The Southern Caribbean Sea experiences a strong upwelling process along the coast from about 61°W to 75.5°W and 10-13°N. In this dissertation three aspects of this upwelling system are examined: (A) A mid-year secondary upwelling that was previously observed in the southeastern Caribbean Sea between June-July, when land based stations show a decrease in wind speed. The presence and effects of this upwelling along the whole southern Caribbean upwelling system were evaluated, as well as the relative forcing contribution of alongshore winds (Ekman Transport, ET) and wind-curl (Ekman Pumping, EP). (B) Stronger upwelling occurs in two particular regions, namely the eastern (63-65°W) and western (70-73°W) upwelling areas. However, the eastern area has higher fish biomass than the western area (78% and 18%, respectively, of the total small pelagic biomass of the southern Caribbean upwelling system). The upwelling dynamics along the southern Caribbean margin was studied to understand those regional variations on fish biomass. (C) The most important fishery in the eastern upwelling area off Venezuela is the Spanish sardine (Sardinella aurita). The sardine artisanal fishery is protected and only takes place up to ~10 km offshore. The effects of the upwelling cycle on the spatial distribution of S. aurita were studied. The main sources of data were satellite observations of sea surface temperature (SST), chlorophyll-a (Chl) and wind (ET and EP), in situ observations from the CARIACO Ocean Time-Series program, sardine biomass from 8 hydroacoustics surveys (1995-1998), and temperature profiles from the World Ocean Atlas 2005 used to calculate the depth of the Subtropical Underwater core (traced by the 22°C isotherm). The most important results of the study were as follows: (A) The entire upwelling system has a mid-year upwelling event between June-August, besides the primary upwelling process of December-April. This secondary event is short-lived (~5 weeks) and ~1.5°C warmer than the primary upwelling. Together, both upwelling events lead to about 8 months of cooler waters (-3, averaged from the coast to 100 km offshore) in the region. Satellite nearshore wind (~25 km offshore) remained high in the eastern upwelling area (> 6 m s-1) and had a maximum in the western area (~10 m s-1) producing high offshore ET during the mid-year upwelling (vertical transport of 2.4 - 3.8 m3 s-1 per meter of coastline, for the eastern and western areas, respectively). Total coastal upwelling transport was mainly caused by ET (~90%). However, at a regional scale, there was intensification of the wind curl during June as well; as a result open-sea upwelling due to EP causes isopycnal shoaling of deeper waters enhancing the coastal upwelling. (B) The eastern and western upwelling areas had upwelling favorable winds all year round. Minimum / maximum offshore ET (from weekly climatologies) were 1.52 / 4.36 m3 s-1 per meter, for the western upwelling area; and 1.23 / 2.63 m3 s-1 per meter, for the eastern area. The eastern and western upwelling areas showed important variations in their upwelling dynamics. Annual averages in the eastern area showed moderate wind speeds (6.12 m s-1), shallow 22°C isotherm (85 m), cool SSTs (25.24°C), and phytoplankton biomass of 1.65 mg m-3. The western area has on average stronger wind speeds (8.23 m s-1) but a deeper 22°C isotherm (115 m), leading to slightly warmer SSTs (25.53°C) and slightly lower phytoplankton biomass (1.15 mg m-3). We hypothesize that the factors that most inhibits fish production in the western upwelling area are the high level of wind-induced turbulence and the strong offshore ET. (C) Hydroacoustics values of Sardinella aurita biomass (sAsardine) and the number of small pelagics schools collected in the eastern upwelling region off northeast Venezuela were compared with environmental variables (satellite products of SST, SST gradients, and Chl -for the last two cruises-) and spatial variables (distance to upwelling foci and longitude-latitude). These data were examined using Generalized Additive Models. During the strongest upwelling season (February-March) sAsardine was widely distributed in the cooler, Chl rich upwelling plumes over the wide (~70km) continental shelf. During the weakest upwelling season (September-October) sAsardine was collocated with the higher Chl (1-3 mg m-3) found within the first 10 km from the upwelling foci; this increases Spanish sardine availability (and possibly the catchability) for the artisanal fishery. These results imply that during prolonged periods of weak upwelling the environmentally stressed (due to food scarceness) Spanish sardine population would be closer to the coast and more available to the fishery, which could easily turn into overfishing. After two consecutive years of weak upwelling (2004-2005) Spanish sardine fishery crashed and as of 2011 has not recovered to previous yield; however during 2004 a historical capture peak occurred. We hypothesize that this Spanish sardine collapse was caused by a combination of sustained stressful environmental conditions and of overfishing, due to the increased catchability of the stock caused by aggregation of the fish in the cooler coastal upwelling cells during the anomalous warm upwelling season.

Most climate change predictions focus on the response of individual species to changing local conditions and ignore species interactions, largely due to the lack of a sound theoretical foundation for how interactions are expected to change with climate and how to incorporate them into climate change models. Much of the variability in species interaction strengths may be governed by fundamental constraints on physiological rates, possibly providing a framework for including species interactions into climate change models. Metabolic rates, ingestion rates and many other physiological rates are relatively predictable from body size and body temperature due to constraints imposed by the physical and chemical laws that govern fluid dynamics and the kinetics of biochemical reaction times. My dissertation assesses the usefulness of this framework by exploring the community-level consequences of physiological constraints. In Chapter 2, I incorporated temperature and body size scaling into the biological rate parameters of a series of realistically structured trophic network models. The relative magnitude of the temperature scaling parameters affecting consumer energetic costs (metabolic rates) and energetic gains (ingestion rates) determined how consumer energetic efficiency changed with temperature. I systematically changed consumer energetic efficiency and examined the sensitivity of network stability and species persistence to various temperatures. I found that a species' probability of extinction depended primarily on the effects of organismal physiology (body size and energetic efficiency with respect to temperature) and secondarily on the effects of local food web structure (trophic level and consumer generality). This suggests that physiology is highly influential on the structure and dynamics of ecological communities. If consumer energetic efficiency declined as temperature increased, that is, species did best at lower temperatures, then the simulated networks had greater stability at lower temperatures. The opposite scenario resulted in greater stability at higher temperatures. Thus, much of the community-level response depends on what species energetic efficiencies at the organismal-level really are, which formed the research question for Chapter 3: How does consumer energetic efficiency change with temperature? Existing evidence is scarce but suggestive of decreasing consumer energetic efficiency with increasing temperature. I tested this hypothesis on seven rocky intertidal invertebrate species by measuring the relative temperature scaling of their metabolic and ingestion rates as well as consumer interaction strength under lab conditions. Energetic efficiencies of these rocky intertidal invertebrates declined and species interaction strengths tended to increase with temperature. Thus, in the rocky intertidal, the mechanistic effect of temperature would be to lower community stability at higher temperatures. Chapter 4 tests if the mechanistic effects of temperature on ingestion rates and species interaction strengths seen in the lab are apparent under field conditions. Bruce Menge and I related bio-mimetic estimates of body temperatures to estimates of per capita mussel ingestion rates and species interaction strengths by the ochre sea star Pisaster ochraceus, a keystone predator of the rocky intertidal. We found a strong, positive effect of body temperature on both per capita ingestion rates and interaction strengths. However, the effects of season and the unique way in which P. ochraceus regulates body temperatures were also apparent, leaving room for adaptation and acclimation to partially compensate for the mechanistic constraint of body temperature. Community structure of the rocky intertidal is associated with environmental forcing due to upwelling, which delivers cold, nutrient rich water to the nearshore environment. As upwelling is driven by large-scale atmospheric pressure gradients, climate change has the potential to affect a wide range of significant ecological processes through changes in water temperature. In Chapter 5, my coauthors and I identified long-term trends in the phenology of upwelling events that are consistent with climate change predictions: upwelling events are becoming stronger and longer. As expected, longer upwelling events were related to lower average water temperatures in the rocky intertidal. Furthermore, recruitment rates of barnacles and mussels were associated with the phenology of upwelling events. Thus climate change is altering the mode and the tempo of environmental forcing in nearshore ecosystems, with ramifications for community structure and function. Ongoing, long-term changes in environmental forcing in rocky intertidal ecosystems provide an opportunity to understand how temperature shapes community structure and the ramifications of climate change. My dissertation research demonstrates that the effect of temperature on organismal performance is an important force structuring ecological communities and has potential as a tractable framework for predicting the community level effects of climate change. / Graduation date: 2013 / Access restricted to the OSU Community, at author's request, from Nov. 20, 2012 - Nov. 20, 2014

Le golfe de Gascogne (GdG), un milieu riche en processus physiques a été étudié à partir de simulations numériques. L’étude est construite autour d'échelles allant du GdG à la sous méso-échelle. Dans la première partie, nous avons examiné la variabilité interannuelle de la température et de la salinité de surface sur une période de 53 ans : nous avons pu décrire deux tendances en lien avec I'Atlantique Nord-Est. Le refroidissement et la dessalure jusqu'en 1976 seraient liés à la grande anomalie de salinité, le réchauffement et la salinification actuels liés à I'augmentation de CO2 atmosphérique. Le GdG se caractérise par un courant de pente, lberian Poleward Current (lPC) : sa variabilité serait liée au vent du Sud-Ouest qui renforce l'lPC par un courant géostrophique dans le Bassin lbérique. L’un des résultats intéressant trouvé dans les simulations et confìrmé par les observations est l'apparition des anomalies froides liées à des upwellings en alternance avec des anomalies chaudes "La Navidad". Ces upwellings seraient liés au vent de Nord dans le Bassin lbérique mais au courant d'Ouest le long des côtes Nord espagnoles. Dans une deuxième partie, nous nous sommes attachés à la méso et sous méso-échelle à travers la détection des tourbillons et la variabilité des spectres d'énergie. Un indice basé sur le rapport entre I'anomalie de densité de surface et I'anomalie de niveau de la mer permet de détecter les tourbillons de subsurface et de les distinguer des tourbillons de surface. Une application de cet indice à partir des données satellites confirme le potentiel de détection des Slope Water Oceanic eDDIES (tourbillons de subsurface caractéristiques du GdG). La description de l'énergie cinétique turbulente (EKE) dans le GdG montre une variabilité spatiale avec un maximum le long de la côte Nord espagnole liée à I'lPC. Les pentes des spectres (k-4.2 pour la SSH, en k-2.4 pour la SST et en k-2.4 pour l'énergie cinétique) sont différents des observations satellites, mais comparables avec les précédentes études. Ces pentes de spectres ont également une variabilité saisonnière avec un maximum en hiver et un minimum en été, liée au cycle saisonnier de I'EKE. / The Bay of Biscay (BoB), an environment rich in physical processes has been studied from numerical simulations. Thestudy is built around scales from the size of BoB until sub mesoscale.ln the first part, we examined the interannual variability of the sea surface temperature and saliniÇ over a period of 53years: we were able to describe two trends related to the North-East Atlantic. Cooling and freshening until 1976 thatcould be related to the Great Salinity Anomaly and current salinification related to the atmospheric increase of CO2.The Bay of Biscay is characterized by a slope current, the lberian Poleward Current (lPC): its variability is linked to theSouth West wind strengthens the IPC by a geostrophic current in the lberian Basin. One of the interesting results foundin simulations and confirmed by observations is the appearance of cold anomalies related to upwellings and alternatingwith warm anomalies 'La Navidad'. These upwellings could be linked to the north wind in the lberian Basin but to the West current along the northern Spanish coast.ln the second part, we are committed to the meso and sub mesoscale eddies through a method of detection and throughthe variability of energy spectra. An index based on the ratio of surface density anomaly and sea level anomaly allowsdetecting subsurface vortices and distinguishing them from the surface ones. The application of this index from thesatellite data confirms the detection potential of Slope Water Oceanic Eddies (subsurface vortices of BoB).The description of the Eddy Kinetic Energy (EKE) in the BoB. shows a spatial variability with maximum along the Spanishnorth coast linked to the lPC. The slopes of the spectra (k-4.2 for SSH, k-2.4 for SST and k-2.4 for the kinetic energy) are different from satellite observations, but comparable with previous studies. These spectral slopes have a seasonalvariability with a maximum in winter and minimum in summe¡ related to the seasonal cycle of EKE.

Le but de cette thèse est d'analyser plusieurs phénomènes de bord ouest de la Mer d'Arabie : (i) le cycle de vie d'un tourbillon de mésoéchelle persistant, le Great Whirl; (ii) la dynamique d'un écoulement d'eau dense (outflow) formée dans une mer adjacente, l'outflow du Golfe Persique; et (iii) une remontée d'eau profonde (upwelling) saisonnière dans la zone côtière d'Oman. Le point commun entre ces phénomènes est leur localisation sur un bord ouest océanique. Ils sont donc influencés par des forçages locaux (notamment les vents de mousson) et les forçages à distance (ondes de Rossby et tourbillons dérivant vers l'ouest). En particulier, ces derniers vont jouer un rôle particulier car la Mer d'Arabie est située à basses latitudes, ce qui implique une propagation rapide des ondes longues et tourbillons. De plus, des ondes sont continuellement excitées par le régime saisonnier des moussons. Nous avons mis au point des expériences numériques de différentes complexités en utilisant un modèle aux équations primitives. Ces expériences permettent soit de simuler de manière réaliste la dynamique complexe de la Mer d'Arabie, soit d'isoler un processus en particulier. Les résultats principaux peuvent se résumer comme suit : (i) le cycle de vie du Great Whirl est significativement impacté par les ondes de Rossby annuelles. Le rotationnel de la tension de vent joue un rôle important dans le maintien, le renforcement et la barotropisation du tourbillon. (ii) La dispersion de l'Eau du Golfe Persique (Persian Gulf Water, PGW) est déterminée par le mélange induit par les tourbillons de mésoéchelle. Précisément, ces tourbillons entrent dans le Golfe d'Oman (où se déverse la PGW), et interagissent avec la topographie. Ces interactions frictionnelles produisent des bandes de vorticité très intenses dans la couche limite de fond. Celles-ci sont arrachées et forment des tourbillons de sous-mésoéchelle. Ces tourbillons capturent de la PGW initialement située sur la pente continentale et la redistribuent dans le golfe d'Oman. Ce mécanisme donne finalement lieu à du mélange, permettant d'expliquer le gradient de salinité climatologique observé en profondeur. (iii) La dynamique de l'upwelling saisonnier au large d'Oman contraste fortement avec la dynamique des upwelling de bord est (Eastern Boundary Upwelling Systems, EBUS). En effet, les ondes de Rossby se propagent vers le large dans les EBUS et vers la côte dans l'upwelling de bord ouest d'Oman. Ces ondes modulent la réponse en température de l'upwelling forcé par le vent.Dans l'ensemble, ces résultats sont relativement spécifiques à la Mer d'Arabie. La faible extension zonale et la basse latitude de la Mer d'Arabie, ainsi que le régime de mousson des vents saisonniers en font une région particulière. La propagation rapide des ondes et tourbillons et leurs interactions avec le bord ouest façonnent les régimes de turbulence de la Mer d'Arabie. / This PhD aims to investigate some western boundary processes in the Arabian Sea : (i) the life cycle of the socalled Great Whirl, a persistent mesoscale eddy; (ii) the dynamics of the Persian Gulf outflow, a marginal sea dense outflow; and (iii) the seasonal Oman upwelling, a coastal upwelling forced by summermonsoonal winds. The cornerstone of all these phenomena is their locationat a western boundary, which makes then being influenced by both localforcing (e.g., monsoonal winds) and remote forcing (Rossby waves and wesward drifting eddies). Specifically, the later are expected to impact the western boundary dynamics since the low latitude of the Arabian Sea implies a fast westward propagation of long Rossby waves and eddies. Moreover, waves are continously excited by the reversing monsoonal winds. Based on a primitive equation model, we designed numerical experiments of different complexity that allowed to either realistically simulate the dynamics in the Arabian Sea or to isolate some processes.Major findings can be summarized as follows : (i) The Great Whirl life cycle is found to be significantly paced by annual Rossby waves, although the strong monsoonal wind stress curl is of major importance to sustain the structure. (ii) The Persian Gulf Water (PGW) spreading in the Gulf of Oman and the northern Arabian Sea can be explained by the stirring done by eddies entering the Gulf. These remotely formed surface intensifed mesoscale eddies propagate into the Gulf and interact with the topography. Frictional interactions produce intense vorticity strips at the boundary that detach and roll up in the interior, forming submesoscale coherent vortices (SCV). These SCV trap PGW initially located on the slope and redistribute it in the interior. This mechanism of transport ultimately produces mixing that explains the large-scale gradient of salinity in the gulf. (iii) We find that the dynamics of the seasonal upwelling of Oman contrasts with the more deeply studied Eastern Boundary Upwelling Systems (EBUS). In particular, Rossby waves, propagating offshore in EBUS vs. onshore in this western boudary upwelling, are found to modulate the wind driven upwelling and its sea surface temperature response.Overall, these results appear to be rather specific to the Arabian Sea. The short zonal extent and the low-latitude of the Arabian Sea, as well as the seasonally reversing wind forcing are the distinguishing features of this region. Fast waves and drifting eddies and their interactions with the western boundary significantly shape the turbulent regimes of the western Arabian Sea.

Ao largo da costa oeste da Am´erica do Sul o sistema de ressurg^encia costeira apresenta uma excepcional produtividade, um resultado da alta efici^encia do ecossistema biol´ogico em converter a produ¸c~ao prim´aria em biomassa. Esses processos qu´?micobiol ´ogicos s~ao fortemente modulados pelo ambiente f´?sico da regi~ao. Por exemplo, durante eventos do El-Ni~no a produtividade pesqueira ´e fortemente prejudicada em resposta a uma inibi¸c~ao da ressurg^encia costeira. Apesar disso ser um fato j´a bastante conhecido desde ´epocas pr´e-colombianas, muita d´uvida ainda persiste sobre o mecanismo pelo qual o sistema costeiro ´e afetado por esses fen^omenos de grande escala. O presente trabalho foi realizado com o objetivo de se contribuir com esse conhecimento, estudando como ocorreram as mudan¸cas do sistema durante recentes eventos El Ni~no, principalmente na escala intrasazonal. Nessa investiga¸c~ao foram utilizadas diferentes metodologias de an´alise de dados para identificar a natureza dos for¸cantes de oscila¸c~oes significativas observadas durante o El Ni~no 1997-1998. Tamb´em foi investigado se essas oscila¸c~oes foram for¸cadas remotamente e se propagaram como ondas aprisionadas na costa; quais foram suas propriedades espaciais e sua propaga¸c~ao longo da costa. A primeira parte do trabalho consiste na an´alise da variabilidade temporal e a caracteriza¸c~ao das flutua¸c~oes de baixa freq¨u^encia ocorridas durante eventos El Ni~no. Essa an´alise faz uso do espectro de ondeleta aplicado a s´eries temporais longas de dados do n´?vel do mar de esta¸c~oes costeiras distribu´?das ao longo da costa sul do Equador, costa do Peru e costa norte do Chile. O espectro de ondeleta cruzada e o espectro da coer^encia foram usados para avaliar a import^ancia relativa das for¸cantes remota e local das varia¸c~oes do n´?vel do mar associadas com os efeitos do El Ni~no 1997-1998. Na segunda parte, as flutua¸c~oes for¸cadas remotamente foram comparadas com modelos lineares simples e o modelo de Brink e Chapman (1987) para se obter as propriedades das ondas aprisionadas na costa durante o El Ni~no Extraordin ´ario (1997-1998). Uma importante conclus~ao dessas an´alises foi a constata¸c~ao de que variabilidades intrasazonais ao largo da costa do Peru durante esse El Ni~no foi, em parte significativa, o resultado da atua¸c~ao de for¸cantes remotas e locais, associadas com ondas equatoriais incidindo na Costa Oeste da Am´erica do Sul e com o efeito do vento, respectivamente. / Off the western coast of South America, the coastal upwelling system presents an exceptional productivity, a result of the high efficiency of the biological ecosystem in converting the primary production into biomass. These chemical and biological processes are strongly modulated by the physical environment. For instance, during El Ni~no events the fishery productivity is strongly impaired in response to an inhibition of the coastal upwelling. In spite of this fact be widely known since pre-Colombian times, much doubts still persist about the mechanism by means of which the coastal system is affected by the large scale phenomena. The present work was idealized with the intention of contributing for this knowledge, studying how occurred the changes in the system during recent El Ni~no events, with emphasis on the intraseasonal scale. In this investigation, different methodologies of data analysis were used to identify the nature of the forcing which produced significant oscillations observed during the El Ni~no 1997-1998. Also, it was investigated if these oscillations were remotely forced and propagated along the coast as coastal trapped waves; what were their spatial and propagation properties. The first part of the work consists of an analysis of the temporal variability and the characterization of the low-frequency fluctuations occurred during the El Ni~no events. This analysis uses the wavelet spectrum applied to time series of sea level data collected off southern Equador, Peru and northern Chile. The cross wavelet spectrum and coherence spectrum were used to evaluate the relative importance of local and remote forcing of the sea level, associated with effects of the 1997-1998 El Ni~no. In the second part, the fluctuations remotely forced wer compared with results of simple linear models and with the model of Brink and Chapman (1987), in order to obtain the coastal trapped waves properties during that extraordinary El Ni~no event. One important conclusion of these analyses was the confirmation that the intraseasonal variability observed offo Peru during the 1997-1998 El Ni~no was, in part, the result of remote and local forcing, associated with equatorial waves and the wind effect, respectively.

O presente trabalho teve como finalidade estudar os efeitos das variações do vento resultantes de ocorrências do fenômeno El Niño-Southern Oscillation (ENSO) nos padrões da circulação superficial ao longo da costa peruana, através da análise de dados observados e de modelagem numérica. É enfocado o período 1991-2000, quando ocorreram fortes eventos La Niña (1996-97, 1998-2000) e El Niño (1997- 98). Esses eventos tiveram fortes impactos em escala global mas muito pouco se sabe sobre os impactos locais na estrutura da termoclina e no ciclo da dinâmica de Ekman ao longo da costa peruana. Os dados analisados no presente estudo foram dados coletados em duas radiais ao longo das latitudes 5 S e 15 S. O modelo oceânico utilizado foi o Modelo da Universidade de Princeton (Princeton Ocean Model-POM). As simulações numéricas foram forçadas com produtos de vento relativos ao período 1991-2000. Essas simulações reproduziram satisfatoriamente os padrões médios da circulação na região de estudo, confirmando que nas áreas próximas da costa o vento é o principal mecanismo gerador de ressurgência ou subsidência. Os resultados mostraram que durante a ocorrência do forte evento El Niño 1997-1998, nas radiais de 5 S e 15 S, houve uma drástica alteração do sistema de ressurgência costeira em resposta às variações do vento. / The objective of the present work was to study the effects of variability in the wind due to the El Niño-Southern Oscillation events in the surface circulation patterns of the Peruvian coast, by means of data analysis and numerical modeling. It is focused the period 1991-2000, when it was observed strong La Niña (1996-1997, 1998-2000) and El Niño events (1997-98). These events had strong effects worlwide but very little is known on the local impacts on the circulation, thermocline structure and the Ekman dynamics of the Peruvian coast. The data analysed in the present study were collected on transects along 5 S and 15 S. The model used was an implementation of the Princeton Ocean Model (POM). The numerical simulations were forced with wind products relative to the period of interest (1991-2000). The simulations reproduced satisfactorily the mean circulation patterns in the study area, confirming that the in the nearshore region the wind is the main driving mechanism for coastal upwelling/downwelling. The results show that during the strong 1997-1998 El Niño, a drastic alteration of the coastal upwelling system ocurred in response to the changes in the wind.

Um modelo numérico tridimensional foi adaptado para o estudo da circulação gerada pelo vento sobre a plataforma continental da Região Sul do Brasil durante o verão e o outono. O modelo oceânico de Princeton (POM) é usado em um domínio limitado com grade curvilínea de alta-resolução, além de linha de costa e batimetria realísticas. As forçantes são variáveis no tempo e incluem tensão de cisalhamento do vento, descarga fluvial, campos termohalinos e nível médio do mar. Os objetivos são simular a resposta da plataforma a estas forçantes, comparar os resultados do modelo com observações oceanográficas afim de estabelecer a confiança do modelo, e subseqüentemente analisar os campos resultantes para ajudar na compreensão do comportamento da circulação observada. As variáveis do modelo mostram boa concordância geral com as correspondentes observações. É encontrado um claro padrão sazonal da circulação gerada pelo vento sobre a plataforma, com fluxos de superfície para sul e para o largo durante o verão, resultando em ressurgência de águas frias de fundo, e para norte e para a costa durante o outono, resultando em subsidência de águas menos densas. Tal circulação aumenta a estabilidade vertical no verão, devido ao aquecimento de superfície e ressurgência de águas frias, e aumenta a mistura vertical e a homogeneidade vertical no outono, devido à subsidência de águas menos densas, a qual causa forte aceleração da pluma fluvial existente na plataforma interna. Os resultados também demonstram que os cabos e promontórios da Região Sul do Brasil desempenham papel fundamental na variabilidade ao longo da costa do fluxo de ressurgência, incluindo o estabelecimento de jatos costeiros baroclínicos, os quais tendem a se separar da costa ao sul de cabos, mostrando uma circulação tridimensional de ressurgência, com complexa dependência espaço-temporal. Durante o outono, o empilhamento de subsidência, somado à presença da pluma fluvial, estabelece uma corrente costeira barotrópica, chamada aqui de \'\'Corrente Costeira de Rio Grande\'\', causando a intrusão de águas frias a partir do sul para a plataforma continental brasileira. A avaliação dos campos de velocidade mostra que o transporte de Ekman pleno se desenvolve na região ao largo das frentes de ressurgência ou subsidência, deixando em direção à costa uma zona dominada por fricção, com estratificação e velocidades perpendiculares fracas. Esta região é denominada aqui de \'\'plataforma interna\'\' e se estende da costa até a isóbata de 30m no verão e até a isóbata de 45m no outono, com esta variação resultando de variações na profundidade da camada de Ekman. / A three-dimensional numerical model has been adapted for the study of the wind-driven flow on the continental shelf off southern Brazil during the summer and autumn seasons. The Princeton Ocean Model (POM) is used in a limited area domain with a high-resolution (eddy resolving) curvilinear grid, with realistic coastline and bottom topography. The forcings include time-varying wind stress, termohaline fields, fluvial discharges and mean sea level. The objectives of the study are to simulate the response of the shelf circulation to these forcings, to compare model results with oceanographic observations, to stablish the model confidence, and to subsequently analyse the model fields a contribution to understand the behavior of the observed flow. The model variables show overall good agreement with corresponding observations. It is found a clear seasonal pattern of wind-driven circulation over the shelf, with surface offshore and southward net flow during summertime, which results in bottom cold water upwelling, and onshore and northward net flow during autumntime, which results in less dense water downwelling. This circulation enhances vertical stability in summer, due to solar surface heating and cold water bottom upwelling, and enhances vertical mixing and unstratified conditions in autumn due to downwelling of less dense water, which causes strong downwind acceleration of river plume innershelf waters. The results also demonstrate that coastal capes and promontories play a dominant role in causing alongshore variability of the upwelling flow over southern Brazil, including the setup of alongshore coastal baroclinic jets, which tend to separate from the coast south of capes, showing a three-dimensional upwelling circulation, with complex time and space dependence. During autumntime, the downwelling wind setup, in addition to the river plume waters, set up a barotropic coastal current, named here \'\'Rio Grande Coastal Current\'\', causing the intrusion of southern colder water over the Brazilian shelf. The evaluation of velocity fields shows that full Ekman transport develops in the region offshore of the upwelling or downwelling fronts, leaving onshore a region dominated by friction with relatively weak stratification and cross-shore currents. This region is named here \'\'innershelf\'\' and extends seaward from the coast until the 30m isobath in summer and the 45m isobath in autumn, showing this seasonal variation due to Ekman layer depth variations.

O presente trabalho teve como finalidade estudar os efeitos das variações do vento resultantes de ocorrências do fenômeno El Niño-Southern Oscillation (ENSO) nos padrões da circulação superficial ao longo da costa peruana, através da análise de dados observados e de modelagem numérica. É enfocado o período 1991-2000, quando ocorreram fortes eventos La Niña (1996-97, 1998-2000) e El Niño (1997- 98). Esses eventos tiveram fortes impactos em escala global mas muito pouco se sabe sobre os impactos locais na estrutura da termoclina e no ciclo da dinâmica de Ekman ao longo da costa peruana. Os dados analisados no presente estudo foram dados coletados em duas radiais ao longo das latitudes 5 S e 15 S. O modelo oceânico utilizado foi o Modelo da Universidade de Princeton (Princeton Ocean Model-POM). As simulações numéricas foram forçadas com produtos de vento relativos ao período 1991-2000. Essas simulações reproduziram satisfatoriamente os padrões médios da circulação na região de estudo, confirmando que nas áreas próximas da costa o vento é o principal mecanismo gerador de ressurgência ou subsidência. Os resultados mostraram que durante a ocorrência do forte evento El Niño 1997-1998, nas radiais de 5 S e 15 S, houve uma drástica alteração do sistema de ressurgência costeira em resposta às variações do vento. / The objective of the present work was to study the effects of variability in the wind due to the El Niño-Southern Oscillation events in the surface circulation patterns of the Peruvian coast, by means of data analysis and numerical modeling. It is focused the period 1991-2000, when it was observed strong La Niña (1996-1997, 1998-2000) and El Niño events (1997-98). These events had strong effects worlwide but very little is known on the local impacts on the circulation, thermocline structure and the Ekman dynamics of the Peruvian coast. The data analysed in the present study were collected on transects along 5 S and 15 S. The model used was an implementation of the Princeton Ocean Model (POM). The numerical simulations were forced with wind products relative to the period of interest (1991-2000). The simulations reproduced satisfactorily the mean circulation patterns in the study area, confirming that the in the nearshore region the wind is the main driving mechanism for coastal upwelling/downwelling. The results show that during the strong 1997-1998 El Niño, a drastic alteration of the coastal upwelling system ocurred in response to the changes in the wind.

Cette thèse s'intéresse à la dynamique océanique induite par les échelles spatiales et temporelles de l'atmosphère, et du vent en particulier, dans les régions d'upwelling du Benguela et des Canaries. Ces régions sont sous l'influence d'un vent local ou régional, soufflant parallèlement à la côte. "Moteur" principal de la résurgence d'eau froide, ce vent est modulé par des processus physiques à des échelles spatio-temporelles variées. La nature des interactions avec l'atmosphère, l'océan et le continent environnants diffère selon les processus. Depuis deux décennies, des efforts remarquables portent sur la description par télédétection des champs atmosphériques à la surface de l’océan. Un nombre croissant de missions spatiales et des améliorations techniques majeures ont permis de raffiner la résolution horizontale et temporelle des produits disponibles à l'échelle globale. La disponibilité de multiples mesures diffusiométriques grillées, traitées et distribuées par le LOSCERSAT,nous amène dans un premier temps à comparer et analyser la richesse et la finesse des échelles retranscrites par différents produits. Ainsi, plusieurs gammes d'échelles de vent sont différenciées et leurs signatures sur l'upwelling côtier sont étudiées. L'intensité des anticyclones subtropicaux (Sainte Hélène et Açores) module la saisonnalité de l'upwelling le long des côtes Ouest africaines. Les régions centrales des upwellings de l’Atlantique, sous l'influence permanente de ces centres de haute pression, sont ainsi les cellules d'upwelling les plus intenses de chacun des systèmes en termes de pérennité et d'intensité (cellule de Lüderitz et cellule de Dakhla respectivement dans l'hémisphère Sud et l'hémisphère Nord). À l'échelle régionale, ou l'échelle des sous-bassins (O(1000 km)), la variabilité intrasaisonnière du vent est contrôlée par le renforcement ou l'atténuation des anticyclones entraînant à la côte l'activation ou la relaxation d'événements d'upwelling. À des échelles plus petites (O(100 km)), le front caractéristique de température de surface (SST) entre la côte et le large façonne la structure spatiale du vent par des processus de stabilisation/déstabilisation de la colonne d'air. Un vent soufflant en direction de l'équateur et parallèlement à un front de SST aura tendance à diminuer (augmenter) sur le flanc froid (chaud) de ce front. Le rotationnel (la divergence) du vent est directement impacté(e) et répond linéairement, au premier ordre, à la composante du gradient de SST normale (tangentielle) à la direction du vent. Ces rétroactions océaniques sont caractérisées par une échelle temporelle allant de l'hebdomadaire au mensuel. Enfin, de fines échelles du vent sont couramment observées dans les premiers kilomètres de l’océan au voisinage de la côte. L’interface entre le large et le continent est en effet associée à un affaiblissement significatif des vents. L'extension zonale de cette transition (O(10 km)) dépend notamment de l'orographie et de la rugosité de surface du continent adjacent. L'impact d'une telle réduction du vent sur la structure des upwellings côtiers, la dynamique sous-jacente et le transport côte-large de particules est appréhendé à l'aide d'analyses numériques eulériennes et lagrangiennes. / This study focuses on the oceanic response to fine atmospheric spatial and temporal scales, and especially fine wind patterns in the Benguela and Canary upwelling systems. These regions are under the influence of local or regional wind, blowing parallel to the coast. Thewind is the main driver of the cold-water upwelling and is modulated by several physical processes at various scales. The nature of the interactions with the atmosphere, the ocean and the adjacent continent differs according to these processes. For the past 20 years, outstanding efforts have been made in the description and understanding of the atmospheric conditions at the sea surface. An increasing number of space missions and major technical improvements have allowed refinement of the horizontaland temporal resolution of the products available at global scale. The availability of multiple gridded scatterometer measurements,processed and distributed by the LOS-CERSAT, brings us first to compare and analyze the richness and fineness of the scales of a few products. We differentiate several wind scales and study their signatures on coastal upwelling dynamics. The intensity of the subtropical anticyclones (Saint Helena andAzores) modulates the seasonality of the upwelling along the Africanwest coast. The central regions of both upwelling systems are permanently under the influence of these atmospheric highs and,thus, are the most intense upwelling cells of each system, both interms of durability and intensity (Lüderitz and Dakhla cells for the southern and the northern hemisphere, respectively). On a regional scale, or basin scale (O(1000 km)), the intraseasonal wind variability is driven by the strengthening or weakening of these anticyclones, causing the activation or relaxation of upwelling events at the coast.At smaller scales (O(100 km)), the characteristic sea surface temperature (SST) front between the coastal and open ocean shapes the spatial structure of the wind by stabilization/destabilization of the air column. An equatorward-blowing wind parallel to an SST front tends to decrease (increase) on the cold side (warm) of this front. The curl (divergence) of the wind is directly impacted and the first order response varies linearly with the crosswind (downwind) SST gradient. This oceanic feedback is characterized by weekly to monthly temporal scales. Finally, small-scale wind structures are frequently observed in the first kilometers of the coastal ocean. Indeed, the interface between the open ocean and the continent is associated with a significant wind drop-off. The zonal extension of this transition (O(10 km) depends on the orography and on the surface roughness of the adjacent continent. The impact of such a wind reduction on the structure of the coastal upwelling, the underlying ocean dynamics and the cross-shore transport of particles is diagnosed with both Eulerian and Lagrangian numerical analyses.

Um modelo numérico tridimensional foi adaptado para o estudo da circulação gerada pelo vento sobre a plataforma continental da Região Sul do Brasil durante o verão e o outono. O modelo oceânico de Princeton (POM) é usado em um domínio limitado com grade curvilínea de alta-resolução, além de linha de costa e batimetria realísticas. As forçantes são variáveis no tempo e incluem tensão de cisalhamento do vento, descarga fluvial, campos termohalinos e nível médio do mar. Os objetivos são simular a resposta da plataforma a estas forçantes, comparar os resultados do modelo com observações oceanográficas afim de estabelecer a confiança do modelo, e subseqüentemente analisar os campos resultantes para ajudar na compreensão do comportamento da circulação observada. As variáveis do modelo mostram boa concordância geral com as correspondentes observações. É encontrado um claro padrão sazonal da circulação gerada pelo vento sobre a plataforma, com fluxos de superfície para sul e para o largo durante o verão, resultando em ressurgência de águas frias de fundo, e para norte e para a costa durante o outono, resultando em subsidência de águas menos densas. Tal circulação aumenta a estabilidade vertical no verão, devido ao aquecimento de superfície e ressurgência de águas frias, e aumenta a mistura vertical e a homogeneidade vertical no outono, devido à subsidência de águas menos densas, a qual causa forte aceleração da pluma fluvial existente na plataforma interna. Os resultados também demonstram que os cabos e promontórios da Região Sul do Brasil desempenham papel fundamental na variabilidade ao longo da costa do fluxo de ressurgência, incluindo o estabelecimento de jatos costeiros baroclínicos, os quais tendem a se separar da costa ao sul de cabos, mostrando uma circulação tridimensional de ressurgência, com complexa dependência espaço-temporal. Durante o outono, o empilhamento de subsidência, somado à presença da pluma fluvial, estabelece uma corrente costeira barotrópica, chamada aqui de \'\'Corrente Costeira de Rio Grande\'\', causando a intrusão de águas frias a partir do sul para a plataforma continental brasileira. A avaliação dos campos de velocidade mostra que o transporte de Ekman pleno se desenvolve na região ao largo das frentes de ressurgência ou subsidência, deixando em direção à costa uma zona dominada por fricção, com estratificação e velocidades perpendiculares fracas. Esta região é denominada aqui de \'\'plataforma interna\'\' e se estende da costa até a isóbata de 30m no verão e até a isóbata de 45m no outono, com esta variação resultando de variações na profundidade da camada de Ekman. / A three-dimensional numerical model has been adapted for the study of the wind-driven flow on the continental shelf off southern Brazil during the summer and autumn seasons. The Princeton Ocean Model (POM) is used in a limited area domain with a high-resolution (eddy resolving) curvilinear grid, with realistic coastline and bottom topography. The forcings include time-varying wind stress, termohaline fields, fluvial discharges and mean sea level. The objectives of the study are to simulate the response of the shelf circulation to these forcings, to compare model results with oceanographic observations, to stablish the model confidence, and to subsequently analyse the model fields a contribution to understand the behavior of the observed flow. The model variables show overall good agreement with corresponding observations. It is found a clear seasonal pattern of wind-driven circulation over the shelf, with surface offshore and southward net flow during summertime, which results in bottom cold water upwelling, and onshore and northward net flow during autumntime, which results in less dense water downwelling. This circulation enhances vertical stability in summer, due to solar surface heating and cold water bottom upwelling, and enhances vertical mixing and unstratified conditions in autumn due to downwelling of less dense water, which causes strong downwind acceleration of river plume innershelf waters. The results also demonstrate that coastal capes and promontories play a dominant role in causing alongshore variability of the upwelling flow over southern Brazil, including the setup of alongshore coastal baroclinic jets, which tend to separate from the coast south of capes, showing a three-dimensional upwelling circulation, with complex time and space dependence. During autumntime, the downwelling wind setup, in addition to the river plume waters, set up a barotropic coastal current, named here \'\'Rio Grande Coastal Current\'\', causing the intrusion of southern colder water over the Brazilian shelf. The evaluation of velocity fields shows that full Ekman transport develops in the region offshore of the upwelling or downwelling fronts, leaving onshore a region dominated by friction with relatively weak stratification and cross-shore currents. This region is named here \'\'innershelf\'\' and extends seaward from the coast until the 30m isobath in summer and the 45m isobath in autumn, showing this seasonal variation due to Ekman layer depth variations.