Understanding how populations are structured and how they use natural and anthropogenic spaces is essential for effective wildlife management. A total of 510 barren-ground (Rangifer tarandus groenlandicus), 176 boreal (R. t. caribou), 11 mountain woodland (R. t. caribou), and 39 island (R. t. groenlandicus x pearyi) caribou were tracked with satellite collars in 1993-2009 in the Northwest Territories, Nunavut, and northern Alberta. Using satellite location data and hierarchical and fuzzy cluster analyses, I verified that Cape Bathurst, Bluenose-West, Bluenose-East, Bathurst, Beverly, Qamanirjuaq, and Lorillard barren-ground subpopulations were robust; the Queen Maude Gulf and Wager Bay barren-ground subpopulations were distinct. Dolphin and Union island caribou formed one population; boreal caribou formed two distinct subpopulations. Females in robust subpopulations were structured by strong annual spatial affiliation; those in distinct subpopulations were spatially independent and structured by migratory connectivity, movement barriers, and/or habitat discontinuity. An east-west cline in annual-range sizes and path lengths supported the subpopulation structure identified for migratory barren-ground caribou. I analyzed satellite location data to determine parturition dates and activity periods for all caribou ecotypes. For parturition dates I found a north-south cline for boreal caribou, west-east cline for migratory barren-ground caribou, and ecotype and subspecies clines for boreal and barren-ground caribou. Based on annual changes in movement rates I identified eight activity periods for boreal and tundra-wintering, 10 for mountain woodland, and 12 for migratory barren-ground caribou. Based distribution and movements, boreal caribou avoided seismic lines during periods when females and calves were most vulnerable to predators or hunters. They crossed fewer seismic lines and travelled faster when they crossed them than expected. Caribou avoided areas ≤400 m from seismic lines where they could space away from them suggesting that they perceive these as risky areas. I defined secure habitats as areas that were >400 m from anthropogenic linear features. Population growth rates were higher in areas where they had access to secure unburned habitat and where most of that was in patches >500 km2. Critical habitat for boreal caribou is a habitat state that provides “security” from predation risk and facilitates the effectiveness of their anti-predator strategies. / Ecology

Doctor of Philosophy (PhD) / A renewed focus on generalised pollinator systems has inspired a conceptual framework which highlights that spatial and temporal interactions among plants and their assemblage of pollinators can vary across the individual, population, regional and species levels. Pollination is clearly a dynamic interaction, varying in the number and interdependence of participants and the strength of the outcome of the interaction. Therefore, the role of variation in pollination is fundamental for understanding ecological dynamics of plant populations and is a major factor in the evolution and maintenance of generalised and specialised pollination systems. My study centred on these basic concepts by addressing the following questions: (1) How variable are pollinators in a generalised pollination system? To what degree do insect visitation rates and assemblage composition vary spatially among populations and temporally among flowering seasons? (2) How does variation in pollinators affect plant reproductive success? I chose to do this using a model system, Trachymene incisa subsp. incisa (Apiaceae), which is a widespread Australian herbaceous species with simple white flowers grouped into umbels that attract a high diversity of insect visitors. The Apiaceae are considered to be highly generalist in terms of pollination, due to their simple and uniform floral display and easily accessible floral rewards. Three populations of T. incisa located between 70 km and 210 km apart were studied over 2-3 years. The few studies investigating spatial and temporal variation simultaneously over geographic and yearly/seasonal scales indicate that there is a trend for more spatial than temporal variation in pollinators of generalist-pollinated plants. My study showed both spatial and temporal variation in assemblage composition among all populations and variation in insect visitation rates, in the form of a significant population by year interaction. However, removing ants from the analyses to restrict the assemblage to flying insects and the most likely pollinators, resulted in a significant difference in overall visitation rate between years but no difference in assemblage composition between the Myall Lakes and Tomago populations. These results indicate more temporal than spatial variation in the flying insect visitor assemblage of T. incisa. Foraging behaviour provides another source of variation in plant-pollinator interactions. Trachymene incisa exhibits umbels that function as either male or female at any one time and offer different floral rewards in each phase. For successful pollination, pollinators must visit both male and female umbels during a foraging trip. Insects showed both preferences and non-preferences for umbel phases in natural patches where the gender ratio was male biased. In contrast, insects showed no bias in visitation during a foraging trip or in time spent foraging on male and female umbels in experimental arrays where the gender ratio was equal. Pollinator assemblages consisting of a mixture of different pollinator types coupled with temporal variation in the assemblages of populations among years maintains generalisation at the population/local level. In addition, spatial variation in assemblages among populations maintains generalisation at the species level. Fire alters pollination in T. incisa by shifting the flowering season and reducing the abundance of flying insects. Therefore, fire plays an important role in maintaining spatial and temporal variation in this fire-prone system. Although insect pollinators are important in determining the mating opportunities of 90% of flowering plant species worldwide, few studies have looked at the effects of variation in pollinator assemblages on plant reproductive success and mating. In T. incisa, high insect visitation rates do not guarantee high plant reproductive success, indicating that the quality of visit is more important than the rate of visitation. This is shown by comparing the Agnes Banks and Myall Lakes populations in 2003: Agnes Banks received the highest visitation rate from an assemblage dominated by ants but produced the lowest reproductive output, and Myall Lakes received the lowest visitation rate by an assemblage dominated by a native bee and produced the highest seedling emergence. Interestingly, populations with different assemblage composition can produce similar percentage seed set per umbel. However, similar percentage seed set did not result in similar percentage seedling emergence. Differences among years in reproductive output (total seed production) were due to differences in umbel production (reproductive effort) and proportion of umbels with seeds, and not seed set per umbel. Trachymene incisa is self-compatible and suffers weak to intermediate levels of inbreeding depression through early stages of the life cycle when seeds are self-pollinated and biparentally inbred. Floral phenology, in the form of synchronous protandry, plays an important role in avoiding self-pollination within umbels and reducing the chance of geitonogamous pollination between umbels on the same plant. Although pollinators can increase the rate of inbreeding in T. incisa by foraging on both male and female phase umbels on the same plant or closely related plants, most consecutive insect movements were between plants not located adjacent to each other. This indicates that inbreeding is mostly avoided and that T. incisa is a predominantly outcrossing species, although further genetic analyses are required to confirm this hypothesis. A new conceptual understanding has emerged from the key empirical results in the study of this model generalised pollination system. The large differences among populations and between years indicate that populations are not equally serviced by pollinators and are not equally generalist. Insect visitation rates varied significantly throughout the day, highlighting that sampling of pollinators at one time will result in an inaccurate estimate and usually underestimate the degree of generalisation. The visitor assemblage is not equivalent to the pollinator assemblage, although non-pollinating floral visitors are likely to influence the overall effectiveness of the pollinator assemblage. Given the high degree of variation in both the number of pollinator species and number of pollinator types, I have constructed a model which includes the degree of ecological and functional specialisation of a plant species on pollinators and the variation encountered across different levels of plant organisation. This model describes the ecological or current state of plant species and their pollinators, as well as presenting the patterns of generalisation across a range of populations, which is critical for understanding the evolution and maintenance of the system. In-depth examination of pollination systems is required in order to understand the range of strategies utilised by plants and their pollinators, and I advocate a complete floral visitor assemblage approach to future studies in pollination ecology. In particular, future studies should focus on the role of introduced pollinators in altering generalised plant-pollinator systems and the contribution of non-pollinating floral visitors to pollinator assemblage effectiveness. Comparative studies involving plants with highly conserved floral displays, such as those in the genus Trachymene and in the Apiaceae, will be useful for investigating the dynamics of generalised pollination systems across a range of widespread and restricted species.

No description available.

The genetic diversity and genetic structure of populations, and the processes shaping gene flow within and between populations, are influenced by the landscapes they occur within. Within terrestrial landscapes, rivers and their riparian habitat are among the most dynamic, diverse and complex of landscapes and their linear structure appears as an interlinking feature across large landscapes. This thesis took a landscape genetics approach to examine the influence of river landscape features on Alnus glutinosa populations, a widespread keystone tree species of European riparian ecosystems. By accounting for the differing dispersal mechanisms of A. glutinosa (wind and water), landscape effects on seed- and pollen-mediated gene flow, genetic diversity, demographic and genetic structure were identified at different spatial scales of a large UK river catchment. Widespread gene flow within and between A. glutinosa populations was identified with no apparent limitation of wind-mediated pollen dispersal. Hydrochorous dispersal of seed between populations was evident, and found to increase genetic connectivity between riparian populations; however an isolation by distance effect was identified between populations located further apart from each other. No pattern of genetic diversity was found, with high levels of genetic diversity identified at all spatial and temporal scales. At the river-catchment scale no genetic clustering was observed, either within or between the six rivers studied. Demographic structuring within A. glutinosa populations was evident, and correlated with distance from the main river channel. Interactions between seed dispersal, hydrological disturbance, colonisation, and historical influences are discussed in relation to fine-scale spatial genetic structure between A. glutinosa sapling and adult generations. Central to the landscape genetics approach taken in this thesis was the incorporation of key A. glutinosa life history attributes. By incorporating gene flow analyses, species ecology and landscape features, the research presented here furthers our understanding of riverine landscape influences on their riparian populations at different spatial scales and can be used to inform management principles.

No description available.

Kinlock is a freshwater mussel shell ring site located in Sunflower County in the Mississippi Delta. Little work has been done at freshwater mussel shell rings, and therefore little is known about them. This thesis uses four different data collection methods to answer questions of chronology, site layout, etc. These four methods are controlled surface collection, excavation, coring, and magnetometry. Based on the results of these methods, Kinlock was found to be a Woodland period mussel shell ring with a later Mississippian period component built on top of the shell. This later component consisted of five mounds situated around a plaza. It was also found that the plaza was planned and maintained from the Woodland period through the Mississippian period, until the site was abandoned.

Juniper (Juniperus spp.) encroachment into sagebrush (Artemisia spp.) and bunchgrass communities has reduced understory plant cover and allowed juniper trees to dominate millions of hectares of semiarid rangelands. Trees are mechanically masticated or shredded to decrease wildfire potential and increase desirable understory plant cover. When trees are masticated after a major increase in tree population density and associated decrease in perennial understory cover, there is a risk that invasive annual grasses will dominate because they are highly responsive to the increased resource availability that commonly follows removal of the main resource user. To determine if tree mastication increases resource availability and subsequently favors invasive annual or perennial grasses, we compared soil temperature, water, and nutrient microenvironmental conditions and seedling establishment and growth. We used the major rangeland weed, cheatgrass (Bromus tectorum L.), to represent invasive annual grasses and Anatone bluebunch wheatgrass (Pseudoroegneria spicata (Pursh) A. Löve), a natural accession of native bluebunch wheatgrass, to represent the perennial grasses of the sagebrush-bunchgrass plant community. These comparisons were made between and within paired-adjacent masticated and untreated areas at three locations in Utah dominated by Utah juniper (Juniperus osteosperma (Torr.) Little). Juniper tree mastication generally increased resource availability with masticated areas having greater soil temperature, soil water availability, and soil N supply rates than untreated areas. Prior to juniper tree mastication litter mounds were not found to be resource islands probably because juniper trees themselves were using subcanopy soil water and nutrients. After juniper tree mastication and elimination of these predominant resource users, litter mounds served as resource islands with greater soil water availability and N supply rates than bare interspaces during the critical time for seedling establishment in spring. Plant growth followed in line with greater resource availability after tree mastication with masticated areas having more productive although fewer invasive-annual and perennial grass seedlings than untreated areas. These results suggest that increases in resource availability and warmer spring temperatures associated with mastication will not necessarily favor invasive annual over perennial grass seedling establishment. Resilience of the sagebrush-bunchgrass community to return to dominance after juniper control will likely be greatly influenced by how much of the sagebrush-bunchgrass community remains following tree control and the intensity of propagule pressure by invasive species. If only invasive annuals remain when the trees are treated then invasive annuals would be expected to dominate the post-treatment plant community especially with their ability to establish inside litter mounds unless they were also controlled and perennial grasses planted at the time of treatment.

Syftet med denna uppsats var att undersöka om det är möjligt att förutse var de okända fångstgropar befinner sig i landskapet med hjälp av karaktärsdragen från de kända lämningarna. Målet med detta arbete är att identifiera och registrera okända fångstgropar för att skydda dem från att skadas av skogsbruket. Denna studie förhåller sig till ett undersökningsområde (kommunerna Karlskoga och Degerfors) men diskuterar även andra fångstgropar i Sverige. Under detta arbete har de kända fångstgroparna nyttjats på flera olika sätt; med hjälp av terrängskuggning granskades området vid fångstgroparna i syfte att se om det fanns ytterligare gropar i närheten samt att studera hur en grop kunde synas i terrängskuggningen. Sedan fältbesöktes de kända fångstgroparna för att kunna sammanställa deras karaktärsdrag och med hjälp av detta, samt tidigare forskning och kartanalyser kunde fem karaktärsdrag uppmärksammas. I slutändan upptäcktes åtta okända fångstgropar, varav tre är kvalitetssäkrade av Ulf Eriksson, arkeolog på Skogsstyrelsen och är/kommer bli registrerade i Kulturmiljöregistret. Uppsatsen diskuterar också det nuvarande kunskapsläget om fångstgropar till olika bytesdjur, samt jakt- och fångstmetoderna vargskall och varggård. Detta har dessvärre inte kunnat nå en slutsats då det fanns alldeles för många osäkra faktorer i tidigare forskning som påverkade uppsatsen negativt. / This paper aims to investigate whether it is possible to predict the location of the unknown pits in the landscape using the characteristics of the known remains. The goal of this work is to identify and register unknown pitfalls to protect them from being damaged by forestry. This study relates to an investigation area (the municipalities of Karlskoga and Degerfors) but also discusses other catch pits in Sweden. During this work, the known catch pits have been used in several different ways; with the help of terrain shading, the area at the trapping pits was examined to see if there were additional pits nearby and to study how a pit could be seen in the terrain shading. The known trapping pits were then visited in the field to be able to compile their characteristics. With the help of previous research and map analyses, five characteristics could be noticed. In the end, seven unknown trap pits were discovered, three of which are quality assured by Ulf Eriksson, archaeologist at the Swedish Forest Agency. The thesis also discusses the current state of knowledge about trapping pits for various prey animals, as well as the hunting and trapping methods wolf hunting and wolf enclosure. Unfortunately, this has not been able to reach a conclusion as too many uncertain factors in previous research affected the essay negatively.

Ce mémoire a pour but de documenter et de comprendre le schème d’établissement des Iroquoiens du Saint-Laurent qui ont séjourné sur le site Irving à la fin du Sylvicole supérieur. La recherche a donc été organisée en trois volets. La première interrogation portait sur la compréhension du schème d’établissement à l’échelle du site. Pour atteindre l’objectif, une analyse de l’assemblage culturel du site a été effectuée et l’emplacement du site a été étudié. Le deuxième volet portait sur des relations possibles entre deux villages iroquoiens contemporains de la région, Droulers et Mailhot-Curran, et le site Irving. Des comparaisons morpho-stylistiques de la céramique ont donc été faites entre les trois sites. Finalement, le troisième volet de recherche était d’évaluer le type d’établissement du site Irving. Des comparatifs entre sites iroquoiens du Sylvicole supérieur récent ont donc été effectués. Nous avons répertorié divers types de sites documentés tels que des villages, des hameaux, des hameaux saisonniers et des camps spécialisés. Nous sommes venus à la conclusion que le site Irving est un hameau saisonnier horticole qui était probablement rattaché à un village environnant, possiblement celui de Mailhot-Curran. / The purpose of this thesis is to document and understand the settlements pattern of the St. Lawrence Iroquoians with a particular focus on those living at the Irving site during the Terminal late Woodland period. The research is comprised of three parts. The first research objective was to document the settlement pattern characteristics at the Irving site. In order to achieve this objective, an analysis of the site's cultural materials was carried out in reference to the site’s size and location. The second research topic focused on the possible relationships between two contemporary archaeological villages in the region, Droulers and Mailhot-Curran, and the Irving site, in which a morpho-stylistic comparison of the ceramics was conducted between these three sites. Finally, the third research topic focused on identifying the type of settlement site of the Irving site. Comparisons between Iroquoians sites of the Terminal late Woodland period and Irving were therefore carried out. In exploring various types of sites documented elsewhere, such as villages, hamlets, seasonal hamlets and specialized camps. We came to the conclusion that the Irving site appears to be have been a seasonal horticultural hamlet that was probably associated with a nearby village, possibly Mailhot-Curran.

Graduation date: 2006 / Habitat loss causes a reduction in available resources for wildlife, alters the configuration of remaining habitat, and may isolate wildlife populations. White-breasted nuthatches (Sitta carolinensis) are experiencing long-term population declines in the Willamette Valley of Oregon, where they are historically associated with oak woodlands. As secondary cavity-nesters, white-breasted nuthatches may be limited by the availability of existing cavities for nesting and roosting. Oak vegetation in the Willamette Valley has changed since European-American settlement times from vast areas of open oak savanna to isolated closed-canopy stands separated by agricultural fields. We examined nuthatch density, nest cavity selection, and nest success in relation to oak woodland structure and landscape context. We conducted point transect surveys in 3 strata: woodland interiors, large woodland edges, and small woodlands. We located and monitored nuthatch nests and sampled vegetation at nest locations and matching random locations around each nest. Woodland structure and edge density were measured at a 178-m radius (home range) scale, and landscape context was measured using vegetation cover within a 1-km radius around point transects and nests. We used program DISTANCE to fit detection functions and calculate nuthatch densities. We used conditional logistic regression to compare nest locations with random locations, and analyzed nest success with Mayfield logistic regression. White-breasted nuthatch density was significantly higher in small woodlands than in edges of large woodlands, which had higher nuthatch density than woodland interiors. Density of nuthatches increased with a combination of oak cover within a 1-km radius of the point, edge density within a 178-m radius, and number of oak trees >50 cm diameter at breast height (dbh) within a 100-m radius. Nest cavities were situated in oak trees containing more cavities than random oak trees that had cavities, and oak trees used as nest trees had a larger dbh than oak trees within random plots. Local woodland structure at nest locations was characterized by larger trees, measured by greater mean dbh, canopy cover, and basal area of oaks than random locations within the home range. Nest success in natural cavities was 71% and was not predicted by attributes of nest cavities, nest trees, local woodland structure at nests, woodland structure at the home range scale, or landscape context. These results suggest that the most suitable habitat for white-breasted nuthatches in the Willamette Valley includes oak woodlands in close proximity to one another with a high proportion of edge and mature oak trees. Managers should preserve trees containing cavities and large oak trees whenever possible. Thinning of small oaks and removal of conifers in oak woodlands to create more open, savanna-like conditions may also promote the development of larger oaks with more spreading branches, providing more opportunities for cavities to form and more foraging surface area for nuthatches.