I studied the acquisition of foraging skills by lambs eating shrub or grass in three experiments. The general approach was to isolate those skills involved in prehending forage from those related to the acceptance of novel foods. Treatment lambs received 15 times more exposure to grass or shrub than did control lambs. Lambs were tested in 2.5 x 2.5 meter monocultures of shrub or grass 5 min/d, on two separate occasions. Height, bulk density and spatial arrangement of plant material were controlled during testing. In Experiment 1, I studied the acquisition of foraging skills by lambs on monocultures of shrubs. I found that inexperienced lambs ingested less forage per unit time than experienced lambs because they had less developed prehension skills. Lambs were more successful at plucking individual leaves than breaking twigs from branches or stripping 1 eaves. Lambs emphasized prehension patterns that were most successful. In Experiment 2, I explored the extent to which foraging skills gained on shrub or grass are specific to shrub or grass. Grass-experienced lambs were more successful at prehending and ingesting grass than were shrub experienced lambs. No statistical differences were observed between shrub- and grass-experienced lambs at prehending and ingesting shrub although numerical values were higher for shrub- than grass-experienced lambs. I hypothesize that a relationship exists between plant form, prehension pattern and foraging experience. In Experiment 3, I studied the degree to which lambs experienced with grass or shrub vary in their ability to prehend and ingest vegetative and flowering grass. Grass- and shrub-experienced lambs ingested more flowering than vegetative grass per unit time. Grass-experienced lambs were more efficient than shrub-experienced lambs at prehending and ingesting vegetative and flowering grass. Shrub-experienced lambs experienced more difficulty at prehending flowering compared to vegetative grass while grass-experienced lambs did not.

When crested wheatgrass (Agropyron desertorum) grows in mixture with sagebrush (Artemisia tridentata), its production declines. Its production increases when grown in mixture with fourwing saltbush (Atriplex canescens), according to previous reports. This study investigated soil water extraction and potassium (K) nutrition of the two shrubs to identify possible causes of the differential responses of crested wheatgrass. Crested wheatgrass had reduced, rather than increased, nitrogen (N) and K yield in mixture with fourwing saltbush. No differences in N and phosphorous (P) concentrations were observed between sagebrush and fourwing saltbush, but fourwing saltbush had a much higher K concentration and returned nearly twice as much K to the soil as sagebrush by throughfall and litterfall. Throughfall additions were much greater than those from litterfall. AK-fertilization/water-stress, two-factor greenhouse experiment was conducted with crested wheatgrass. High- and medium-K-fertilization treatments had highest tissue K concentration, but biomass yield was reduced in waterstressed plants with high K-fertilization. A difference of 1.56 MPa in osmotic adjustment was observed between waterstressed plants with high K-fertilization and irrigated, low-K-fertilization plants. These results suggest that K accumulation in fourwing saltbush may be a factor for enhanced crested wheatgrass productivity. Crested wheatgrass grown in mixture with fourwing saltbush had lowered predawn and mid-day xylem water potentials compared with monoculture and sagebrush mixture plots, but no other treatment differences were observed for any species. Fourwing saltbush monoculture plots had the most uniform water extraction rates and may compete less for water than sagebrush when crested wheatgrass extraction rates are highest.

Die Vegetationskomposition und –struktur, beispielsweise die unterschiedliche Architektur von Bäumen, Sträuchern, Gräsern und Kräutern, bietet ein großes Spektrum an Habitaten und Nischen, die wiederum eine hohe Tierdiversität in den Savannensystemen des südlichen Afrikas ermöglichen. Dieses Ökosystem wurde jedoch über Jahrzehnte weltweit durch intensive anthropogene Landnutzung (z.B. Viehwirtschaft) nachhaltig verändert. Dabei wurden die Zusammensetzung, Diversität und Struktur der Vegetation stark verändert. Überweidung in Savannensystemen führt zu einer Degradation des Habitates einhergehend mit dem Verlust von perennierenden Gräsern und krautiger Vegetation. Dies führt zu einem Anstieg an vegetationsfreien Bodenflächen. Beides, sowohl der Verlust an perennierenden Gräsern und krautiger Vegetation sowie der Anstieg an vegetationsfreien Flächen führt zu verbesserten Etablierungsbedingungen für Sträucher (z.B. Rhigozum trichotomum, Acacia mellifera) und auf lange Sicht zu stark verbuschten Flächen. Die Tierdiversität in Savannen ist hiervon entscheidend beeinflusst. Mit sinkender struktureller Diversität verringert sich auch die Tierdiversität. Während der Einfluss von Überweidung auf die Vegetation relativ gut untersucht ist sind Informationen über den Einfluss von Überweidung auf die Tierdiversität, speziell für Reptilien, eher spärlich vorhanden. Zusätzlich ist sehr wenig bekannt zum Einfluss auf die Populationsdynamik (z.B. Verhaltensanpassungen, Raumnutzung, Überlebensrate, Sterberate) einzelner Reptilienarten. Ziel meiner Doktorarbeit ist es den Einfluss von Überweidung durch kommerzielle Farmnutzung auf die Reptiliengemeinschaft und auf verschiedene Aspekte der Populationsdynamik der Echse Pedioplanis lineoocellata lineoocellata zu untersuchen. Hinsichtlich bestimmter Naturschutzmaßnahmen ist es einerseits wichtig zu verstehen welchen Auswirkungen Überweidung auf die gesamte Reptiliengemeinschaft hat. Und zum anderen wie entscheidende Faktoren der Populationsdynamik beeinflusst werden. Beides führt zu einem besseren Verständnis der Reaktion von Reptilien auf Habitatdegradation zu erlangen. Die Ergebnisse meiner Doktorarbeit zeigen eindeutig einen negativen Einfluss der Überweidung und der daraus resultierende Habitatdegradation auf (1) die gesamte Reptiliengemeinschaft und (2) auf einzelne Aspekte der Populationsdynamik von P. lineoocellata. Im Teil 1 wird die signifikante Reduzierung der Reptiliendiversität und Abundanz in degradierten Habitaten beschrieben. Im zweiten Teil wird gezeigt, dass P. lineoocellata das Verhalten an die verschlechterten Lebensbedingungen anpassen kann. Die Art bewegt sich sowohl häufiger als auch über einen längeren Zeitraum und legt dabei größere Distanzen zurück. Zusätzlich vergrößerte die Art ihr Revier (home range) (Teil 3). Im abschließenden Teil wird der negative Einfluss von Überweidung auf die Populationsdynamik von P. lineoocellata beschrieben: In degradierten Habitaten nimmt die Populationsgröße von adulten und juvenilen Echsen ab, die Überlebens- und Geburtenrate sinken, währen zusätzlich das Prädationsrisiko ansteigt. Verantwortlich hierfür ist zum einen die ebenfalls reduzierte Nahrungsverfügbarkeit (Arthropoden) auf degradierten Flächen. Dies hat zur Folge, dass die Populationsgröße abnimmt und die Fitness der Individuen verringert wird, welches sich durch eine Reduzierung der Überlebens- und Geburtenrate bemerkbar macht. Und zum anderen ist es die Reduzierung der Vegetationsbedeckung und der Rückgang an perennierenden Gräsern welche sich negativ auswirken. Als Konsequenz hiervon gehen Nischen und Mikrohabitate verloren und die Möglichkeiten der Reptilien zur Thermoregulation sind verringert. Des Weiteren hat dieser Verlust an perennierender Grasbedeckung auch ein erhöhtes Prädationsrisikos zur Folge. Zusammenfassend lässt sich sagen, dass nicht nur Bäume und Sträucher, wie in anderen Studien gezeigt, eine bedeutende Rolle für die Diversität spielen, sondern auch das perennierende Gras eine wichtige Rolle für die Faunendiversität spielt. Weiterhin zeigte sich, dass Habitatdegradation nicht nur die Population als gesamtes beeinflusst, sondern auch das Verhalten und Populationsparameter einzelner Arten. Des Weiteren ist es Reptilien möglich durch Verhaltensflexibilität auf verschlechterte Umweltbedingen zu reagieren. / In semi-arid savannah ecosystems, the vegetation structure and composition, i.e. the architecture of trees, shrubs, grass tussocks and herbaceous plants, offer a great variety of habitats and niches to sustain animal diversity. In the last decades intensive human land use practises like livestock farming have altered the vegetation in savannah ecosystems worldwide. Extensive grazing leads to a reduction of the perennial and herbaceous vegetation cover, which results in an increased availability of bare soil. Both, the missing competition with perennial grasses and the increase of bare soils favour shrub on open ground and lead to area-wide shrub encroachment. As a consequence of the altered vegetation structure and composition, the structural diversity declines. It has been shown that with decreasing structural diversity animal diversity decline across a variety of taxa. Knowledge on the effects of overgrazing on reptiles, which are an important part of the ecosystem, are missing. Furthermore, the impact of habitat degradation on factors of a species population dynamic and life history, e.g., birth rate, survival rate, predation risk, space requirements or behavioural adaptations are poorly known. Therefore, I investigated the impact of overgrazing on the reptile community in the southern Kalahari. Secondly I analysed population dynamics and the behaviour of the Spotted Sand Lizard, Pedioplanis l. lineoocellata. All four chapters clearly demonstrate that habitat degradation caused by overgrazing had a severe negative impact upon (i) the reptile community as a whole and (ii) on population parameters of Pedioplanis l. lineoocellata. Chapter one showed a significant decline of regional reptile diversity and abundance in degraded habitats. In chapter two I demonstrated that P. lineoocellata moves more frequently, spends more time moving and covers larger distances in degraded than in non-degraded habitats. In addition, home range size of the lizard species increases in degraded habitats as shown by chapter three. Finally, chapter four showed the negative impacts of overgrazing on several population parameters of P. lineoocellata. Absolute population size of adult and juvenile lizards, survival rate and birth rate are significantly lower in degraded habitats. Furthermore, the predation risk was greatly increased in degraded habitats. A combination of a variety of aspects can explain the negative impact of habitat degradation on reptiles. First, reduced prey availability negatively affects survival rate, the birth rate and overall abundance. Second, the loss of perennial plant cover leads to a loss of niches and to a reduction of opportunities to thermoregulate. Furthermore, a loss of cover and is associated with increased predation risk. A major finding of my thesis is that the lizard P. lineoocellata can alter its foraging strategy. Species that are able to adapt and change behaviour, such as P. lineoocellata can effectively buffer against changes in their environment. Furthermore, perennial grass cover can be seen as a crucial ecological component of the vegetation in the semi-arid savannah system of the southern Kalahari. If perennial grass cover is reduced to a certain degree reptile diversity will decline and most other aspects of reptile life history will be negatively influenced. Savannah systems are characterised by a mixture of trees, shrubs and perennial grasses. These three vegetation components determine the composition and structure of the vegetation and accordingly influence the faunal diversity. Trees are viewed as keystone structures and focal points of animal activity for a variety of species. Trees supply animals with shelter, shade and food and act as safe sites, nesting sites, observation posts and foraging sites. Recent research demonstrates a positive influence of shrub patches on animal diversity. Moreover, it would seem that intermediate shrub cover can also sustain viable populations in savannah landscapes as has been demonstrated for small carnivores and rodent species. The influence of perennial grasses on faunal diversity did not receive the same attention as the influence of trees and shrubs. In my thesis I didn’t explicitly measure the direct effects of perennial grasses but my results strongly imply that it has an important role. If the perennial grass cover is significantly depleted my results suggest it will negatively influence reptile diversity and abundance and on several populations parameters of P. lineoocellata. Perennial grass cover is associated with the highest prey abundance, reptile diversity and reptile abundance. It provides reptiles both a refuge from predators and opportunities to optimise thermoregulation. The relevance of each of the three vegetation structural elements is different for each taxa and species. In conclusion, I can all three major vegetation structures in the savannah system are important for faunal diversity.

The Carrizo-Wilcox aquifer is a valuable groundwater resource, situated in a semi-arid landscape of Southwest Texas, where over-use by dependent farming practices has lowered aquifer levels. In semi-arid regions, rates of groundwater recharge are predominantly low due to high potential evapotranspiration rates; however, least understood is the role that vegetation plays in soil-plant-water dynamics. Vegetation management potentially plays a major role in countering the loss to recharge because evapotranspiration (ET) varies with vegetation type and cover. The conversion from shrubland to grassland likely reduces rooting depths and total plant cover. Subsequently, deep drainage (percolation below the root zone) will likely increase and lead to groundwater recharge, at least temporarily. The primary aims of the study were to identify those biotic and abiotic factors facilitating deep drainage and to examine differences in recharge for the years following clearing of natural shrub vegetation. Soil chloride was examined to estimate long-term recharge rates, since its concentration in the soil is influenced by the movement of water. Short-term soil moisture trends were also monitored for any water movement deep in the soil profile in response to individual rain events. Rooting depths decreased following removal of vegetation; yet root biomass unexpectedly increased due to successful grass establishment during the first five years after treatment. Soil properties did not vary between treatments, indicating that the majority of chloride differences seen were a consequence of vegetation change. Peak and total soil chloride concentrations were expected to decrease and occur deeper in the soil profile 15-30 years following the clearing of woody vegetation. Total chloride decreased by up to 65% after 30 years and resulted in an estimated 14.9 mm/yr more recharge compared to adjacent untreated controls. Evidence in this study suggest that much of this chloride is leached during the first five years following treatment and that more leaching occurs in especially wet periods. During the wet 2007 growing season, soil moisture below the root zone increased by up to 17% after vegetation clearing. The results of this study indicate that hydrologic changes following brush removal were evident in this system and are likely to positively influence groundwater recharge in the long-term.

Die derzeitige Markt- und Preisentwicklung bei Schnittblumen gibt neue Impulse für eine Eigenproduktion. Besonders für Einzelhandelsgärtnereien bietet sich die energie- und investitionsarme Produktion im Freiland an. Innerhalb des Projektes "Sortimentssichtung und Erarbeitung von Anbauverfahren zur Erzeugung von Freilandschnittblumen und -schnittgrün von Stauden mit Absatzschwerpunkten vor und nach den Sommerferien" im Fachbereich Gartenbau Dresden-Pillnitz der LfL konnten Anbauhinweise und Sortimentsempfehlungen für den Freilandschnittanbau unter hiesigen Bedingungen erarbeitet werden. Die über einen längeren Zeitraum ermittelten Ertragskennzahlen am vollsonnigen Standort sowie die Daten zur Aufbereitung, zur Schnittblumenqualität und Vasenhaltbarkeit der Stauden ermöglichten die Zusammenstellung der in der Schriftenreihe dokumentierten umfang- und abwechslungsreichen Empfehlungssortimente. Sie erleichtern dem Erwerbsanbauer die Arten- und Sortenauswahl entsprechend der betriebsspezifischen Bedingungen, betriebswirtschaftlichen Überlegungen und des Kundenklientel. Im Mittelpunkt stehen dabei Stauden für die absatzstärkeren Zeiträume vor und nach den Sommerferien. Bei ausgewählten Arten und Sorten kann der Rückschnitt des Austriebes Mitte Mai zur Blühterminverspätung genutzt werden. Diesbezügliche Versuchsergebnisse sind den Sortimentsempfehlungen im Bericht angefügt. Innerhalb des Projektes wurden auch intensive Anbauversuche mit Montbretien und Clematis als Schnittblume erfolgreich durchgeführt. Als Ergebnis werden Sortimentsempfehlungen und Hinweise zum Kulturablauf und zur Produktion im Freiland gegeben. Wertvolle Anbauergebnisse bezüglich des Einsatzes von Xylit als Mulchstoff zur Reduzierung des Unkrautaufkommens runden die Projektergebnisse ab.

This three part study attempted to enhance our understanding of vegetation change and its potential effects on ecohydrology in drylands. The first study developed a method to measure the velocity of shallow overland flow. Under rainfall simulation, dye tracers were applied to runoff and photographed to calculate mean surface velocity. Results showed this approach was a significant improvement explaining 13% more of the variation in mean velocity compared to traditional methods. Results from the first study were used to compare hydraulic parameters on shrub- and grass-dominated plots in the second study. Previous research has suggested microtopography in shrublands acts to concentrate flow, leading to increased runoff velocity compared to grasslands. However, present findings showed that flow velocities were similar on many grass and shrub plots; only plots with ground cover > 90% exhibited significantly lower flow velocities, and some shrub-dominated plots had lower flow velocities than grass-dominated plots implying that horizontal water flux is reduced under certain states of woody plant encroachment. In terms of ground cover characteristics, velocity increased rapidly with increases in the fraction of bare soil, up to a value of ~20% bare soil. Above ~20% bare soil, basal gap became a dominant factor suggesting a possible threshold where spatial metrics related to the distance between plants become important indicator of shallow flow velocity. The third study tested an approach to quantify woody plant canopy metrics over large areas. Radar has been used to map biomass in forests but few studies have examined open canopy ecosystems. Field measurements of shrublands were compared to satellite images to identify the relationship between radar signal and height and cover of woody vegetation. Results indicated that radar signal increased positively with shrub height or shrub volume explaining 74% and 90% of the variation, respectively. The effect of surface roughness and sub-canopy species on radar signal appears reduced when images are collected at large incidence angles.

No description available.

Doctor of Philosophy(PhD) / Soil contamination with heavy metal(loid)s is a major environmental problem that requires effective and affordable remediation technologies. The utilisation of plants to remediate heavy metal(loid)s contaminated soils has attracted considerable interest as a low cost green remediation technology. The process is referred to as phytoremediation, and this versatile technology utilises plants to phytostabilise and/or phytoextract heavy metal(loid)s from contaminated soils, thereby effectively minimising their threat to ecosystem, human and animal health. Plants that can accumulate exceptionally high concentrations of heavy metal(loid)s into above-ground biomass are referred to as hyperaccumulators, and may be exploited in phytoremediation, geobotanical prospecting and/or phytomining of low-grade ore bodies. Despite the apparent tangible benefits of utilising phytoremediation techniques, a greater understanding is required to comprehend the ecophysiological aspects of species suitable for phytoremediation purposes. A screening study was instigated to assess phytoremediation potential of several fern species for soils contaminated with cadmium (Cd), chromium (Cr), copper (Cu), nickel (Ni), lead (Pb) and zinc (Zn). Hyperaccumulation was not observed in any of the studied species, and in general, species excluded heavy metal uptake by restricting their translocation into aboveground biomass. Nephrolepis cordifolia and Hypolepis muelleri were identified as possible candidates in phytostabilisation of Cu-, Pb-, Ni- or Zn-contaminated soils and Dennstaedtia davallioides appeared favourable for use in phytostabilisation of Cu- and Zn-contaminated soils. Conversely, Blechnum nudum, B. cartilagineum, Doodia aspera and Calochlaena dubia were least tolerant to most heavy metals and were classified as being least suitable for phytoremediation purposes Ensuing studies addressed the physiology of arsenic (As) hyperaccumulation in a lesser known hyperaccumulator, Pityrogramma calomelanos var. austroamericana. The phytoremediation potential of this species was compared with that of the well known As hyperaccumulator Pteris vittata. Arsenic concentration of 3,008 mg kg–1 dry weight (DW) occurred in P. calomelanos var. austroamericana fronds when exposed to 50 mg kg–1 As without visual symptoms of phytotoxicities. Conversely, P. vittata was able to hyperaccumulate 10,753 mg As kg–1 DW when exposed to 100 mg kg–1 As without the onset of phytotoxicities. In P. calomelanos var. austroamericana, As was readily translocated to fronds with concentrations 75 times greater in fronds than in roots. This species has the potential for use in phytoremediation of soils with As levels up to 50 mg kg–1. Localisation and spatial distribution of As in P. calomelanos var. austroamericana pinnule and stipe tissues was investigated using micro-proton induced X-ray emission spectrometry (µ-PIXE). Freeze-drying and freeze-substitution protocols (using tetrahydrofuran [THF] as a freeze-substitution medium) were compared to ascertain their usefulness in tissue preservation. Micro-PIXE results indicated that pinnule sections prepared by freeze-drying adequately preserved the spatial elemental distribution and tissue structure of pinnule samples. In pinnules, µ-PIXE results indicated higher As concentration than in stipe tissues, with concentrations of 3,700 and 1,600 mg As kg–1 DW, respectively. In pinnules, a clear pattern of cellular localisation was not resolved whereas vascular bundles in stipe tissues contained the highest As concentration (2,000 mg As kg–1 DW). Building on these µ-PIXE results, the chemical speciation of As in P. calomelanos var. austroamericana was determined using micro-focused X-ray fluorescence (µ-XRF) spectroscopy in conjunction with micro-focused X-ray absorption near edge structure (µ-XANES) spectroscopy. The results suggested that arsenate (AsV) absorbed by roots was reduced to arsenite (AsIII) in roots prior to transport through vascular tissues as AsV and AsIII. In pinnules, AsIII was the predominant species, presumably as aqueous-oxygen coordinated compounds. Linear least-squares combination fits of µ-XANES spectra showed AsIII as the predominant component in all tissues sampled. The results also revealed that sulphur containing thiolates may, in part sequester accumulated As. The final aspect of this thesis examined several ecophysiological strategies of Ni hyperaccumulation in Hybanthus floribundus subsp. floribundus, a native Australian perennial shrub species and promising candidate in phytoremediation of Ni-contaminated soils. Micro-PIXE analysis revealed that cellular structure in leaf tissues prepared by freeze-drying was adequately preserved as compared to THF freeze-substituted tissues. Elemental distribution maps of leaves showed that Ni was preferentially localised in the adaxial epidermal tissues and leaf margin, with concentration of 10,000 kg–1 DW in both regions. Nickel concentrations in stem tissues obtained by µ-PIXE analysis were lower than in the leaf tissues (1,800 mg kg–1 vs. 7,800 mg kg–1 DW, respectively), and there was no clear pattern of compartmentalisation across different anatomical regions. It is possible that storage of accumulated Ni in epidermal tissues may provide Ni tolerance to this species, and may further act as a deterrent against herbivory and pathogenic attack. In H. floribundus subsp. floribundus seeds, µ-PIXE analysis did not resolve a clear pattern of Ni compartmentalisation and suggests that Ni was able to move apoplastically within the seed tissues. The role of organic acids and free amino acids (low molecular weight ligands [LMW]) in Ni detoxification in H. floribundus subsp. floribundus were quantified using high performance liquid chromatography (HPLC) and ultra performance liquid chromatography (UPLC). Nickel accumulation stimulated a significant increase in citric acid concentration in leaf extracts, and based on the molar ratios of Ni to citric acid (1.3:1–1.7:1), citric acid was sufficient to account for approximately 50% of the accumulated Ni. Glutamine, alanine and aspartic acid concentrations were also stimulated in response to Ni hyperaccumulation and accounted for up to 75% of the total free amino acid concentration in leaf extracts. Together, these LMW ligands may complex with accumulated Ni and contribute to its detoxification and storage in this hyperaccumulator species. Lastly, the hypothesis that hyperaccumulation of Ni in certain plants may act as an osmoticum under water stress (drought) was tested in context of H. floribundus subsp. floribundus. A 38% decline in water potential and a 68% decline in osmotic potential occurred between water stressed and unstressed plants, however, this was not matched by an increase in accumulated Ni. The results suggested that Ni was unlikely to play a role in osmotic adjustment in this species. Drought stressed plants exhibited a low water use efficiency which might be a conservative ecophysiological strategy enabling survival of this species in competitive water-limited environments.

As an ecological disturbance agent, wildfire is highly responsive to spatial and temporal variables. At the landscape scale, the spatiotemporal characteristics of fire are influenced by climate, which is non-stable and subject to oscillations, and by weather, which affects the intensity of burning and the severity of fire effects over short time periods. Like wildfire, vegetation communities and fuels are similarly influenced and modified by climate and weather, which configure not only the type of burning possible at any given time, but also the fire effects produced by burning. Furthermore, ignition sources, both natural and anthropogenic, vary over space and time, whether they are from natural ignitions in response to weather and climate factors, and anthropogenic ignitions, which are much less constrained. Within a given physiographic setting, the spatiotemporal conditions of wildfire can be understood as fire regimes, which can be expressed in terms of typical fire frequency, season of burning, ignition source, duration, fire size, patch size, fire rotation, and return interval. As a dynamic assemblage, vegetation communities, combustible fuels, ignition sources, and fire regimes are arrayed over topographic features at landscape scales. Humans are able to affect various and multiple components of this dynamic assemblage. Humans have the most direct control over ignitions, both in terms of adding ignitions to the weather and climate-modulated background of natural ignitions, or by suppressing anthropogenic ignitions and suppressing fires that do start. Humans can also manipulate vegetation communities and fuel complexes, either promoting or diminishing the chance for fires to burn and spread. Humans have far less control over weather and climate, although the enhanced greenhouse effect is beginning to be expressed in terms of climate change and unusually extreme weather, including weather variables that drive fire growth and spread, including low humidity, high temperatures, and increased winds. The objectives of this dissertation were to: (1) investigate the influences of several waves of human occupation on temporal fire regime characteristics in the Chiricahua Mountains across major topographic settings and forest types, and to detect cessation of widespread, low-intensity wildfire in specific locations; (2) account for the mechanisms by which the U.S. state has managed fires by managing anthropogenic ignitions, which has contributed to long-term deviation in formerly frequent fire regimes; (3) examine the effects from the return of large and severe wildfire following decades of fire prevention and suppression by categorizing the ecological trajectories of montane forests following mixed-severity reburning; and (4) understand how the post-disturbance recovery of burned areas is influenced by the response of shrub species to variations in fire severity, with the post-fire regeneration strategies of shrub species driving differences in patch structure and total cover. Major findings and contributions of this research include identifying distinct anthropogenic influences on temporal fire regime characteristics in several forest types in an Arizona Sky Island, including shortened fire frequencies during times of conflict between Apaches and colonizing forces. We found that periods of conflict with Spanish and later American militaries exerted an influence on fire frequencies, with higher-elevation forests burning more frequently than pre-Apache periods or during episodes of peacetime. We also find that single-tree fires, which are likeliest to express anthropogenic ignitions, do not significantly correspond to multi-year patterns in climate (Appendix A). We found that the U.S. state used Smokey Bear and other wildfire prevention media as a pyropolitical instrument aimed at simultaneously managing people, landscapes, and flammable forests, in the process binding proper fire behavior to notions of citizenship, territory, and ecology. The wildfire prevention campaign, with Smokey Bear as its avatar, was successful because it remained flexible in the face of social, economic, and environmental change within the United States, but the ultimate result of this success is an environmental feedback loop by which fire prevention and suppression produce the need for ever-greater state response (Appendix B). We found that mixed-severity reburning has differential effects on various structural and demographic components of vegetation communities, with trees, shrubs, and regeneration responding differently according to plant functional traits. Although the effects of recent disturbance tend to overwrite prior disturbance, mixed severity fire produces different response in plant communities than single events. Repeated high and moderate severity fire suppress tree regeneration and shift the community to shrub dominance, with recovery in pine-oak vegetation types dominated by resprouting species and by resprouting species in mixed conifer. Unburned areas contain different vegetation communities, with pine-oak forests increasing the proportions of fire-intolerant species typical of mixed conifer, as well as oaks and other potentially shrubby species growing as trees (Appendix C). Finally, we found that the proportion of the landscape dominated by shrub cover and the structure of shrub patches is influenced strongly by fire severity, with high severity burns producing the largest shrub patches as well greater area:stem ratios. Unburned areas have the lowest amount of shrub cover, and shrub patches tend to be single-stemmed, indicating that sufficiently long fire-free periods can produce monopodial trees that would otherwise grow in shrub form. The effects of antecedent disturbance can be seen in shrub patch structure, suggesting that the organizing effects of fire can persist for decades (Appendix D). Together, this body of work underscores the means by which human activities interact with the natural world to produce historic fire regimes, and the ecological communities that arise following long periods of fire regime disruption.

Namaqualand is world renowned for its mass displays of annual wildflowers occurring in highly disturbed areas. Leysera gnaphalodes is a short-lived perennial shrub that encroaches into this wildflower display, lessening the aesthetic appeal. For this reason populations of L. gnaphalodes need to be kept as small as possible. This is usually achieved by tilling the area regularly, but a less disruptive method would be preferable. Alternatives to this approach are explored. The effect of many interacting factors needed to be examined over long periods of time so that alternative management strategies could be evaluated. Ecological modelling was used as it is ideally suited to this purpose. A review of modelling and its application in ecology is given, which includes a description of the modelling process and a discussion of different types of models and their applications. It was hypothesised that grazing and low rainfall, in addition to tilling, could control the population size of L. gnaphalodes. Data was used from an eight-year study conducted to determine the effects of tilling, grazing and environmental factors on the seedbank and population size of L. gnaphalodes. A rule-based mechanistic mathematical model based on the logistic growth curve was constructed to describe the population dynamics of this species. The model-fit was evaluated using Pearson’s correlation coefficients and graphs, and it proved to be a good model. Tilling and low rainfall were both found to be effective in reducing populations of L. gnaphalodes but grazing had no reducing effect. Simulations based on the model were run to test three different basic management strategies under stochastic rainfall conditions. The management strategy, which most effectively controlled the population was to till the lands whenever the population of L. gnaphalodes reaches of exceeds a relative frequency of 45%. Multivariate statistical models were constructed to determine the effects of all of these factors on the population of L. gnaphalodes. Tilling was confirmed to be effective in reducing the population, but grazing was found to have no effect. Low rainfall was also effective in controlling the population but has the disadvantages of being out of management control and also affecting the desirable wildflowers. / Dissertation (MSc (Botany))--University of Pretoria, 2005. / Plant Science / unrestricted