Despite an abundance of archaeological material recovered from sites in Island Southeast Asia, the timing and route by which cultigens first arrived in Wallacea remains unclear. Many of the staple crops now grown on these islands were domesticated in mainland Asia, and were deliberately introduced by humans at an unknown point during the Holocene, through several possible routes. In this study, the δ 13C, δ15N and δ18O values of subfossil bones and teeth attributed to Sus celebensis and Sus scrofa are analyzed. These materials, which span the last 5160 years at Liang Bua, Flores, Indonesia are used to determine if and when there was a shift towards agricultural intensification, and whether this intensification included the integration of domesticated C4 crops. The δ13C and δ15N values of the bone and dentin collagen samples indicate an abrupt shift towards enrichment in 13C and depletion in 15N at some time between 5160 and 2750 yBP. This hints at changes in human subsistence patterns that may have included the clearing of forests, and the integration of nonendemic C4 cultigens such as foxtail millet (Setaria italica) onto the island. No statistically significant variation in the δ 18O values of the enamel carbonate samples over time is observed, suggesting that once they appeared on Flores, semidomesticated pigs became an important part of the island ecosystem, and were bred and raised on Flores instead of being continuously imported from elsewhere.

Fieldwork in the Zemplen Mountain area of north-eastern Hungary showed that there are at least eight geological sources of obsidian here, five of which have obsidian of a workable quality. There are a further three sources in the Slovak Zemplen, all of which provide workable obsidian. Sources in Central Slovakia are highly devitrified and not useable, and reported sources in Rumania had been discounted earlier (Nandris, 1975). Forty-six samples of obsidian from the Zemplen sources, and 293 pieces from 87 archaeological sites in Central and Eastern Europe, were analysed by neutron activation analysis for 15 trace and two major elements. The trace elements used included those which are geochemically likely to show the greatest variation between different obsidian sources, and which are not badly affected by devitrification and hydration of the obsidian, for example the rare earth elements. The analytical data was processed using Cluster Analysis. 242 of the archaeological samples came from Slovak sources, 22 from Hungarian sources, 9 from Lipari and 5 from Melos. In addition, 6 samples were tentatively assigned to Carpathian sources, and 9 could not be assigned to any source. Obsidian from the Zemplen Mountains was distributed up to a distance of approximately 480 km from the sources; it was used extensively in Slovakia and Hungary and reached southern Poland, Austria, Moravia, central Yugoslavia, north-east Italy and central Rumania. Obsidian use in central and eastern Europe began in the Mousterian period. The earliest pieces analysed were Aurignacian and came from Hungarian sources. Later, in the Gravettian, Slovakian sources began to be exploited and remained predominant until obsidian use declined sharply in the Later Neolithic, and Copper and Bronze Ages. The Carpathian obsidian distribution overlaps with the Liparian distribution at one site in north-east Italy. There is no evidence for an overlap with Aegean or Near Eastern sources. The rate of fall off of obsidian away from the sources suggests a down-the-line trading mechanism.

A methodology for ageing foetal and neonatal cattle is developed, involving radiographic examination of infant mandibles for early developmental stages in molariform teeth; tooth-wear methodologies are imprecise at this stage before wear commences. Known-age modern bovine foetal and neonate material are collected as a control assemblage for method development (n=73); six Neolithic to Norse era assemblages from Orkney are examined using the modified technique together with standard tooth-wear analysis and other methodologies. Foetal and died-at-birth material is diagnosed at most sites using the new technique, together with a range of other peri-natal age-groups. Ageing at this early stage is highly relevant in the diagnosis of milking as a palaeoeconomy: the accepted view is that unwanted (male) calves were slaughtered to maximise milk for human consumption, hence a surfeit of neonate calf remains, as at the study sites. The diagnosis of foetal and died-at-birth material challenges this view, suggesting that attritional causes may have contributed to deaths at this stage. Although milking was probably carried out at most of the study sites, this may have been combined with slaughter of cattle for meat in a pragmatic exploitation strategy. Literary research shows possible attritional causes of abortion and early death in calves, in particular dietary insufficiency in pregnant cows, microbial infections, and also inadequate colostrum uptake. Additionally, research is used to consider the challenges to health that early milking might have posed, to the calf as mentioned, but also to the cow, where three main health issues are highlighted: infertility, mastitis and lameness. / The attached files include the Landscape pages and appendices V and VI. Not included are the jpeg Mandible files. A cover sheet was not available.

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No / Farming transformed societies globally. Yet, despite more than a century of research, there is little consensus on the speed or completeness of this fundamental change and, consequently, on its principal drivers. For Northern Europe, the debate has often centered on the rich archaeological record of the Western Baltic, but even here it is unclear how quickly or completely people abandoned wild terrestrial and marine resources after the introduction of domesticated plants and animals at approximately 4000 calibrated years B.C. Ceramic containers are found ubiquitously on these sites and contain remarkably well-preserved lipids derived from the original use of the vessel. Reconstructing culinary practices from this ceramic record can contribute to longstanding debates concerning the origins of farming. Here we present data on the molecular and isotopic characteristics of lipids extracted from 133 ceramic vessels and 100 carbonized surface residues dating to immediately before and after the first evidence of domesticated animals and plants in the Western Baltic. The presence of specific lipid biomarkers, notably omega-(o-alkylphenyl)alkanoic acids, and the isotopic composition of individual n-alkanoic acids clearly show that a significant proportion ( approximately 20%) of ceramic vessels with lipids preserved continued to be used for processing marine and freshwater resources across the transition to agriculture in this region. Although changes in pottery use are immediately evident, our data challenge the popular notions that economies were completely transformed with the arrival of farming and that Neolithic pottery was exclusively associated with produce from domesticated animals and plants.

Traditionally archaeology has referred to the anthropic sediments accumulated around prehistoric settlements with the blanket term ‘midden’. This is now recognised as an inadequate term to describe the complex formation processes and functions represented in these sediments. This thesis reviewed the body of evidence accumulated over the past century of research into Neolithic and Bronze Age settlements on the islands of the Scottish North Atlantic seaboard and extrapolated the many occurrences of ‘midden’. Several contexts emerged for these sediments including interior floors, hearths, exterior occupational surfaces, dumped deposits, building construction materials and abandonment infill. In addition, ‘midden’ is described added to cultivated soils to form fertile anthrosols. The way in which prehistoric communities exploited this material for agriculture and construction has been described through geoarchaeological research which implied that to past communities ‘midden’ was a valuable resource. This led to the formation of a model based upon a human ecodynamics framework to hypothesise sediment formation pathways. Rescue excavation at the Links of Noltland, Westray provided an opportunity to conduct a holistic landscape and fine resolution based study of Neolithic and Bronze Age settlement to test this model. The research incorporated auger survey, archaeological and geoarchaeological excavation, thin section micromorphology and SEM EDX analyses. Sediments identified in literature review and recovered from the field site were described using this toolkit and set within a cultural and environmental context. Results demonstrate that anthropic materials were incorporated into all contexts examined. Discrete burning and maintenance activities were found to have taken place during the gradual accumulation of open-air anthropic sediments whilst incorporation of fuel residues and hearth waste into floors lead to the gradual formation of ‘living floors’ inside structures. An unexpected discovery was evidence of animal penning within late Neolithic/Early Bronze Age settlement and the in situ burning of stabling waste. Three types iv | P a g e of land management strategy which relied upon the input of anthropic sediments were evidenced and the range and extent of anthropic inclusions in the landscape recorded. Spatial interpolation of auger survey data utilised a new sub-surface modelling technique being developed by the British Geological Survey to explore soil stratigraphic relationships in 3D. SEM EDX analysis supported micromorphological analysis providing chemical data for discrete inclusions and assisting in the identification of herbivore dun ash and the Orcadian funerary product ‘cramp’. SEM EDX analysis was also applied to fine organo-mineral material for statistical testing of nutrient loadings across context groups. It was found that anthropic sediments were enriched in macro and intermediate plant nutrients Mg, P, K, S and Ca compared to geological controls, and the application of anthropic material to cultivated soils improved soil fertility for the three observed land management practices. The versatility of anthropic sediments was explored through discussion of context groups based upon the results of this research and the potential significance of this material to prehistoric communities is explored.

Über die Ätiologie und Epidemiologie der Erkrankungen des Respirationstraktes im Frühneolithikum Mitteleuropas gibt es bislang keine umfassenden Studien. Die Häufigkeit, mögliche Geschlechts- und Altersunterschiede, Populationsunterschiede und Erkenntnisse über mögliche Ursachen und auslösende Faktoren der Atemwegserkrankungen zur Zeit der Bandkeramik sind von besonderem Interesse. Zudem soll aufgezeigt werden wie wichtig es ist, alle Strukturen zu untersuchen, die den „knöchernen Respirationstrakt“ repräsentieren. Von den ersten Ackerbauern und Viehzüchtern aus Wandersleben (Thüringen, Kreis Gotha) lagen insgesamt 112 erwachsene Individuen zur Untersuchung vor. Rippen und Schädel wurden mit paläopathologischen Methoden untersucht. Diese waren neben einer makroskopischen Begutachtung der Knochen, röntgenologische, endoskopische, lupenmikroskopische, lichtmikroskopische und rasterelektronenmikroskopische Untersuchungen, um eine verlässliche Diagnosestellung zu gewährleisten. 100 % (n = 71/71) der befundbaren Individuen zeigten Spuren chronischer Erkrankungen im Bereich der knöchernen Strukturen, die die oberen Atemwege umgeben. Bei 76,8% (n = 53/69) der Individuen mit befundbaren Rippen konnten Spuren von chronischen Erkrankungen aufgezeigt werden. Signifikante Geschlechts- oder Altersunterschiede bestehen insgesamt nicht. Bei der linearbandkeramischen Population aus Wandersleben ist davon auszugehen, dass es sich in vielen Fällen um Chronifizierungen von Erkältungskrankheiten und um die Folgen einer vergleichbar schlechten Luftqualität hauptsächlich im Haus handelt. Dazu haben die damaligen Lebensumstände, vor allem aber die sesshafte Lebensweise und Wirtschaftszweige mit Ackerbau und Viehzucht, maßgeblich beigetragen.

The complex demographic changes that underlie the expansion of anatomically modern humans out of Africa have important consequences on the dynamics of natural selection and our ability to detect it. In this thesis, I aimed to refine our knowledge on human population history using ancient genomes, and then used a climate-informed, spatially explicit framework to explore the interplay between complex demographies and selection. I first analysed a high-coverage genome from Upper Palaeolithic Romania from ~37.8 kya, and demonstrated an early diversification of multiple lineages shortly after the out-of-Africa expansion (Chapter 2). I then investigated Late Upper Palaeolithic (~13.3ky old) and Mesolithic (~9.7 ky old) samples from the Caucasus and a Late Upper Palaeolithic (~13.7ky old) sample from Western Europe, and found that these two groups belong to distinct lineages that also diverged shortly after the out of Africa, ~45-60 ky ago (Chapter 3). Finally, I used East Asian samples from ~7.7ky ago to show that there has been a greater degree of genetic continuity in this region compared to Europe (Chapter 4). In the second part of my thesis, I used a climate-informed, spatially explicit demographic model that captures the out-of-Africa expansion to explore natural selection. I first investigated whether the model can represent the confounding effect of demography on selection statistics, when applied to neutral part of the genome (Chapter 5). Whilst the overlap between different selection statistics was somewhat underestimated by the model, the relationship between signals from different populations is generally well-captured. I then modelled natural selection in the same framework and investigated the spatial distribution of two genetic variants associated with a protective effect against malaria, sickle-cell anaemia and β⁰ thalassemia (Chapter 6). I found that although this model can reproduce the disjoint ranges of different variants typical of the former, it is incompatible with overlapping distributions characteristic of the latter. Furthermore, our model is compatible with the inferred single origin of sickle-cell disease in most regions, but it can not reproduce the presence of this disorder in India without long-distance migrations.

In South Sweden the third millennium BC is characterised by coastal settlements of marine hunter-gatherers known as the Pitted Ware culture, and inland settlements of the Battle Axe culture. This thesis outlines the history of research of the Middle Neolithic B in general and that of the pottery and burial practices in particular. Material culture must be understood as the result of both conscious preferences and embodied practices: technology can be deliberately cultural just as style can be un-selfconscious routine. Anthropological and ethnoarchaeological research into craft and the transmission of learning in traditional societies shows how archaeologists must take into consideration the interdependence of mind and body when interpreting style, technology and change in prehistory. The pottery crafts of the Pitted Ware and Battle Axe cultures were not just fundamentally different technologically, but even more so in the attitudes toward authority, tradition, variation and the social role of the potter in the community. The Battle Axe beakers represent a wholly new chaîne opératoire, probably introduced by a small group of relocated Beaker potters at the beginning of the period. The different attitudes toward living bodies is highlighted further in the attitudes toward the dead bodies. In the mortuary ritual the Battle Axe culture was intent upon the creation and control of a perfect body which acted as a representative of the idealised notion of what it was to belong to the community. This focus upon completeness, continuation and control is echoed in the making of beakers using the ground up remains of old vessels as temper. In contrast, the Pitted Ware culture people broke the bodies of the dead by defleshing, removal of body parts, cremation, sorting, dispersal and/or reburial of the bones on the settlements. The individuality of the living body was destroyed leaving the durable but depersonalised bones to be returned to the joint collective of the ancestors. Just as the bodies were fragmented so were the pots, sherds and bases being deposited in large quantities on the settlements and occasionally in graves. Some of the pots were also tempered with burnt and crushed bones. At the end of the Middle Neolithic the material and human remains show evidence of a growing effort to find a common ground in the two societies through sharing certain mortuary rituals and making beakers with a mix of both traditions, stylistically and technologically.

Genf, Universit́e, Diss.