The Mountain Hemlock (MH) zone includes all subalpine forests along British Columbia’s coast. It occurs at elevations where most precipitation falls as snow and the growing season is less than 4 months long. The zone includes the continuous forest of the forested subzones and the tree islands of the parkland subzones (Figure 1). Old-growth stands are populated by mountain hemlock, Pacific silver fir, and Alaska yellow-cedar, and are among the least-disturbed ecosystems in the world. Canopy trees grow slowly and are commonly older than 600 years, while some Alaska yellow-cedars may be up to 2000 years old. Understanding regeneration patterns in the MH zone has become increasingly important as logging continues towards higher elevations of the zone where snowpacks are deeper.

Magistro darbe tiriama Nemuno kilpų regioninio parko Punios šilo sumedėjusią augaliją. Darbo objektas – Punios šilas bei jame esantys gamtinis rezervatas ir botaninis – zoologinis draustinis. Dabar Punios šilas – viena svarbiausių Nemuno kilpų regioninio parko dalių. Darbo tikslas – ekologinis Nemuno kilpų regioninio parko Punios šilo sumedėjusios augalijos vertinimas. Darbo metodika – Išanalizavus literatūrinę medžiagą ir remiantis Nemuno kilpų regioniniame parke esančiame Punios šile atliktais sumedėjusios augalijos tyrimais, stebėjimais nustatytas medžių rūšių išsidėstymas, jų pasiskirstymas kvartaluose. Buvo nustatyta medynų ekologinė būklė. Tyrimai buvo vykdomi visuose 48 Punios šilo kvartaluose. Kiekvienam kvartalui sudaryta atskira lentelė, iš kurios atsispindi bendras užimamas kvartalo plotas hektarais, sklypų skaičius, medynų sudėtis, būdinga augavietė, medynų amžiaus vidurkis, skalsumas, medynų tūris. Darbo rezultatai – nustatyta esama sumedėjusios augalijos rūšinė sudėtis, medynų įvairovė, ekologinė būklė. Punios šilo gamtinių objektų būklė yra gera. Pagal medynų charakteristiką Punios šile pušynai užima didžiausią plotą, eglynai per puse mažesnį, o ąžuolynai ir beržynai beveik vienodą plotą. / Post–graduate work examined the Nemunas loops regional park the forest of Punia woody vegetation. Object of the work - forest of Punia with nature reserve and botanical - zoological preserve that are in it. Now forest of Punia - one of the most important part of the Nemunas Loops Regional Park. Aim of the work - the ecological assessment of Nemunas loops regional park forest of Punia woody vegetation. Method of the work - analysis of literary material and on the basis of the Nemunas Loops Regional Park located in forest of Punia for the woody vegetation studies, the observations set out in the distribution of tree species, their distribution areas. It was found the ecological status of the woods. Studies were conducted in all 48 blocks of forest of Punia. For different block was made separate table, which is reflected in the total area in hectares occupied by block, parcel number, stand composition, characterized place of growth, average age of stands, the volume of stands. Results of the work - the existing woody vegetation species composition, stand diversity, the ecological status was set. Forest of Punia natural objects are in good condition. Under the stands testimonial pine forests occupy the largest area of the forest of Punia, fir - a half lower, and the oak woods, and birch forest takes near the same area.

Thesis (MScFor)--Stellenbosch University, 2013. / ENGLISH ABSTRACT: The practice of combining row and selective thinning in commercial pine plantation silviculture carries the risk of unwanted irregularities in tree distribution within the stand. This situation is aggravated with poor tree selection during marking. The potential consequences of poor tree selection are gaps created along row removals, which are necessary for access to harvesting operations. These gaps lead to spatially asymmetric growing space among adjacent trees. The effect of irregular stand structures on tree morphology and growth are investigated in this study, and are based on two stands of Pinus patula, (Schiede ex Schlechtendal et Cham.) in Langeni plantation, South Africa. This study focuses on two aspects. Firstly, a comparison between trees grown in all-sided and one-sided spatial competition situations in order to assess if there are differences in growth and selected quality parameters. Secondly, the mitigation of irregular structures using a simulation based study on changing the planting geometry in order to investigate the effect on harvesting in terms of stand impact, simulated harvesting productivity and harvesting system costs. Results showed that trees grown in an irregular competitive status have significantly larger crown diameters, crown lengths, longer and thicker branches, disproportionately one sided crown growth and a reduction in space-use efficiency. Simulations indicated that changing planting geometry from the current 2.7m x 2.7m to 2.3m x 3.1m and 2.4m x 3m would result in up to a 20% reduction of machine trail length and fewer rows being removed for machine access. The simulation of harvesting thinnings showed that various planting geometry alternatives increased harvesting productivity by 10% to 20% and reduced overall thinning harvesting cost by up to 11%. This study successfully investigated the factors that potentially negatively affect saw timber quality and volume production of the stand at final felling. It also illustrated the applicability of simulation methods for testing harvesting scenarios and developing economically viable alternatives. / AFRIKAANSE OPSOMMING: Die praktiese kombinasie van ryuitdunning en seleksiedunning in kommersiële denneplantasies dra die risiko van ongewensde onreelmatighede in die verspreiding van bome in die opstand. Hierdie situasie word vererger deur swak boomseleksie tydens die merk van dunnings. Die potensiële gevolge van swak boomseleksie is die ontstaan van onreelmatige gapings tussen boomkrone, veral langs die rydunnings, wat nodig is vir toegang tydens die ontginning van die hout. Dit lei daartoe dat die bome langs die dunningsrye asimmetriese ruimtes het om in te groei. Die effek van onreelmatige opstandstrukture op boom-morfologie en -groei word in hierdie studie ondersoek in twee Pinus patula, (Schiede ex Schlechtendal et Cham.) vakke te Langeni plantasie, Suid-afrika. In die studie word daar gefokus op twee aspekte. Eerstens word bome wat onder toestande van eweredige ruimetlike kompetisie groei vergelyk met die wat onder toestande van eensydige ruimtelike kompetisie groei om sodoende vas te stel of daar verskille is in die groeipatroon aan die hand van geselekteerde gehalteparameters. Tweedens word daar gefokus op die verbetering van onreelmatige opstandstrukture deur gebruik te maak van ’n simulasie-gebasseerde studie om veranderinge in die aanplantingsgeometrie te ondersoek met die doel om die effek van plantspasieering op ontginningsimpakte, gesimuleerde ontginningsproduktiwiteit en -sisteem koste te bepaal. Die resultate het getoon dat bome wat onder toestande van onreelmatige spasieering en kompetisie groei krone met groter deursnee asook langer lengtes ontwikkel, langer en dikker takke het, disproporsionele, eensydige kroongroei en ’n reduksie in ruimte-gebruik toon, wat die groeidoeltreffendheid nadelig beinvloed. Simulasies met betrekking tot die verandering in boomaanplantgeometrie vanaf die huidige 2.7m x 2.7m na 2.3m x 3.1m en 2.4m x 3m het gedui op ’n reduksie van 20% in die masjienpadafstand en na minder rye wat uitgehaal moes word om die toegang van masjiene moontlik te maak. Die simulasie van die ontginning van dunnings het getoon dat verskillende aanplantgeometriealternatiewe die ontginningsproduktiwiteit met 10% tot 20% verbeter het, en die algehele dunningsoeskoste met tot 11% verminder het. In hierdie studie is die faktore, wat die gehalte van saaghoutkwaliteit en volume tydens die finale oes van die plantasie potensieel negatief mag beinvloed, suksesvol ondersoek. Dit illustreer ook die geskiktheid van simulasietoepassings vir die toets van ontginningsalternatiewe en die ontwikkelling van meer ekonomies voordelige praktyke .

In this thesis I present and test a methodology for developing a stand scale index of structural complexity. If properly designed such an index can act as a summary variable for a larger set of stand structural attributes, providing a means of ranking stands in terms of their structural complexity, and by association, their biodiversity and vegetation condition. This type of index can also facilitate the use of alternative policy instruments for biodiversity conservation, such as mitigation banking, auctions and offsets, that rely on a common currency – the index value – that can be compared or traded between sites. My intention was to establish a clear and documentable methodology for developing a stand scale index of structural complexity, and to test this methodology using data from real stands.¶ As a starting point, I reviewed the literature concerning forest and woodland structure and found there was no clear definition of stand structural complexity, or definitive suite of structural attributes for characterising it. To address this issue, I defined stand structural complexity as a combined measure of the number of different structural attributes present in a stand, and the relative abundance of each of these attributes. This was analogous to approaches that have quantified diversity in terms of the abundance and richness of elements. It was also concluded from the review, that stand structural complexity should be viewed as a relative, rather than absolute concept, because the potential levels of different structural attributes are bound within certain limits determined by the inherent characteristics of the site in question, and the biota of the particular community will have evolved to reflect this range of variation. This implied that vegetation communities with naturally simple structures should have the potential to achieve high scores on an index of structural complexity.¶ I proposed the following five-stage methodology for developing an index of stand structural complexity: 1. Establish a comprehensive suite of stand structural attributes as a starting point for developing the index, by reviewing studies in which there is an established relationship between elements of biodiversity and structural attributes. 2. Develop a measurement system for quantifying the different attributes included in the comprehensive suite. 3. Use this measurement system to collect data from a representative set of stands across the range of vegetation condition (highly modified to unmodified) and developmental stages (regrowth to oldgrowth) occurring in the vegetation communities in which the index is intended to operate. 4. Identify a core set of structural attributes from an analysis of these data. 5. Combine the core attributes in a simple additive index, in which attributes are scored relative to their observed levels in each vegetation community.¶ Stage one of this methodology was addressed by reviewing a representative sample of the literature concerning fauna habitat relationships in temperate Australian forests and woodlands. This review identified fifty-five studies in south-east and south-west Australia, in which the presence or abundance of different fauna were significantly (p&lt0.05) associated with vegetation structural attributes. The majority of these studies concerned bird, arboreal mammal, and ground mammal habitat requirements, with relatively fewer studies addressing the habitat requirements of reptiles, invertebrates, bats or amphibians. Thirty four key structural attributes were identified from these fifty-five studies, by grouping similar attributes, and then representing each group with a single generic attribute. This set, in combination with structural attributes identified in the earlier review, provided the basis for developing an operational set of stand level attributes for the collection of data from study sites.¶ To address stages two and three of the methodology, data were collected from one woodland community –Yellow Box-Red Gum (E. melliodora-E. Blakelyi ) – and two dry sclerophyll forest communities – Broadleaved Peppermint-Brittle Gum (E. dives-E. mannifera ), Scribbly Gum-Red Stringybark (E. rossii E. macrorhyncha ) – in a 15,000 km2 study area in the South eastern Highlands Bioregion of Australia. A representative set of 48 sites was established within this study area, by identifying 24 strata, on the basis of the three vegetation communities, two catchments, two levels of rainfall and two levels of condition, and then locating two sites (replicates) within each stratum. At each site, three plots were systematically established, to provide an unbiased estimate of stand level means for 75 different structural attributes.¶ I applied a three-stage analysis to identify a core set of attributes from these data. The first stage – a preliminary analysis – indicated that the 48 study sites represented a broad range of condition, and that the two dry sclerophyll communities could be treated as a single community, which was structurally distinct from the woodland community. In the second stage of the analysis, thirteen core attributes were dentified using the criteria that a core attribute should:¶ 1. Be either, evenly or approximately normally distributed amongst study sites; 2. Distinguish between woodland and dry sclerophyll communities; 3. Function as a surrogate for other attributes; 4. Be efficient to measure in the field. The core attributes were: Vegetation cover &lt0.5m Vegetation cover 0.5-6.0m; Perennial species richness; Lifeform richness; Stand basal area of live trees; Quadratic mean diameter of live stems; ln(number of regenerating stems per ha+1); ln(number of hollow bearing trees per ha+1);ln(number of dead trees per ha+1);sqrt(number of live stems per ha &gt40cm dbh); sqrt(total log length per ha); sqrt(total largelog length per ha); Litter dry weight per ha. This analysis also demonstrated that the thirteen core attributes could be modelled as continuous variables, and that these variables were indicative of the scale at which the different attributes operated.¶ In the third and final stage of the analysis, Principal Components Analysis was used to test for redundancy amongst the core attributes. Although this analysis highlighted six groupings, within which attributes were correlated to some degree, these relationships were not considered sufficiently robust to justify reducing the number of core attributes.¶ The thirteen core attributes were combined in a simple additive index, in which, each attribute accounted for 10 points in a total index value of 130. Attributes were rescaled as a score from 0-10, using equations that modelled attribute score as a function of the raw attribute data. This maintained a high correlation (r > 0.97, p< 0.0001) between attribute scores and the original attribute data. Sensitivity analysis indicated that the index was not sensitive to attribute weightings, and on this basis attributes carried equal weight. In this form my index was straightforward to apply, and approximately normally distributed amongst study sites.¶ I demonstrated the practical application of the index in a user-friendly spreadsheet, designed to allow landowners and managers to assess the condition of their vegetation, and to identify management options. This spreadsheet calculated an index score from field data, and then used this score to rank the site relative to a set of reference sites. This added a regional context to the operation of the index, and is a potentially useful tool for identifying sites of high conservation value, or for identifying sites where management actions have maintained vegetation quality. The spreadsheet also incorporated the option of calculating an index score using a subset of attributes, and provided a measure of the uncertainty associated with this score.¶ I compared the proposed index with five prominent indices used to quantify vegetation condition or habitat value in temperate Australian ecosystems. These were: Newsome and Catling’s (1979) Habitat Complexity Score, Watson et al.’s (2001) Habitat Complexity Score, the Site Condition Score component of the Habitat Hectares Index of Parkes et al. (2003), the Vegetation Condition Score component of the Biodiversity Benefits Index of Oliver and Parkes (2003), and the Vegetation Condition Score component of the BioMetric Assessment Tool of Gibbons et al. (2004). I found that my index differentiated between study sites better than each of these indices. However, resource and time constraints precluded the use of a new and independent data set for this testing, so that the superior performance of my index must be interpreted cautiously.¶ As a group, the five indices I tested contained attributes describing compositional diversity, coarse woody debris, regeneration, large trees and hollow trees – these were attributes that I also identified as core ones. However, unlike these indices, I quantified weeds indirectly through their effect on indigenous plant diversity, I included the contribution of non-indigenous species to vegetation cover and did not apply a discount to this contribution, I limited the direct assessment of regeneration to long-lived overstorey species, I used stand basal area as a surrogate for canopy cover, I quantified litter in terms of biomass (dry weight) rather than cover, and I included the additional attributes of quadratic mean diameter and the number of dead trees.¶ I also concluded that Parkes et al. (2003), Oliver and Parkes (2003), and Gibbons et al. (2004), misapplied the concept of benchmarking, by characterising attributes in terms of a benchmark range or average level. This ignored processes that underpin variation at the stand level, such as the increased development of some attributes at particular successional stages, and the fact that attributes can respond differently to disturbance agents. It also produced indices that were not particularly sensitive to the differences in attribute levels occurring between stands. I suggested that a more appropriate application of benchmarking would be at the overarching level of stand structural complexity, using a metric such as the index developed in this thesis. These benchmarks could reflect observed levels of structural complexity in unmodified natural stands at different successional stages, or thresholds for structural complexity at which a wide range of biota are present, and would define useful goals for guiding on-ground management.

Old-growth stands are important for management, conservation, wildlife, recreation, and maintaining biological diversity in forested landscapes. However, we are lacking the information needed to adequately identify and characterize old-growth stands. This is especially true for high elevation, interior forests. The characterization of stand structure and regeneration pattern will help in the development of site-specific guidelines for identifying old growth stands and restoring some of the old-growth characteristics in managed stands. This pamphlet presents a synopsis of a study investigating stand structure and regeneration of old-growth stands in the Moist Cold Engelmann Spruce - Subalpine Fir (ESSFmc) Subzone near Smithers, B.C. The three stands selected for the study were located on zonal sites, each in different watersheds, and the stands were established after fire. The criteria used for selection were: i) absence of lodgepole pine, ii) presence of advanced regeneration, and iii) abundant snags and coarse woody debris. These stands were presumed to represent the old-growth stage of stand development or the final (climax) stage of secondary succession.

The structure of a forest stand is characterized by: (a) species composition, (b) age, (c) size (diameter and height), and (d) spatial (horizontal and vertical) arrangement of the trees. Depending on the species, site, and disturbance history, the stand structure varies with time, thus providing a snapshot of a particular development stage. Research on growth and stand structure has shown that the spatial distribution of trees is one of the key determinants of stand productivity. Forest inventories and ecological surveys carried out in British Columbia (BC) have shown that the structure of naturally established, unmanaged stands varies from simple (single-species, single-storied, and even-aged) to complex (multi-species, multi-storied, and uneven-aged). Only a few studies have quantitatively characterized this range of structural complexity, with nearly all studies focusing on old-growth stands. BC forest policy requires that harvested areas be regenerated with a mixture of tree species whenever a mixture is suited to the site. This policy is based upon the assumption that under appropriate conditions, increases in stand productivity, reliability, and/or biodiversity can be attained in mixed-species stands. This assumption has not yet been tested for forest ecosystems. One mechanism by which different tree species can reduce crown competition for light is through vertical separation (the development of multiple canopy strata). Canopy stratification is not easily recognized in mixed-species stands, particularly when species have similar shade tolerance and height growth patterns, and no quantitative methods have been developed to detect stratification. The diameter frequency distribution of two-storied stands have been characterized by inverted J-shaped as well as modal curves. Although it would be more appropriate to characterize stand structure by height frequency distributions, these distributions have not been developed. We suggest that (i) a stand is stratified if there are distinct, quantitatifiable modes in the size distribution; either diameter, height, or crown height, and (ii) height or crown height distributions will be the most sensitive measures. To characterize the structure of western hemlock (Tsuga heterophylla (Raf.) Sarg.) (Hw) and western redcedar (Thuja plicata Donn ex D. Don in Lamb.) (Cw) second-growth stands, and to investigate its influence on tree growth, we (1) described and compared size (diameter, height, and crown height) frequency distributions in single- and mixed-species stands, (2) determined whether mixed-species stands develop a stratified canopy, and (3) examined whether interactions between hemlock and redcedar affect tree growth.

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