Baraba during the Last Glacial Maximum and therefore the site may have acted as a potential refugium for more mesic communities. There was a notable decline in Casuarinaceae during the Holocene at Lake Baraba and Kings Waterhole, a trend that has been found at a number of sites from southeastern Australia.

Made available in DSpace on 2014-06-11T19:32:21Z (GMT). No. of bitstreams: 0 Previous issue date: 2007-08-31Bitstream added on 2014-06-13T19:42:53Z : No. of bitstreams: 1 oliveira_lgf_dr_rcla.pdf: 2551069 bytes, checksum: acd81b14b612f8d3d79d2b7529438352 (MD5) / Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) / O trabalho apresenta uma revisão dos conchostráceos da Formação Rio do Rasto (Bacia do Paraná, Permiano) e discussões substanciadas, em parte, pelos resultados dos experimentos realizados com representantes viventes cultivados em aquários. Concluiu-se que diversas características das carapaças normalmente usadas para a classificação dos conchostráceos fósseis não são diagnósticas para a distinção de táxons porque variam de acordo com fatores tafonômicos, ecológicos e ontogenéticos. Os conchostráceos vivem em águas continentais, são extremamente esteno-halinos e, ao contrário de alguns paradigmas da literatura, não vivem obrigatoriamente em corpos d'água efêmeros, toleram a presença de peixes predadores em seu habitat e duas espécies do mesmo gênero num único corpo d'água. Tais observações coadunam com o registro litológico e paleontológico da parte inferior da Formação Rio do Rasto, depositada em condições lacustres. A parte superior, onde há mais fácies eólicas e evidências de aumento de aridez, apresenta diversidade maior de conchostráceos, incluindo Hemicycloleaia mitchelli, uma espécie que permite realizar correlações cronoestratigráficos com a Austrália e Rússia. Sua idade provavelmente é neotatariana (=wuchiapingiana). Outros táxons, como Monoleiolophus unicostatus, também foram considerados nas correlações, estimando-se que a Formação Rio do Rasto esteja compreendida no Permiano médio-superior. / Conchostracans of the Rio do Rasto Formation (Paraná Basin, Permian) are revised and discussions based on experiments with living conchostracans are presented. One conclusion is that many carapace characteristics frequently used for classification change according to taphonomic, ecologic and ontogenetic factors. The conchostracans live in continental fresh waters and, on the contrary of some paradigms of the literature, the organisms are not restricted to ephemeral water bodies and tolerate the presence of predatory fishes and two species of one single genus. These observations combine well with the lithological and paleontological record of the lower Rio do Rasto Formation deposited under lacustrine conditions. The upper part, formed under more arid conditions, presents bigger conchostracan diversity and includes Hemicycloleaia mitchelli. This species may be used for wide correlations with Australia and Russia, and its age probably is Late Tatarian (= Wuchiapingian). Other taxa, as Monoleiolophus unicostatus, were also used for correlations and the age of the formation may be interpreted as mid to late Permian.

The Neogene history of Caribbean deep-sea benthic foraminifera was investigated by calculating changes in paleoproductivity, diversity and paleobiogeography ~26 to 2 Ma, which includes the progressive closure of the Central American Seaway. Three hypotheses were tested: (1) Paleoproductivity values prior to closure of the Central American Seaway are similar in both the Caribbean and equatorial Pacific and then diverge by the time of early shoaling events; (2) Diversity values of benthic foraminifera prior to the closure of the Central American Seaway were similar in the Caribbean and EEP, and had changed by the time of early shoaling; and (3) during the Miocene and into the Pleistocene, the progressive constriction of the CAS affected deep-sea benthic foraminiferal assemblages by increasing their dissimilarity between the Caribbean and equatorial Eastern Pacific. These hypotheses were tested with 104 samples from five Caribbean and EEP deep-sea cores by calculating paleoproductivity with multiple proxies, determining diversity indices and calculating biogeographic similarity coefficients. The data supported the first two hypotheses: The greatest change in paleoproductivity occurred at ~8 Ma during seaway constriction, when values diverged between the Caribbean and EEP. After complete seaway closure at ~4 Ma, the Caribbean became oligotrophic, noted by a decrease in high-organic flux species, and an increase in Nuttalides umbonifera, an indicator species. The largest changes in species-level diversity occurred with the barrier to deep-water flow at ~12 Ma, and Caribbean diversity increased at ~8 Ma with seaway constriction. However, the third hypothesis was rejected: Increases in assemblage similarity actually occurred during most major paleoceanographic events, with the only decrease in Caribbean-EEP similarity occurring at ~12 Ma, coincident with a drop in diversity and emplacement of the Panama isthmian sill. Thus, the barrier to deep-water flow at ~12 Ma affected the composition of tropical American benthic foraminifera more than the largest change in paleoproductivity at ~8 Ma, or closure of the Central American Seaway at ~4 Ma.

Many organisms are known to reach high levels of endemism and biodiversity in the temperate forests of Southern Appalachia, especially in the dense forests and rugged terrain of the Blue Ridge physiographic province. Many plants and fungi reach their highest levels of biodiversity in these mountains, as does one group of vertebrates: the lungless salamanders, Plethodontidae. This family of salamanders hosts the most species of any other group of salamanders on earth and has adapted to a wide range of habitats. Only two of the approximately twenty-seven known genera are not found in North or South America, and while we know much about the modern-day biology of this family, few fossils older than ~15,000 years have been recovered, complicating our understanding of the historical distribution of this group and the timing of key evolutionary events within the family. Recently discovered salamander fossils from the Gray Fossil Site provide the foundation of this project. We describe these exceptionally large plethodontid remains to a new genus that belongs to the group containing dusky salamanders, or desmognathans. The morphology of the fossil material resembles Phaeognathus hubrichti, an extant burrowing species from southern Alabama with a suite of primitive characteristics. Comparison of the fossil material to modern desmognathans using geometric statistical methods has revealed that the extinct form was likely similar in lifestyle to P. hubrichti, but considerably larger. It was unparalleled in the southern Appalachians in terms of size and ecology, and reveals a more complex evolutionary history for desmognathan salamanders.

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No description available.

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La sous-famille des Suinae est largement répandue en Afrique au Plio-Pléistocène et a été abondamment utilisée pour corréler biochronologiquement les sites à hominidés en se basant sur l'évolution morphologique rapide des troisièmes molaires dans différentes lignées. À partir d'un échantillon important de suinés africains actuels, les schémas de variation morphologique crânio-mandibulaire et dentaire sont quantifiés par morphométrie géométrique. Ce référentiel actuel sert à estimer la variabilité morphologique dans le registre fossile. La révision de la paléobiodiversité du genre Kolpochoerus (le suiné plio-pléistocène le plus abondant) dans les bassins du Turkana et de l'Awash s'appuie sur l'étude anatomique et morphométrique de matériels publié et inédit découverts dans la basse vallée de l'Omo (formation de Shungura) et dans la vallée moyenne de l'Awash en Éthiopie. Une nouvelle espèce fossile est décrite, les trajectoires évolutives au sein de chaque espèce sont quantifiées, et les interprétations biostratigraphiques sont révisées. Les liens de parenté au sein du genre Kolpochoerus sont décrits grâce à l'analyse cladistique. L'étude de la paléoécologie des suinés africains est basée sur l'anatomie comparée et la biogéochimie des isotopes stables (carbone et oxygène). Les suinés fossiles présentent une gamme de taille importante, ainsi que des régimes alimentaires et des habitats variés. Les nouvelles connaissances acquises sur leur biodiversité, leur phylogénie, et leur écologie permettent de proposer une histoire paléobiogéographique des Suinae en Afrique. / The subfamily Suinae is widespread in Plio-Pleistocene deposits in Africa, and was heavily used as a biochronological tool to correlate hominid-bearing sites based on the rapid morphological evolution of third molars in different lineages. A large sample of extant African suines enables to quantify patterns of variation in cranio-mandibular and dental morphology using geometric morphometrics. This modern referential is used to estimate the morphological variability in the fossil record. Revision of the paleobiodiversity of the genus Kolpochoerus (the most abundant Plio-Pleistocene suine) in Turkana and Awash basins is based on anatomical and morphometric studies of published and unpublished material discovered in the Lower Omo Valley (Shungura Formation) and in the Middle Awash Valley in Ethiopia. A new fossil species is described, evolutionary trajectories within each species are quantified, and biostratigraphic interpretations are revised. Phylogenetic relationships within the genus Kolpochoerus are described by the cladistic analysis. Paleoecological study of African suines is based on comparative anatomy and stable isotopic biogeochemistry (carbon and oxygen). Fossil suines display a wide range of body size, as well as various diets and habitats. The new results produced regarding their biodiversity, phylogeny, and ecology, enable to reconstruct a paleobiogeographic history of Suinae in Africa.