Previous biostratigraphic studies in the Indian Ocean, equatorial Pacific, Atlantic Ocean and the Caribbean Sea document the temporal disappearance of Reticulofenestra pseudoumbilicus in the late Miocene (Tortonian age). This interval has been subsequently defined as the R. pseudoumbilicus paracme. While the cause of the event remains unknown, the geographical extent and global synchronicity of the interval represent an open subject for research. Answers to these questions could help establish the biostratigraphic reliability of the interval and its use for global correlation. In this study, 49 deep-sea drilling sites spread across all major ocean basins were compiled into a biostratigraphic database and studied to evaluate the global extent and timing of the R. pseudoumbilicus paracme. To ensure accurate comparison of the studied sites, the timing of the event was estimated at all sites using the age estimates generated through Undatable MatLab software, which made use of established nannofossil bioevents as age-depth tie points for the analysis. Ages of all nannofossil bioevents were updated to the latest geological time scale. This systematic analysis reveals a prominent distribution of the paracme in the tropical and subtropical regions. The timing of the event in the tropical region appears to be synchronous, with the base around 8.8 Ma and the top of the event around 7.2 Ma. Sites in the equatorial Pacific, central Atlantic and the Indian Oceans record a similar duration of the event. The paracme had a shorter duration towards the southern higher latitudes. Although timings in southern high latitudes are diachronous compared to the tropics, the duration is consistent between sites (Sites 1088 and 1264) in the southern South Atlantic, thereby establishing a new time frame for the event in the region. This study narrows down on ODP Site 806 for a critical evaluation of the nannofossil assemblages, accumulation rates and Reticulofenestra size variation in the late Miocene, including the paracme. Thirty-four samples were collected and analysed for absolute abundance and nannofossil accumulation rates (NAR). Absolute abundance and NAR reveal three distinct stages, herein described as generally low, high, and “crashed” export production of nannofossils. The period of high NAR is characteristic of the late Miocene biogenic bloom interval (ca. 9–4.4 Ma). Importantly, the R. pseudoumbilicus paracme does not only coincide with the start of the biogenic bloom but it was also associated with remarkable changes in Reticulofenestra assemblages. Just before the base of the paracme, all except the smallest (&lt;3 µm in coccolith size) Reticulofenestra groups experienced a sharp decline in abundance. Subsequently, the relative abundance of the Reticulofenestra morphospecies indicates the dominance of bloom-forming small and medium Reticulofenestra during the late Miocene biogenic bloom interval. The end of the biogenic bloom interval is concurrent with a decrease in the contribution of small Reticulofenestra. The exact biotic and abiotic processes that led to these major shifts in plankton composition are yet to be discovered, but the findings presented herein show that the mechanisms behind the temporary disappearance of R. pseudoumbilicus in the tropics most likely involved the interplay between a broad range of ecological changes and evolutionary processes affecting the Reticulofenestra assemblages. / <p>The work for this thesis was financially supported by the Erasmus Mundus Joint Master Degree PANGEAprogramme.</p>

No description available.

Paleomagnetic directions have been obtained from 190 Middle Miocene (12-20 Ma) mafic volcanic flows in 16 mountain ranges in the Mojave-Sonora desert region of western Arizona and southeastern California. These flows generally postdate Early Miocene tectonic deformation accommodated by low-angle normal faults but predate high-angle normal faulting in the region. After detailed magnetic cleaning experiments, 179 flows yielded characteristic thermal remanent magnetism (TRM) directions. Because of the episodic nature of basaltic volcanism in this region, the 179 flows yield only 65 time-distinct virtual geomagnetic poles (VGPs). The angular dispersion of the VGPs is consistent with the angular dispersion expected for a data set that has adequately averaged geomagnetic secular variation. The paleomagnetic pole calculated from the 65 cooling unit VGPs is located at 85.5°N, 108.9°E within a 4.4° circle of 95% confidence. This pole is statistically indistinguishable (at 95% confidence) from reference poles calculated from similar-age rocks in stable North America and from a paleomagnetic pole calculated from similar-age rocks in Baja and southern California. From the coincidence of paleomagnetic poles from the Mojave-Sonora and adjacent areas, we can conclude that: (1) vertical-axis tectonic rotations have not accompanied high-angle normal faulting in this region; (2) there has been no latitudinal transport of the region since 12-20 Ma; and (3) long-term nondipole components of the Miocene geomagnetic field probably were no larger than those of the recent (0-5 Ma) geomagnetic field. In contrast, paleomagnetic data of other workers indicate vertical-axis rotations of similar-age rocks in the Transverse Ranges, the Eastern Transverse Ranges, and the Mojave Block. We speculate that a major discontinuity in the vicinity of the southeastward projection of the Death Valley Fault Zone separates western areas affected by vertical-axis rotations from eastern areas that have not experienced such rotations.

In my diploma thesis I observed microbial activity in Miocene sediments collected at two different depths - 30 and 150 meters - in the "Družba" open-mine pit, which is located in Sokolov Brown Coal Basin. There were three different types of sediment: lamellar, amorphous and transitional. Samples were obtained under sterile conditions and following treatments were applied to the transitional sediment: addition of glucose, wetting and freezing. The aim of my study was to observe the activity of the original microflora at various depths and the impact of environmental factors that can affect the quarried claystones after being put on heaps. Microbial activity was evaluated by measuring released CO2 (titration). Obtained values of microbial activity did not show any differences between the respiration of sediments of different ages (depths), but differences between different types of sediments were significant, the highest values were found in lamellar sediment. Affection of samples then showed biota activity response suggesting that microbial communities at lower depths are limited by nutrients. Because the application of glucose did not show any noteworthy effect, we can assume that these communities are specialize on present fossil organic matter they are able to decompose, which is important especially in...

Clays and clay mineral distribution studies are important for understanding the geological history of the Gulf of Mexico Basin, but few studies document any subsurface clay mineral distribution in the Gulf of Mexico. Shale samples from nine wells (30 samples) in the South Timbalier protraction were selected near known paleontological markers identifying the Miocene, Pliocene, and Pleistocene boundaries. Bulk mineralogy of each sample, determined by XRD, is primarily mixed-layer smectite and illite with a minor amount of kaolinite. The mixed-layer mineralogies are end-member smectite, mixed-layer smectite, mixed-layer illite, and end-member illite. These clay mineral fractions do not correlate with age. The illite mixed layer percentage correlates with depth, but the correlation decreases when depth is converted to temperature. However, the illite mixed layer fraction does not exhibit a strong correlation in this multiwell study when compared to a single well study in Ship Shoal using identical methods (Totten et al., 2002).

Three kinds of processes are known to modify the geographical spatial arrangement of organisms: dispersal, extinction and vicariance. The Southern Hemisphere has an intriguing and complicated geological history that provides an ideal backdrop to study these processes. This thesis focuses on three historical events that illustrate these processes: the proposed marine inundation of New Zealand in the Oligocene, the asteroid impact at the K – Pg boundary, and the continental breakup of Gondwana. It investigates what impact these events had on species diversification by studying the phylogenetic relationships of two groups of taxa – the family Anostostomatidae (insects), and Neoaves (birds). Anostostomatidae were studied in relation to the Oligocene drowning and the break up of Gondwana as they have a wide southern distribution, found on all “Gondwanan” fragments with the exception of Antarctica, and are thought represent an ancient lineage that predates the Gondwanan breakup. Birds, in particular Neoaves, were studied in relation to the asteroid impact at the K – Pg boundary. Although birds are mobile and many circumnavigate the globe between seasons, they are suggested to have originated in the Southern Hemisphere in Gondwanan times, and subsequently undergone range expansion and diversification around the world. In order to address the relationship (if any) between modern biotic diversity and historical geological events, phylogenetic relationships were determined and where possible, molecular clock analysis carried out. Timing information provided by molecular clock analysis is important as it enables distinction between opposing hypotheses such as vicariance and dispersal. In Chapter Two, the phylogenetic relationships within the family Anostostomatidae are investigated. One of the most controversial times in New Zealand’s geological history is during the Oligocene. Some suggest that the lack of fossils and evidence for recent dispersal of numerous taxa support the notion that all modern biota reached the region during the last 25 million years. Anostostomatidae were chosen as they represent a group of insects that are thought to be ancient and there is little published data in the literature. Previous studies focused on the relationships within Hemideina and Deinacrida suggesting that these groups diversified in the early Miocene. The data presented here are from mitochondrial (COI and 12S) and nuclear (18S and 28S) sequences. Molecular dating using a relaxed clock as implemented in BEAST suggest that in fact some lineages were present at or shortly after continental breakup and could have survived throughout this turbulent time. As there were no definitive fossils to use for calibration points, geological events were used as calibration points for the molecular clock. Mutation rates obtained from the different analyses were compared to those published for other insects in an attempt to identify the most likely model. Both maximum likelihood and Bayesian analyses support the presence of three distinct ecological groups in New Zealand; Hemiandrus (ground weta), Anisoura/Motuweta (tusked weta) and Hemideina–Deinacrida (tree–giant weta). With regards to their Australasian relatives (taxa from Australia and New Caledonia) it appears that the family is divided with the most northern New Zealand taxa (tusked weta) more closely related to New Caledonian taxa while all other New Zealand taxa are more closely related to Australian taxa. There does not appear to be any link between the Australian and New Caledonian taxa studied here. Results should be viewed with caution however as an increased mutation rate was observed in the New Caledonian-tusked weta lineage, something future studies will have to address. Chapter Three presents new sequence data and phylogenetic analyses that go towards resolving the apparent basal polytomy of neoavian birds. This chapter includes analyses carried out on previously published data with the addition of nine new mitochondrial genomes. My contribution to this larger project was to perform the phylogenetic analysis and to sequence three of the nine mitochondrial genomes. The genomes I sequenced were the Southern Hemisphere species: dollar bird (Eurystomus orientalis), Owlet nightjar (Aegotheles cristatus cristatus) and great potoo (Nyctibius grandis). The inclusion of these nine new genomes allows assessment and comparison of the six hypothesised groups reported in Cracraft (2001). First an improved conditional down-weighting technique is described reducing noise relative to signal, which is important for resolving deeper divergences. Second, a formula is presented for calculating probabilities of finding predefined groupings in the optimal tree. Maximum likelihood and Bayesian based phylogenetic analyses were carried out and in addition, dating using a relaxed molecular clock was performed in BEAST. Results suggested that the six groups suggested by Cracraft (2001) represent robust lineages. The results suggested that one group, the owls, are more closely related to other raptors, particularly accipitrids (buzzards/eagles) and the osprey rather than the Caprimulgiformes, which could indicate morphological convergent evolution. In addition, a group termed shorebirds appears to be distinct from the large group referred to as ‘Conglomerati’ to which previous publications have suggested they belong. The ‘Conglomerati’ is the least well studied group and may actually comprise of at least three subgroups (as suggested by Cracraft). Within the three suggested groups, Cracraft grouped shorebirds with pigeons and sandgrouse, neither of which (pigeons or sandgrouse) were analysed here. So although the shorebirds are at least close to the ‘Conglomerati’ and may be within that group, their exact position is still not clear. The molecular dating reported here utilised two fossil calibrations (Vegavis and Waimanu), for which there is relatively little dispute as to age or the lineage to which they belong. Calibrations resulting from BEAST analyses suggest that at least 12 distinct lineages were present prior to the K – Pg boundary, a finding supported by previous studies. Robust phylogenies will allow future studies to investigate not only the relationships within Neoaves, but look more closely at the biological and ecological evolution of the group. Chapter Four for the first time investigates whether the phylogenetic relationships within the family Anostostomatidae follow the conventionally accepted order and timing of Gondwanan breakup. Following the initial restults for taxa studied in Australasia (Chapter Two) an attempt to resolve family relationships in a wider spatial (geographic) context was carried out to determine if Australasian taxa are monophyletic when other members of the family are included. Again both maximum likelihood and Bayesian phylogenetic analyses were carried out on both mitochondrial (COI and 12S) and nuclear (18S and 28S) sequences. In this chapter, datasets included samples from across the geographic range of Anostostomatidae (South Africa, Madagascar, South America, Australia, New Caledonia and New Zealand), and two clades were observed, congruent with earlier findings. Sequence divergence within geographic regions was found to be relatively high in the mitochondrial genes (COI and 12S) while low in the nuclear ribosomal RNA genes (18S and 28S) as expected given their relative mutation rates. Under the vicariance paradigm, phylogenetic relationships should follow the order of continental breakup, but this was not found. Further, if dispersal and colonisation were continuous, no geographic substructure is expected, however distinct geographic substructure within clades was consistently observed. This interesting phylogenetic pattern may be a case of convergent evolution or paraphyletic sampling which highlights taxonomic issues of the group. Future studies need to include not only molecular data but information on morphology, ecology and behaviour along with the implementation of biogeographic programs that can test alternative hypotheses (such as dispersal and vicariance) directly. Also, the inclusion of the recently reported fossil from the subfamily Euclydesinae (Martins-Neto 2007) should allow for more accurate date estimates within the family. Taken as a whole the results presented in this thesis suggest that microevolutionary processes are sufficient to explain modern diversity without the need to invoke abiotic events. The three cases investigated here - marine inundation, asteroid impact and continental drift - all appear to have had only a limited effect on the diversity of taxa studied. To reach even stronger conclusions future studies should incorporate different data (for instance nuclear genes, intron position, and genome structure) and use biogeographic software capable of including ecological, morphological and habitat information.

Four multicyclic complex calderas and smaller ignimbrite shields located within the Altiplano Puna Volcanic Complex of the Central Andes (APVC) erupted 13000 km�� of magma within the last 11 Ma. One of the largest and most complex of these is the Cerro Guacha Caldera. Ar-Ar age determinations and paleomagnetic directions suggest that the Cerro Guacha Caldera was formed by two major eruptions, caldera collapse, resurgence cycles and several smaller eruptions. Two major ignimbrites (> 600 km��) are found with ������Ar-�����Ar from biotites and sanidines of 5.65 �� 0.01Ma for the 1300 km�� (magma volume) Guacha ignimbrite and 3.49 �� 0.01Ma for the 800 km�� Tara Ignimbrite. The last major eruption occurred on the western flank producing the 1.72 �� 0.02 Ma Puripica Chico Ignimbrite with a volume of approximately 10 km��. Characteristic remanent magnetization data (ChRM) for these ignimbrites show that the Guacha has reverse polarity, while the Tara is normally polarized and the magnetic fingerprints have allowed their current full extents to be identified. A conspicuous lineament of volcanic structures in the eastern part of the caldera, bordering a caldera moat, filled out welded ignimbrites and sedimentary lacustrine sequences suggest an earlier 60x40 km outer collapse associated with the Guacha explosive episode. A central graben formed on the Guacha welded ignimbrite is related to a first episode of resurgence. Evidence of a second 30 x15 km inner collapse includes offset of welded Guacha ignimbrites and alignment of lava domes associated with the Tara ignimbrite. A second resurgence episode is suggested by the presence of an uplifted central block consisting primarily of welded Tara ignimbrite. As a whole the three ignimbrites (Guacha, Tara and Puripica Chico) share the same petrological and geochemical characteristics: high-K series, compositional ranges from dacite to rhyolite, with andesitic members present as lavas (for the Guacha and Puripica Chico Ignimbrites) and as pumices (for the Tara Ignimbrite). Highest silica content is found in the Chajnantor dome. Rayleigh modeling for Ba, Rb and Sr suggests at least 60% of crystal fractionation to account for the compositional variation between the Guacha andesite and the Chajnantor dome. Dy/Hb ratio increases with time from the Guacha andesite to the Negreal andesite suggesting stabilization of garnet owing to crustal thickening. Fe-Ti exchange geothermometry for the Tara Ignimbrite yielded log fO��� values ranging from -13.06 to -13.38 and temperatures of 714�� to 801��C. Amphibole geobarometry yielded pressures ranging from 150 to 180 MPa equivalent to 5.3 and 6.4 km depth respectively for the Tara Ignimbrite; the pressures range between 133 to 242 MPa, equivalent to 5.0 to 9.2 km depth for the Guacha Ignimbrite. The zircon saturation method yielded saturation temperatures of 716�� and 705��C for the Guacha and Chajnantor dome respectively and 784��C for the Tara Ignimbrite. The zircon crystallization range for the magmas of the Cerro Guacha Caldera is 1.25 Ma for the Guacha Ignimbrite; 1.09 Ma for the Puripica Chico Ignimbrite and 0.95 Ma for the Tara Ignimbrite. Recycling of antecrystic zircons within the caldera magmas is continuos through time. / Graduation date: 2012

The Miocene Nullarbor Limestone is the most recent formation in the Cenozoic Eucla Group and was deposited in the Eucla Basin, southern Australia, at ~38°S paleolatitude during the early to middle Miocene. The rocks form the modern surface of the vast, karsted Nullarbor Plain. Older Eucla Group marine carbonates (Eocene-earliest Miocene) are cool-water in nature and dominated by bryozoans and echinoderms. The Nullarbor Limestone is subtropical in composition and rich in coralline algae (rhodoliths and articulated types), large and small benthic foraminifera and molluscs. Diverse zooxanthellate corals are also present but not numerous. Deposition is interpreted to have taken place in three main paleoenvironments: rhodolith gravels, seagrass banks, and open seafloors. The Southern Ocean extended inboard ~450 km from the shelf edge during Nullarbor Limestone deposition. Interpreted paleodepths ranged from the top to the base of the photic zone, implying a small slope over a wide shelf. The Miocene Eucla platform is therefore interpreted to have been epeiric in nature. Paleoenvironment distribution is explained using epeiric platform sedimentation patterns and comparisons with modern environments. Open seafloor environments, the deepest settings, are thought to have been below fair-weather wave base. Rhodolith gravels accumulated at intermediate depths, where waves frequently swept the seafloor. Seagrass banks developed in the shallowest waters farthest inboard, where wave energy had been largely dissipated. Diverse corals, large benthic foraminifera and micrite envelopes inboard and in the western part of the basin support the notion of paleotemperatures generally above 20°C, the upper limit of subtropical carbonate accumulation. Although deposition occurred during the Miocene Climatic Optimum, a simple overall temperature increase cannot completely account for the subtropical nature of these sediments at mid-latitudes. Tropical components decrease from west to east, implying a temperature gradient, probably due to the warm proto-Leeuwin Current. Thus, these subtropical carbonates were deposited at mid-latitudes and their presence did not simply reflect a change in global climate. / Thesis (Master, Geological Sciences & Geological Engineering) -- Queen's University, 2010-12-23 16:05:47.981

xiii, 198 p. : ill. (some col.) / Whether or not a causal relationship exists between climate and mammal body size is one of the longest-standing and most intractable questions in ecology. The classic model of body size evolution (Bergmann's Rule) holds that body size is driven by temperature, but more recent hypotheses have suggested that other climatic variables or biotic interactions may play a more important role. The use of paleoecological data to address this question allows variables that are tightly correlated in modern ecosystems to be teased apart and allows body size patterns to be observed through time, adding an extra dimension to analyses. This dissertation details the findings of two paleoecological tests of Bergmann's Rule in the Oligo-Miocene (30-5 Ma), one tracking body size and climate through time in the northwestern United States and another tracking geographic body size trends through time along the west coast of North America. In both cases, body size was analyzed in three representative families of mammals: equids, canids, and sciurids. Such large-scale analyses are dependent on fossils that can be placed in a reliable taxonomic, geologic, and temporal context, and this dissertation also focuses on a reevaluation of the canid fauna of Oregon's Juntura Formation that places a critically important Late Miocene carnivore fauna in just such a context. Two genera of canids - Epicyon and Carpocyon - are described from the fauna for the first time, with important implications for regional biostratigraphy. The body size analyses show no consistent relationship between body size and any climatic variable. Further, body size patterns vary widely between taxa at several levels, suggesting that one universal driver of body size evolution does not exist. Not only is there no evidence for Bergmann's Rule in Oligo-Miocene mammals, but comparative analyses of geographic body size patterns in the modern genera Odocoileus, Canis, and Spermophilus fail to show the latitudinal gradients upon which Bergmann's Rule is predicated. The apparent existence of such trends in some taxa may be the result of anthropogenic extirpation at low latitudes, further underscoring the importance of including paleontological data when formulating models predicting the response of biotic variables to environmental change. / Committee in charge: Dr. Samantha Hopkins， Chair； Dr. Gregory Retallack， Member； Dr. Rebecca Dorsey， Member； Dr. Stephen Frost， Outside Member

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No. of bitstreams: 1 Dissertação - Eliezer Senna Gonçalves Júnior.pdf: 13704881 bytes, checksum: 1d71d2269743a9193eaff46fd6fd9f0b (MD5) Previous issue date: 2013-08-30 / OUTRAS / In Careiro-da-Várzea, Careiro-Castanho, Manaquiri and Autazes areas, near Manaus, three asymetric quaternary fluvial terraces levels overlains the Alter do Chão (Cretaceous) and Novo Remanso (Miocene) siliciclastic sediments. Locally, these geological basement outcrops as an elongated stripe about 150 kilometers length toward NE-SW, in heights between 100 and 50 meters sustained by three chemically distinct lateritic paleosoils levels. The quaternary fluvial terraces developed between 240.000 and 2.000 years BP extends more than 100 kilometers between 80 e 10 meters heights composing the Amazonas-Solimões Fluvial System alluvial plain on the studied reach. Generally, these deposits present scroll-bars morphology, with low drainages density, characterized by sparse plain secondary channels and rounded lakes. Internally, they are constituted by pairs of layers made of sand and mud (silt and clay) forming Inclined Heterolitic Stratification (IHS) within the meander point bars. The TS deposits presents pink to reddish colors and shows pedogenesis features, while the TI and TInf deposits tends to present brownish to greyish colors due the rich organic matter and bioturbation features. The TS deposits occupies de south-central portion of the study area extending around 100 kilometers being constituted mainly of mud deposits, commonly used by the locals to brick and tiles manufacture. The TI and TInf deposits occur as narrow and elongated stripes of maximum 30 kilometers width, composing the active floodplain of the actual Solimões-Amazonas system. The point bars with IHS deposits, the terraces asymmetric distribution and the register of paleochannels features near the mouth of Purus River, suggests the predominance of a meander pattern in the study area between 240.000 and 6.000 years BP. The Solimões-Amazonas Fluvial System developed the last 6.000 years to an anastomosed-anabranching stable river related to the rise of sea level during the Late Holocene and the climatic changes that boosted the development of the rainforest, furthering the stabilization of margins on this fluvial system. / Na região entre os municípios do Careiro-da-Várzea, Careiro-Castanho, Manaquiri e Autazes, ao sul de Manaus, ocorrem três níveis de terraços fluviais quaternários, de distribuição assimétrica, desenvolvidos sobre as rochas siliciclásticas (arenitos, pelitos e conglomerados) das formações Alter do Chão (Cretáceo) e Novo Remanso (Mioceno), que compõem o embasamento geológico desta região. Localmente, este embasamento aflora como uma faixa alongada de direção NE-SW, com cerca de 150 km de comprimento e 15 km de largura, entre cotas de 100 e 50 metros sustentadas por três níveis de paleossolos lateríticos quimicamente distintos. Os terraços fluviais desenvolvidos entre 240.000 e 2.000 anos AP, se estendem por mais de 100 km e situam-se em cotas variando entre 80 e 10 metros de altitude, constituindo as planícies aluviais do sistema fluvial Solimões-Amazonas, no trecho estudado. Em geral, apresentam morfologia deposicional de barras de acresção lateral, com baixa densidade de drenagem, caracterizada por esparsos canais secundários de planície, além de lagos de vários tamanhos e formas. Internamente, os depósitos são constituídos principalmente de areia e lama (silte e argila), em diferentes proporções, que compõem os pares de estratificação heterolítica inclinada (EHI) das barras em pontal. Os depósitos do Terraço Superior (TS) apresentam coloração rosada a avermelhada e exibem feições de pedogênese, enquanto os depósitos dos Terraços Intermediário (TI) e Inferior (TInf) apresentam coloração marrom a cinza médio, com feições de bioturbação, sendo ricos em matéria orgânica. O TS ocupa a porção centro-sul da área de estudo, com cerca de 100 km de extensão, sendo constituído principalmente por depósitos finos (silte e argila), usados na fabricação da cerâmica vermelha (tijolos e telhas) pelas comunidades locais. Os TI e TInf ocorrem como faixas estreitas e alongadas de até 30 km de extensão, que compõem a planície ativa do sistema fluvial Solimões-Amazonas, estando, portanto, sujeitos as inundações periódicas. As barras em pontal com EHI, a distribuição assimétrica dos terraços, o registro de paleocanais na planície aluvial na região da foz do Rio Purus são evidências da predominância do estilo fluvial meandrante, com migração geral para norte, entre 240.000 e 6.000 anos AP. No Holoceno Tardio o sistema fluvial Solimões-Amazonas evoluiu para um estilo anastomosado-anabranching relativamente estável em resposta ao aumento do nível do mar que barrou sua foz e ao aumento da umidade que proporcionou o desenvolvimento vegetal e subsequentemente a fitoestabilização das margens e aumento da carga de finos em suspensão.