京都大学 / 0048 / 新制・課程博士 / 博士(地球環境学) / 甲第21978号 / 地環博第185号 / 新制||地環||37(附属図書館) / 京都大学大学院地球環境学舎地球環境学専攻 / (主査)准教授 真常 仁志, 教授 舟川 晋也, 教授 西前 出 / 学位規則第4条第1項該当 / Doctor of Global Environmental Studies / Kyoto University / DGAM

This study describes the contemporary educational methods of Mongolian nomads from chosen sample locations of the Hangai mountains. It has been completed on the basis of three field research trips conducted in these locations. Recordings of guided interviews and notes based upon involved observation serve as the source material for this study. Mongolian culture is at a turning point, as it is influenced by many elements of the world of Western civilization as well as by modern technologies; these latter are becoming a part of the nomad's daily life to an ever greater extent. These elements are revealed through the comparison of past and contemporary educational methods, children's games and the recollections of elder nomads. It also reveals itself in the comparison of past and contemporary educational institutions. Nevertheless, traditional culture and certain aspects of ethno pedagogy still remain very much alive. It is largely the elders who regard these traditions with much esteem and try to pass them down to younger generations.

京都大学 / 0048 / 新制・課程博士 / 博士(社会健康医学) / 甲第20623号 / 社医博第81号 / 社新制||医||9(附属図書館) / 京都大学大学院医学研究科社会健康医学系専攻 / (主査)教授 木原 正博, 教授 中川 一路, 教授 平家 俊男 / 学位規則第4条第1項該当 / Doctor of Public Health / Kyoto University / DFAM

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The purpose of this study was to record a set of Mongolian monosyllabic word lists that could be used to obtain a word recognition score. A word list was developed of 190 frequently used monosyllabic words which had been selected by a group of raters. Native male and female Mongolian talkers were utilized to make digital recordings. The 190 words were divided into 10 lists of 19 words. The lists were presented to 20 normally hearing subjects at 10 different intensity levels ranging from -5 to 40 dB HL in 5 dB increments. An S-curve distribution was used to divide the words among three lists based upon the results of the word recognition testing. Word lists were assembled formulating 3 lists of 50 words and 6 half-lists of 25 words using an S-curve distribution. Logistic regression was used to calculate the psychometric functions for each list. The mean psychometric function slopes for the male and female word lists were 6.19%/dB and 5.17%/dB respectively. The 50% threshold was 14.47 dB HL for the final adjusted male and female lists.

Since the early 1900s, efforts have been made to catalogue the stoneflies of Mongolia. Taxonomic work from 1960 to 1980 greatly expanded basic lists of stoneflies in Mongolia, but no comprehensive survey or synthesis of this dispersed literature has been completed. In conjunction with a modern survey of the aquatic insects of Mongolia, we collected Plecoptera on a series of expeditions to the Selenge (north) and Altai (west) regions of Mongolia. A total of 48 species distributed in 24 genera and 8 families were documented, including 3 of the 5 Mongolian endemics, 2 new species records for Mongolia, and 1 species new to science. The majority of the fauna are representative of the East Palearctic region. The 800+ species records were used to validate historical species lists, document species ranges with geo-referenced localities, and create various identification tools to be used by researchers and Mongolian natives with a broad range of taxonomic experience. These identification tools include a records database, genera-level key to nymphs, species diagnosis pages, species range and predicted Ecological Niche Modeling (ENM) maps, and a custom "Level of Expectedness" (LOE) report useful for determining an unbiased projected species list for local areas. These tools are primarily intended for use by Mongolian scientists, sampling teams, and community water quality monitoring groups, as well as general use by researchers interested in biogeography, ecology, and water quality applications of Mongolian Plecoptera. With this work, we hope to equip Mongolians with the scientific resources to protect their valuable and vulnerable water resources.

The purpose of this paper is to study and evaluate the Japaenese program relative to the Mongol question. Some questions to consider would be: What relationship was there between the Hsingan provinces and the Manchukuo government and why? What specific Mongol programs did the Japanese promote and why? What were the results? How did the Mongols react? What designs did the Japanese have in Outer Mongolia and Inner Mongolia relative to her key position in Eastern Mongolia? Why and how did China and Russia react to Japan's plans and what were the results? How did the Mongol react to the three Asiatic Powers at this period? In an overall view, what were the successes and failures of the Japanese in their attempt to solve the Mongol question?

The present thesis elucidates facts about a prominent steppe plant’s evolutionary history (i.e., geographic origin, migration route, distribution of genealogical lineages and polyploidization events) and contemporary population divergence (i.e., genetic diversity and differentiation, impacts of abiotic and biotic factors). Artemisia frigida has been chosen as the target species, with Mongolia as the focus study region. Because A. frigida is widely distributed in the both New and Old Worlds, it was a suitable candidate for the phylogeographic study. Moreover, because of its dominance in many communities in Mongolian steppes and tolerance for cold, drought and mechanical disturbances (grazing), evaluating the effect of environmental factors and grazing pressures on its population genetics was profitable. The overall goal of this thesis was to assess the effects of paleo- and current climate, and land use changes on the distribution of A. frigida’s genealogical lineages and genetic variations. The thesis is divided into two main parts: (i) Chapter 3 focuses on Phylogeography. Within this, section 3.3 depicts a study on the phylogeography of A. frigida, covering samples from its distributional range across the northern hemisphere. The study resulted in Asia being the species’ main origination and diversification center, and the species spread northwards to the Russian Far East and eventually crossed the Bering Strait to North America. Among four geographical regions sampled, seven genetic lineages were found, with Middle Asia having the most diverse populations. According to our phylogenetic analysis, two populations of Kazakhstan in Middle Asia represented the most likely ancestral diploids, and subsequent polyploidization events have occurred on several occasions independently. The observed phylogeographic patterns of the species showed that paleoclimate, especially glaciation events of the Quaternary has predominantly affected species’ current distribution, along with the expansion and contraction of the Eurasian steppe. The second part (Chapter 4) is dedicated to Population genetics to reveal the effects of the current climate and land use on population genetic variation. Three studies were conducted at local and regional levels, focusing on Mongolia. The first study (Section 4.1- review of local literature) was done to offer background information about Mongolian steppes, and the effects of climate and grazing on the steppe vegetation. As a result, steppe vegetation responded to grazing in different ways, depending on the interplay of local environmental factors. In particular, an overall negative effect of grazing was found in desert, dry and high mountain steppes, but no or even positive effects in meadow and mountain steppes. The study highlighted the importance of the interaction effect of local environmental conditions and grazing in Mongolian steppe vegetation. The second study (Section 4.2) employed large scale climatic gradient and local scale grazing gradients to assess the effects of grazing and environmental factors on the population genetics of A. frigida. Precipitation gradient covered 110 – 300mm difference of mean annual precipitation from central to southern Mongolia. While three levels of grazing gradient, such as heavy, moderate, and least grazed sites were examined. According to the study, grazing in overall, had no substantial effect on the genetic diversity of A. frigida, while environmental factors, i.e., summer precipitation and soil phosphorous content, promoted high genetic diversity. Genetic differentiation among populations across large climatic gradients was extremely low, suggesting the existence of considerable gene flow among populations across the steppes of Mongolia. The third study (Section 4.3) employed grazing exclosures to evaluate the genuine effects of grazing. Because Mongolia has a long-term nomadic pastoralism history, and grazing of large herbivores is already an integral part of the steppe vegetation. Thus, we utilized reference site fences along the Trans-Mongolian Railway (TMR), where fences have been built and maintained since 1955, resulting in over 60 years of grazing exclusion. In addition, we supplemented this with data from Hustai National Park (HNP), where three fences were established in 2003. As a result, we found a significant positive impact of grazing on the genetic diversity of A. frigida, implying that a certain level of grazing is beneficial for the species. While no grazing effect on the population genetic differentiation was detected, but climatic and soil variables strongly influenced population genetic structure. In summary, this thesis provided an in-depth investigation of the phylogeography and population genetics of the species A. frigida, which can stand as an exemplar for other Eurasian steppe species. Paleoclimate had largely shaped the current distribution pattern of the species, while contemporary climate and environmental heterogeneity promoted species’ polyploidization and genetic variation. Grazing by large herbivores showed no detrimental effect, or even a positive impact on the genetic diversity of A. frigida. Artemisia frigida populations in Mongolia are thus apparently genetically ‘healthy’, in spite of pervasive grazing in the region. Climate variables and environmental heterogeneity had a substantial impact on the species’ both genetic diversity and differentiation, indicating its higher sensibility to climate change than to land use change. The findings of the thesis could be valuable in understanding species genetic variation under global land use and climate changes.:Summary 4 List of Abbreviations 6 Chapter 1. Introduction 8 Chapter 2. Material and methods 10 2.1. Study region: Mongolian steppe 10 2.2. Focus species: Artemisia frigida Willd. 11 2.3. Molecular markers 12 Chapter 3. Phylogeography 13 3.1. Eurasian steppe and its evolutionary history 13 3.2. Artemisia L. (Asteraceae) as model plant for phylogeography 16 3.3. A case study: Phylogeography of Artemisia frigida Willd. 19 Chapter 4. Plant population genetics under changing climate and grazing……. 40 4.1. Climate – grazing interactions in Mongolian steppe 41 4.2. Climate – grazing interactions on plant population genetics 59 4.3. Effect of grazing exclusion on plant populations genetics 83 Chapter 5. Overarching synthesis and discussion 101 5.1. Molecular markers: pros and cons 102 5.2. A review of phylogeographical studies on Eurasian steppe plants 104 5.3. A review on plant population genetic studies in Mongolia 109 5.4. Outlook 113 Acknowledgements 114 References 115 Curriculum vitae 137 Confirmation 143