Epibiosis is an ecological relationship that has been described as one of the closest possible associations in marine ecosystems. In the space limited rocky intertidal, mussel beds provide important secondary space for barnacles. The epibiotic relationship between mussels and barnacles on the south-east coast of South Africa was considered at different scales, from large-scale, natural patterns of epibiosis on the rocky shore, to fine-scale settlement choices of barnacles and the effects on the condition and growth rates of individual mussels. Mussel and barnacle assemblages were generally stable over a 12-month period. The tracking of individual mussels with and without barnacle epibionts resulted in a significant increase in mortality rate of mussels with epibionts over 12 months (two-way ANOVA, p = 0.028). Barnacles on rocks, as well as on mussels, were also tracked with no significant effect of substratum on mortality of barnacles (two-way ANOVA, p = 0.119). Prevalence and intensity of barnacle infestations was also examined in relation to coastline topography on two co-occurring mussel species, the indigenous Perna perna and invasive Mytilus galloprovincialis. The results were complex, but bay status had significant effects on prevalence and intensity for both mussel species, depending on the time and zone. The effect of bay in relation to time was particularly relevant for M. galloprovincialis (four-way nested ANOVA, Season X Site(Bay): p = 0.0002), where summer prevalence was higher than that of winter in bays, regardless of zone, while in open coast sites, the effect of season was only significant in the mid zone. Patterns of intensity generally showed higher values in summer. Substratum preference by barnacles was investigated by recording settlement, survival and mortality of Chthamalus dentatus barnacles on various treatments. There was a strong preference for the rock-like plastic substratum by primary settlers (pair-wise tests of PERMANOVA: Dead < Rock mimic (p = 0.0001); Replica < Rock mimic (p = 0.019) and Live < Rock mimic (p = 0.0001)). This indicates that barnacles settle on mussel shells only as a secondary choice and that micro-topography is an important variable in barnacle settlement. The effect of barnacle epibiosis on condition index and growth of P. perna and M. galloprovincialis was also examined as a direct indication of the health of mussels subjected to the biological stress of epibiosis. Although not significant (PERMANOVA: P. perna: p(perm) = 0.890; M. galloprovincialis: p(perm) = 0.395), growth for both mussel species was slower for barnacle-infested individuals in summer, which is the main growing season for mussels in the region. Results from condition index calculations, however, showed no negative impacts of epibiotic barnacles (three-way ANCOVA: P. perna: p = 0.372; M. galloprovincialis: p = 0.762). Barnacle epibionts create a new interface between the mussel and its environment and this interaction can affect other members of the community. The possibility of the barnacle epibiont causing increased drag also needs further investigation. Biological processes operating within a wide range of physical stressors drive the interactions on the rocky shore, such as epibiosis. Overall, the results of this study suggest that the epibiotic relationship between mussels and barnacles on the south-east coast of South Africa does not significantly affect the mussel species present and that barnacles only use mussel shells as a secondary choice of substratum.

Thesis (MTech (Nature Conservation))--Cape Peninsula University of Technology, 2010. / Many urban rivers are heavily engineered and no longer function naturally. The City of Cape Town has designed and implemented many stormwater and river management projects. Very little monitoring has occurred as to whether these engineering projects and remediation measures, have had a positive effect on our urban river ecosystem function. The study investigated the influence of specific engineering interventions such as the placement or rocks in stream to create weirs, gabion lining of stream channels, removal of canal walls, establishment of artificial wetlands, and approaches to urban river management, on river ecosystem function of the Keysers River, Little Lotus River, Langevlei Canal, Silvermine River, Moddergat River and Big Lotus River, as measured by specific indices including water chemistry and aquatic community structure (macroinvertebrates and diatoms).

The most recent surveys in South Africa estimate that invasions are still increasing, despite substantial clearing efforts. Riparian systems in South Africa are particularly vulnerable to invasion by woody IAPs. This thesis addresses the restoration of alien‐invaded riparian systems, by investigating the factors that facilitate or constrain spontaneous recovery and influence the trajectories of succession. These factors include invasion history and management history, especially the use of fire. A seedling emergence approach was used to test the presence of a viable pre‐fire seedbank, and the effect of fire on the seed bank. The efficacy of some active restoration interventions was also tested, with the aim to return invasion‐resistant, indigenous vegetation with a structure and function representative of uninvaded sites. The findings of this study indicated the presence of a viable and persistent riparian soil seed bank, even after 30 years of intermtittent invasion as well as two fire cycles under invasion. It shows that the management practice of fell‐and‐burn resulted in high soil temperatures, and that this reduced the indigenous soil seed bank density, especially in the upper soil layer. Clear germination sequences and patterns of emergence over time for different species were observed during this study, with many species exhibiting delayed emergence relative to the timing of the fire event. It is proposed that manipulation of the season of fire could be used to selectively optimise the order of arrival and therefore superior recruitment of some species over others in the Eastern Cape fynbos, and thus alter the trajectories of recovery of vegetation towards a more desired state. Active restoration in the form of indigenous seed and plant additions resulted in a significantly higher indigenous cover after seven months, compared to a control (passive restoration) or restoring with grass. Indigenous cover and composition was also strongly influenced by lateral zonation, and some key guilds and species were missing or present in much lower densities compared to reference sites. Grass restoration significantly suppressed the regeneration of A. longifolia, as well as the regeneration of indigenous species. Biotic resistance can thus be achieved through restoration, and it could be a powerful tool in the management of IAPs, although the deliberate introduction of grass after clearing in fynbos also reduces biodiversity and could have unforeseen consequences to riparian function.

Rehabilitation is a dynamic process influenced by factors related to more than one field of ecology. It is therefore necessary to consider all these components when assessing the rehabilitation, although in the initial stages the successful revegetation of the disturbed areas is the most important criterion. Richards Bay Minerals, on whose mining site this project was carried out, is dredge mining heavy minerals on the north coast of Natal, where they have rehabilitated mined areas since 1978. This project has been carried out to establish: 1) The success of their dune forest rehabilitation using quantitative techniques. 2) The available seed bank in their rehabilitation stands. 3) The similarities in the succession taking place in rehabilitation stands compared to the revegetated stands in the vicinity of Richards Bay. 4) The best methods for creating alternative vegetation communities, especially grasslands, with a high species diversity on the mined tailings. This study reviews only the success of rehabilitation of the natural vegetation but other studies focusing on the insect, reptile, mammal and bird populations are also being undertaken by other researchers. No particular method of determining the success of vegetation rehabilitation has been chosen by restoration ecologists. Therefore in this study a broad range of quantitative techniques were used to show whether successional changes are occurring in the vegetation and physical environment. The results obtained from sampling the rehabilitated vegetation have shown that both the species richness and diversity are increasing as the returned vegetation matures. Levels of soil properties such as Sodium, Phosphate, Calcium and percentage organic matter have also risen with increasing stand age. Community complexity is also increasing with stand age, and TWINSPAN and DECORANA plots have separated out the differently aged stands based on their differences. A "pilot" study was done on the seed bank present in the rehabilitation stands. This has shown the presence of large amounts of early successional, mostly herbaceous species. Seeds of later successional and woody species were scarce which may be a result of the sampling intensity used. However seeds of late successional ground cover species were found in the older stands. Comparisons between the natural revegetation of disturbed areas in the vicinity of Richards Bay and the rehabilitation stands revealed similarities in both species composition and complexity. Species richness and diversity values are comparatively similar for the younger revegetated and older rehabilitation stands, and lWlNSPAN and DECORANA analysis techniques clustered the samples recorded from these areas in close proximity on their relative plots. The oldest revegetated sites contain a number of species found in the rehabilitated vegetation but as Acacia karroo has thinned-out in these stands many of these other species are now mature individuals. Attempts at rehabilitating an area of grassland at Richards Bay Minerals has not produced satisfactory species diversity and experimental manipulations were used to try and increase the diversity of the existing Eragrostis curvula dominated community. Of the several treatments used for the manipulation, a combination of burning and further topsoiling was the most successful in reducing Eragrostis importance and in increasing the species richness. Grassland topsoil spread directly onto the bare tailings produced an extensive vegetation covering over a short period but species richness was not significantly greater than for the existing Eragrostis dominated grassland, and further treatments and management needs to continue if this technique is to be employed. Only a limited amount of alien infestation of the rehabilitated areas was evident from the sampling undertaken during this research. As the removal of alien plants is an ongoing process and the rehabilitation stands are continuously monitored to identify any new invaders, this is not expected to become a problem. From the results of work done overseas and the rehabilitation carried out in South Africa it appears that it is possible to return natural vegetation communities on mined areas. That this is a lengthy process is to be expected but by manipulating the vegetation and continuously monitoring the process it may be possible to speed up development. Areas in need of further research have been identified based on the findings of this project. This will help to reinforce the undertaking of management proposals that will enhance the vegetation recovery and the success of the rehabilitation programme.

The structuring of ant assemblages in a Southern African savanna was investigated using data from the only long-term, large-scale savanna fire experiment in Africa. A comprehensive survey of three habitats in the Kruger National Park (KNP) revealed a total of 169 ant species from 41 genera. The sampling efficiency and consistency of pitfall traps and Winkler samples for inventory, bioindicator and ecological studies in savanna habitats was compared using ants. Pitfall traps were more efficient and productive than Winkler sampling for epigaiec ants, with a greater total species richness and higher abundance of ants recorded. Suggestions were made to improve Winkler sampling output, and to allow quantitative data to be collected. With the structuring of local assemblages, competition was the most significant local factor tested. The relationship between ant dominance and ant species richness was consistent across three continents. A model developed to test mechanisms that could be responsible for the form of this relationship supported the hypothesis that competitive exclusion by dominant ants at least partially reduces species richness. Stress was only partially responsible for low dominance and low species richness, while scatter in the data points is related to patchiness of ants at baits. These findings contrast strongly with previous claims regarding the relationship between richness and dominance. Habitat complexity was not found to play an important role in determining ant assemblage body size in this savanna system. The size-grain hypothesis (Kaspari&Weiser 1999) which predicts that environmental rugosity results in positive allometric scaling of leg length on body length because of changes in locomotion costs, was tested by comparing the body sizes of ants from areas of contrasting habitat complexity. No support for the hypothesis was found. Phylogenetic independent contrast methods did however support the allometric relationship found by Kaspari and Weiser (1999). Ant assemblages in KNP exhibited a remarkable degree of resistance, and in some cases resilience, to burning. Species richness or abundance did not vary with different burning treatments, although ant assemblage composition was sensitive to burning treatment. This difference, however, was only pronounced between burnt and unburnt plots, not between burning treatments. The degree of response of ant assemblages is likely to be related to two main contributory factors: mean annual rainfall and changes in vegetation structure with burning, and the assemblage's history of association with fire. An overview of published research on the effects of fire on fauna in Southern Africa was undertaken. Few studies have examined the effects of fire on amphibians or reptiles and few experimental studies have been undertaken using an experimental fire regime applied over appropriately long time intervals. Most studies provided no information on the scale of the study. Replication was often not reported, and was generally inadequate. Information on the effects of fire on fauna in Southern Africa is fragmentary, and consequently informed management decisions regarding the consequences of burning policies on the conservation of biodiversity both within and outside protected areas are problematic. Recommendations and suggestions for improving fire research are given. / Thesis (DPhil (Zoology))--University of Pretoria, 2005. / Zoology and Entomology / unrestricted

The main aim of this study was to assess the “top-down” and “bottom-up” control of bacterial production in the small temporarily open/closed East Kleinemonde Estuary, situated on the south-eastern coastline of southern Africa. Spatial and temporal patterns in bacterial abundance, biomass and production and the importance of abiotic and biotic factors were investigated over the period May 2006 to April 2007. The trophic interactions between bacteria, phytoplankton, nanoflagellates (< 20 μm), microzooplankton (< 200 μm) and mesozooplankton (< 2 000 μm) were investigated during winter and summer. Bacterial abundance, biomass and production ranged between 1.00 × 10⁹ and 4.93 × 10⁹ cells 1⁻¹, 32.4 and 109 μg C 1⁻¹ and 0.01 and 1.99 μg C 1⁻¹ h⁻¹, respectively. With a few exceptions there were no spatial patterns in the values. Bacterial abundance, biomass and production, however, demonstrated a distinct temporal pattern with the lowest values consistently recorded during the winter months. Nanoflagellate and bacterial abundances were significantly correlated to one another (lower reaches: r = 0.818, p < 0.001; middle reaches: r = 0.628, p < 0.001; upper reaches: r = 0.484, p < 0.05) suggesting a strong predator-prey relationship. The frequency of visibly infected bacterial cells and the mean number of virus particles within each bacterial cell during this study demonstrated no temporal or spatial patterns and ranged from 0.5 to 6.1 % and 12.0 to 37.5 virus particles per bacterium, respectively. Viral infection and lysis was thus a constant source of bacterial mortality throughout the year. The estimated percentage of bacterial production removed by viral lysis ranged between 7.8 and 88.9% of the total which suggests that viral lysis represented a very important source of bacterial mortality during this study. The biological interactions between the selected components of the plankton community demonstrated that among the heterotrophic components of the plankton, the nanoflagellates were identified as the most important consumers of bacteria and small phytoplankton cells (< 20 μm). In the presence of microzooplankton the impact of the nanoflagellates on both the bacteria and phytoplankton was reduced, indicating that larger heterotrophs were preying upon the nanoflagellates. Mesozooplankton, however, appeared to exert the greatest impact on nanoflagellates. In the cascading experiments, the data suggest that mesozooplankton consume nanoflagellates, which resulted in a decrease in the predation impact of these organisms on the bacteria. This result is consistent with predator-prey cascades. The presence of the larger heterotrophs therefore, mediates the interactions between the primary bacterivores, the nanoflagellates, and the bacteria within the temporarily open/closed East Kleinemonde Estuary.

The nutrient dynamics in two contrasting estuaries and in the adjacent nearshore environment along the south-east coast of South Africa was investigated seasonally. Due to an inter-basin transfer of water from the Gariep Dam to the Great Fish River, the Great Fish estuary is a fresh water dominated, terrestrially driven system with an annual fresh water inflow of 250 – 650 x 10⁶ m³ per year. In contrast, the Kariega estuary is a fresh water deprived, marine dominated system with a fresh water inflow estimated at 2.5 – 35 x 10⁶ m³per year. The reduced fresh water inflow into the estuary is attributed to regular impoundments along the Kariega River. Water samples were collected from surface and subsurface layers along the length of the estuaries as well as from a series of transects occupied in the nearshore environment. Samples were analysed for nitrate, nitrite, ammonium, phosphate and silicate. Temperature and salinity were recorded at each station. A Land-Ocean Interactions in the Coastal Zone (LOICZ) budget was constructed for each estuary to describe the role of ecosystem-level metabolism as either a sink or a source of phosphorus, nitrogen and carbon. Seasonal variation in physico-chemical properties and nutrient concentrations in the Kariega estuary was minimal due to constant low inflow, while in the Great Fish estuary, concentrations varied in response to changes in flow rate. Nutrient concentrations were consistently higher in the Great Fish estuary than in the Kariega estuary, largely reflecting differences in fresh water inflow. During periods of high flow (32.92 m³.s⁻¹in the Great Fish River) dissolved inorganic nitrogen (DIN) concentrations in the Great Fish estuary were an order of magnitude higher than those recorded in the Kariega estuary. Results of the LOICZ budgeting procedures revealed that in spite of the contrasting hydrodynamic features, the estuaries behave in largely the same manner – both predominantly sources of nutrients with heterotrophic processes dominating over autotrophic actions and both were net denitrifyers during all surveys. This was, however, due to different sets of processes operating in the two estuaries, namely low nutrient concentrations resulting in microbial activity in the Kariega estuary, and riverine influx of nutrients and phytoplankton combined with a short residence time of the water in the Great Fish estuary. In the marine nearshore environment, higher nutrient concentrations were recorded adjacent to the Great Fish estuary than offshore of the Kariega estuary. This was due to a surface plume of less saline water leaving the Great Fish estuary, which acted as an ‘outweller’ of nutrients. Offshore of the Kariega estuary, on the other hand, the nutrient concentrations were characteristic of marine waters due to a lack of fresh water outflow from the estuary. Nutrient concentrations in the marine environment adjacent to the Kariega estuary were, at times, higher than those recorded within the estuary. This observation supports previous statements which suggest that the Kariega estuary is not an ‘outweller’ of dissolved nutrients and particulate material, but rather an extension of the marine environment.

In spite of the relative simple vegetation structure, the Grassland biome has surprisingly high species diversity. The Grassland biome is also the most transformed biome in South Africa, with cultivation having the largest impact. When croplands are abandoned, secondary succession leads to low diversity Hyparrhenia hirta dominated grassland. A combination of two seed mixtures, two seeding densities and two establishment methods was established in plots on a recently abandoned cropland at Suikerbosrand Nature Reserve to evaluate their effect on secondary succession. The rip plots, where more resources were available between the rip lines, have shown higher densities of relic weeds as well as local perennials, showing some progressive successional movement. However, Hyparrhenia hirta was one of the non-sown perennials increasing in the rip plots. Hyparrhenia invasion and relic weeds were best controlled in the plough plots. Although Hyparrhenia was successfully controlled in plough plots, no secondary succession occurred in these treatments. / Agriculture, Animal Health & Human Ecology / M. Tech. (Nature Conservation)

Thesis (MSc)--University of Stellenbosch, 2003. / ENGLISH ABSTRACT: Arid and semi-arid rangelands of the Nama-Karoo Biome are believed to have changed considerably since the arrival of domestic livestock in the veld. Severe grazing pressure is considered to be one of the prime factors responsible for the perceived degradation of vegetation and concurrent soil degradation. To understand the process of degradation and to make further recommendations for future veld restoration, a large-scale project was undertaken in the Eastern Cape. This project focused on the role that landscape heterogeneity plays in providing refuges for plant species. The key question asked in this project was: What role do these hypothetically less grazed mesas play in the conservation of rangelands in the Nama-Karoo of the Eastern Cape? This study, a component of the larger project, investigated grazing and soil landscape gradients on- and off- the three mesas (Tafelberg, Folminkskop and Buffelskop) in the Middelburg District of the Eastern Cape, South Africa and the possible influences that they might have on the veld. The flats surrounding the mesas were mostly used as grazing camps in contrast to the plateaux of the mesas, which had varied levels of accessibility. In the absence of direct observations, the primary objective of the study was to test the differential dung pellet abundance and impact of animals on different parts of the landscape. As such, dung pellet counts in this study were correlated with surrogates of soil physical properties including bare ground, trampling and litter cover. Variation in dung pellet density was found at the different habitats (flats, slopes and plateaux) of all mesas. The flats to the north-eastern and south-western of Tafelberg mesa were found to be more heavily utilized by livestock and herbivores, while the plateaux and southeastern slopes of Folminkskop and Buffelskop were also utilized by grazers. The Tafelberg mesa was the only study site that was consistent with the hypothesis which stated that grazers would be less concentrated on the plateau compared to the surrounding flats due to its inaccessibility, whilst the high mean dung pellet density on the plateaux of the smaller Folminkskop and Buffelskop mesas due to easier access contradicted the original hypothesis. It appeared that dung pellet density did not clearly turn out to be an indicator of habitat use in this study, but showed where slopes and plateaux were accessible to herbivores, as in the case for the Buffelskop mesa, a higher abundance of dung pellets were found suggesting that higher intensities of habitat use took place. During this study a strong pattern was observed of bare soil patches on the flats surrounding the mesas. There was a decrease in percentage of bare soil along the gradient of the three mesas with a high percentage of bare soil on the plateaux of the mesas. A significant correlation was found between bare soil and dung pellet density. However, the plateaux of Folminkskop and Buffelskop had a high percentage of bare soil compared to the plateau of Tafelberg mesa. Farmers mainly used these smaller mesas as grazing camps for their livestock and herbivores. A positive correlation between bare soil and litter cover of the different habitats was evident in this study. A lower percentage of litter cover at these sites was associated with a high percentage of bare soil. Litter is very important in a healthy vegetation community in terms of nutrient cycling and fertile patches. A detailed assessment of soil chemical and physical properties would reveal, firstly, if vegetation change is better explained by soil or grazing effects and, secondly, if changes in soil have resulted from land use. Differences in macro- and micro-site variations between open-canopy (between shrubs) and closed-canopy (under shrubs) sites for each habitat were determined to differentiate between local scales due to land use and landscape scales due to geomorphology. The results suggested that carbon, phosphorus, calcium, magnesium, copper and manganese levels in soils at landscape scale better explain vegetation changes between habitats. At a local scale (open- and closed-canopy sites) land use was responsible for little changes in soils. Changes in only soil potassium, zinc and boron elements were actually a consequence of local scales due to land use. The soil nutrient content on the slopes appeared to be intermediate between the flats and plateaux of all three mesas. It appeared that dolerite capped Tafelberg and Folminkskop mesas had high silt and clay content, while Buffelskop (sandstone) mesa had a lower silt and clay content. Consequently, the texture and parent material of the soils contributed to the variations in soil nutrient composition between these mesas. High infiltration rate together with low nutrient content on the flats clearly showed that these flats, surrounding the mesas were degraded. The high infiltration rates were caused by high activity by livestock and other indigenous animals on the flats which breaks the surface crusting of bare soil and improve infiltration. It was concluded that high levels of grazing at these sites have also altered the textural and soil properties. Endozoochory dispersal and the deposition of dung pellets in areas of small patch disturbances play an important role in veld regeneration in degraded areas. Dung pellets collected from permanent study sites on the southeastern and northwestern flats and slopes, and all the study sites on the plateau of Tafelberg mesa, was sown in seedling trays, watered and monitored for seedling germination. Species list were then compared to below-ground soil seed bank data and above-ground vegetation data collected by other researchers at the same permanent study sites. Higher seedling percentages were recorded from dung pellets collected on the flats than on the plateau. A total of sixteen species were found to germinate in dung pellets collected on the flats compared to ten species germinating in dung pellets collected on the slopes and two species on the plateau of Tafelberg mesa. The seeds that germinated represent a variety of palatable grasses and shrub species. Aristida sp., Eragrostis bicolor, Eragrostis ch/orome/as and Eragrostis obtusa were palatable grasses recorded for dung pellets collected on the flats. Of the species recorded, Aristida sp., Chenopodium sp. and Pentzia sp. were found in dung pellets but were not recorded in parallel soil seedbank and vegetation studies. Successful restoration of veld conditions requires strict grazing management practices. Germination of seed in dung pellets might be considered to be a valuable means of indicating restoration potential and rangeland conditions for the identification of both degraded and conservation worthy areas. With appropriate land management skills and restoration measures, these challenges can be constructively and creatively faced. / AFRIKAANSE OPSOMMING: Daar word geglo dat die ariede en sernl-arlede weivelde van die Nama-Karoo bioom aansienlik verander het sedert die aankoms van mak lewende hawe in die veld. Swaar weidingsdruk word beskou as een van die vernaamste faktore verantwoordelik vir die waarneembare oorbeweiding van plantegroei en die gelyktydige grond agteruitgang (degradasie). Ten einde die proses van degradasie beter te verstaan, en om verdere aanbevelings te kan maak vir toekomstige veld herstel, is huidiglik 'n grootskaalse projek in die Oos-Kaap aan die gang oor die rol wat landskap ongelyksoortigheid speel in die voorsiening van skuiling aan plant spesies. Hierdie studie ondersoek weiding en grondlandskap gradiënte (op en af) van die drie mesas (Tafelberg, Folminkskop en Buffelskop) in die Middelburg streek van die Oos- Kaap, Suid-Afrika en die moontlike invloed wat hierdie gradiënte op die veld het. Die vlaktes was meestal gebruik vir weidingskampe in teenstelling met die kruin van die mesas. Hierdie studie het duidelike patrone vir habitat-gebruik op die vlaktes, hange en kruine van al die mesas aangetoon. Die vraag wat gestel was, was watter rol speel hierdie hipoteties minder beweide mesas in die bewaring van weivelde in die Nama- Karoo van die Oos-Kaap? Hierdie studie toets nie direk vir mis verspreiding as 'n plaasvervanger vir dier verspreiding in die landskap sedert werklike dier getalle nie oorweeg was nie. In die awesigheid van direkte waarneming, die primêre doelwit was om te toets die differensiaal misdigtheid en die impak van diere op verskillende dele van die landskap. Mis getalle in hierdie studie was gekorreleer met plaasvervangers van fisiese eienskappe insluitent onbedekte grond, vertrapping en droë plant material decking. Variasie in misdigtheid was gevind by die verskillende habitate (vlaktes, hange en kruine) van all die mesas. Die vlaktes aan die noord-oostelike en suid-westelike kante van die Tafelberg mesa was meer hewig benut deur lewende hawe en hêrbivore, terwyl die kruine en suid-oostelike hange van Folminkskop en Buffelskop ook gebruik was deur weidende diere. Die Tafelberg mesa was die enigste studie area wat konsekwent was met die hipotese, terwyl die hoë gemiddeld misdigtheid op die kruine van die kleiner Folminkskop en Buffelskop mesas, as gevolg van makliker toegang weerspreek die oorspronklike hipotese. Dit blyk dat misdigtheid duidelik opkom as 'n nie aanwyser habitat-gebruik in hierdie studie, maar wys well waar hange en kruine toeganklik was vir herbivore, soos in die geval van Buffelskop mesa, oorvloed van mis was gevind wat voorstel dat hoer intensiteit van habitat-gebruik voorgekom het. 'n Duidelike patroon is waargeneem tydens hierdie studie van onbedekte grond areas op die vlaktes rondom mesas. Daar was 'n afname in persentasie onbedekte grond langs die gradiënt van die drie mesas. Die kruine van Folminkskop en Buffelskop toon hoër persentasies onbedekte grond in vergelyking met die kruin van die Tafelberg mesa. Daar is 'n betekenisvolle korrelasie tussen onbedekte grond en misdigtheid gevind. Boere gebruik hierdie kleiner mesas hoofsaaklik as weidingskampe vir hulle lewende hawe en hêrbivore. In hierdie studie is daar 'n positiewe korrelasie gevind tussen onbedekte grond en droë plant material decking van die verskillende habitatte. Droë plantmateriaal bedekking speel 'n baie belangrik rol in 'n gesonde plant gemeenskap in terme van voedingsirkulering en vrugbare kolle. 'n Breedvoerige skatting van grond chemise en fisiese eienskappe wou openbaar eerstens of plantegroei verandering beter verklaar word deur grond of weiding effekte en tweedens, of verandering in grond veroorsaak was deur land-gebruik. Verskille in makro- en mikroterrein-wisseling tussen oop-blaredak (tussenplantruimtes) en toeblaredak (onderplantdekking) terreine vir elke habitat was bepaal om te onderskei tussen lokaal effek as gevolg van land-gebruik en landskap effek as gevolg van geomorfologie. Die resultate stel voor dat koolstof, fosfor, kalsium, magnesium, koper en mangaan vlakke in die grond by landskap effek beter plantegroei verandering tussen habitate verduidelik. By 'n lokaal effek (oop-en toe-blaredak terreine) land-gebruik was verantwoordelik vir min veranderinge in grond. Verandering in grond kalium, sink en baron elemente was werklik 'n gevolg van lokaal effek as gevolg van land-gebruik. Die nutrient inhoud van grond op die hange blyk intermediêr te wees tussen die van die vlaktes en kruine van al drie mesas. Hierdie studie toon ook dat die geologie en topografie van hierdie mesas die grond tekstuur en nutrient samestelling op en af van die mesas beinvloed. Dit wil voorkom asof die dolerite bedekte Tafelberg en Folminkskop mesas, hoë slik en klei materiaal bevat, terwyl Buffelskop (sandsteen) mesa 'n laer slik en klei inhoud het. Gevolglik, tekstueel en ouer materiaal grond dra by tot die variasie in grond nutrient samestelling tussen hierdie mesas. Hoë infiltrasie tempo tesame met lae voedingstof van die vlaktes dui daarop dat die vlaktes rondom die mesas oorbewei was. Hierdie hoë infiltrasie tempo word veroorsaak deur hoë dierlike aktiwieteite op die vlaktes deur lewende hawe en ander inheemse diere wat die oppervlakke van onbedekte grond versteur en infiltrasie verbeter. Die gevolgtrekking word gemaak dat die hoë weidingsvlakke van hierdie terreine ook die tekstuur en ander grond eienskappe verander het. Endozoochoreeuse verspreiding en die mis-afsetting in gebiede van klein areaversteurings, speel 'n belangrike rol in veld herlewing in oorbeweide gebiede. Mis versamel by permanente studie terreine op die SO en NW vlaktes en hange, as ook al die studie terreine op die kruin van die Tafelberg mesa, is gesaai in saailing bakke, besproei en gekontroleer vir saailing ontkieming. 'n Lys van spesies is vergelyk met onder-grondse saadbank gegewens en bo-grondse plantegroei gegewens wat deur ander navorsers by dieselfde studie terreine versamel was. Hoër saailing persentasies is gevind vir mis wat op die vlaktes versamel is as op die kruine. 'n Totaal van sestien geidentifiseerde spesies het uit die mis op die vlaktes ontkiem in vergelyking met die tien spesies wat ontkiem het uit die mis van die hange en twee spesies iut die mis van die kruin van Tafelberg mesa. Die ontkiemde saad verteenwoordig 'n verskeidenheid van vreetbare gras en struik spesies. Aristide sp., Eragrostis bicolor, Eragrostis chloromelas en Eragrostis obtusa is vreetbare grasse wat gevind was in die mis versamelop die vlaktes. Die spesies opname wys dat Aristida sp., Chenopodium sp. en Pentzia sp. Gevind was in die mis, maar nie in die parallelle grond saadbank en plantegroei studies nie. Die suksesvolle herstel van veldtoestande vereis "n streng weidingsbestuur toepassing. Die ontkieming van saad in mis kan beskou word as "n bekostigbare manier om veld herstel potentiaal te bepaal, asook weiveld toestande vir die indentifiseering van beide degradasie en bewaring van waardevolle areas. Met gepaste veldbestuur vaardighede en herstel maatreels, kan hierdie uitdagings konstruktief en kreatief aangespreek word.

Thesis (PhD)--Stellenbosch University, 2012. / ENGLISH ABSTRACT: The restoration of natural capital has ecological, hydrological and economic benefits. Are these benefits greater than the costs of restoration when compared across a range of dissimilar sites? This study examines the impact of restoration at eight case study sites distributed throughout South Africa. The benefits of restoration include improved grazing values and crop yields, improvements in water yield and quality, soil carbon improvements, wild products, lumber, fuelwood and electricity. The impact of restoration on all forms of natural capital (i.e. cultivated, replenishable, renewable and non-renewable) is therefore quantified. The costs of restoration include depreciation on capital expenditure, labour costs, equipment and bond refinancing costs. The literature review done during this study presents three frameworks. The first framework classifies social science using the classification scheme of Burrell and Morgan. It shows that system dynamics modelling and neoclassical economics share the same epistemological and ontological characteristics, both of these fall within the naturalistic paradigm, which also characterises most of scientific research. System dynamics modelling and neoclassical economics, however, digress in the Flood and Jackson classification scheme, which is the second framework for classifying social science. Neoclassical economics is characterised by a small number of elements and few interactions between the elements. Systems dynamics modelling, on the other hand, is characterised by a large number of elements and many interactions between the elements. The nature-freedom ground motive is subject to a number of criticisms, including the fact that it introduces dualistic thinking into the analysis, as well as that it does not adequately address normative or moral issues. The framework of Dooyeweerd, the third framework, is presented as a means of transcending the nature-freedom ground motive. Although the nature-freedom ground motive is largely utilised in this study, the analysis does transcend the traditional economic approach in a number of areas. These include, for example, a focus on transdisciplinary methods, disequilibria, adopting a case study approach, and empirical estimation instead of theoretical models. The restoration case studies in this study are examples of individual complex systems. Eight system dynamics models are developed to model interactions between the economic, ecological and hydrological components of each of the case studies. The eight system dynamics models are then used to inform a risk analysis process that culminates in a portfolio mapping exercise. This portfolio mapping exercise is then used to identify the characteristics and features of the different case study sites based on the risk profile of each sites. This study is the first known application of system dynamics, risk analysis and portfolio mapping to an environmental restoration project. This framework could potentially be used by policymakers confronted with budgetary constraints to select and prioritise between competing restoration projects. / AFRIKAANSE OPSOMMING: Die restorasie van natuurlike kapitaal het ekologiese, hidrologiese en ekonomiese voordele. Maar is hierdie voordele groter as die kostes verbonde aan restorasie wanneer dit oor verskeie ongelyksoortige terreine vergelyk word? Hierdie studie bestudeer die impak van restorasie op agt verskillende studie terreine versprei regoor Suid-Afrika. Die voordele van restorasie sluit die volgende in: beter weiding waardes en oes opbrengste, verbeterde water lewering en water kwaliteit, verbetering van grondkoolstof, wilde produkte, hout, brandstofhout en elektrisiteit. Die impak van restorasie op alle vorme van natuurlike kapitaal (gekultiveerd, aanvulbaar, hernubaar en nie-hernubaar) is daarom gekwantifiseer. Die kostes van restorasie sluit in ‘n vermindering in kapitaal uitgawes, arbeidskoste, toerusting en verband herfinansieringskoste. Die literatuurstudie hou drie raamwerke voor. Die eerste raamwerk klassifiseer sosiale wetenskappe volgens die Burrel en Morgan klassifikasie skema. Dit wys daarop dat dinamiese stelsel modellering en neoklassieke ekonomie dieselfde epistemologiese en ontologiese eienskappe deel; beide val binne die naturalistiese paradigma, wat dan ook meeste wetenskaplike navorsing tipeer. Stelseldinamiese modellering en neoklassieke ekonomie wyk egter af na die Flood and Jackson klassifikasie skema, wat die tweede raamwerk is waarvolgens sosiale wetenskappe geklassifiseer word. Neoklassieke ekonomie word gekenmerk aan 'n klein aantal elemente en 'n beperkte hoeveelheid interaksie. Stelseldinamiese modellering het egter 'n groot aantal elemente met veel meer interaksies tussen hierdie elemente. Die natuur-vryheid grondmotief is onderworpe aan 'n aantal punte van kritiek, insluitende die feit dat dit dualistiese denke in analise inbring. Verder spreek dit ook nie voldoende die normatiewe of morele kwessies aan nie. Die raamwerk van Dooyeweerd, wat dan die derde raamwerk is, word voorgestel as 'n wyse waarop die natuur-vryheid grond-motief getransendeer kan word. Alhoewel die natuur-vryheid grondmotief grootliks gebruik word in hierdie studie, transendeer die analise die tradisionele ekonomiese benadering op 'n aantal gebiede. Hierdie gebiede sluit die volgende in: 'n fokus op transdissiplinere metodes, onewewigtigheid, 'n gevallestudie benadering, en empiriese skatting in plaas van teoretiese modelle. Die restorasie gevallestudies wat in hierdie studie gebruik word is voorbeelde van individuele komplekse sisteme. Agt dinamiese stelsel modelle word ontwikkel om die interaksies tussen ekonomiese, ekologiese en hidrologiese komponente in elke gevallestudie te modelleer. Hierdie agt stelseldinamiese modelle word dan gebruik in 'n risiko analise proses wat uitloop op 'n portefeulje plot oefening. Hierdie portefeulje plot oefening word dan gebruik om eienskappe en kenmerke van verskeie gevallestudie terreine te identifiseer gebaseer op die risiko profiel van elke terrein. Hierdie studie is die eerste bekende toepassing van dinamiese stesels, risiko analise en portefeulje plot tot 'n omgewingsrestorasie projek. Hierdie raamwerk kan potensieël gebruik word deur beleidskrywers wat met begrotings beperkinge gekonfronteer word om tussen restorasie projekte te kies en om hulle te prioritiriseer.