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Magistro darbe tiriama Nemuno kilpų regioninio parko Punios šilo sumedėjusią augaliją. Darbo objektas – Punios šilas bei jame esantys gamtinis rezervatas ir botaninis – zoologinis draustinis. Dabar Punios šilas – viena svarbiausių Nemuno kilpų regioninio parko dalių. Darbo tikslas – ekologinis Nemuno kilpų regioninio parko Punios šilo sumedėjusios augalijos vertinimas. Darbo metodika – Išanalizavus literatūrinę medžiagą ir remiantis Nemuno kilpų regioniniame parke esančiame Punios šile atliktais sumedėjusios augalijos tyrimais, stebėjimais nustatytas medžių rūšių išsidėstymas, jų pasiskirstymas kvartaluose. Buvo nustatyta medynų ekologinė būklė. Tyrimai buvo vykdomi visuose 48 Punios šilo kvartaluose. Kiekvienam kvartalui sudaryta atskira lentelė, iš kurios atsispindi bendras užimamas kvartalo plotas hektarais, sklypų skaičius, medynų sudėtis, būdinga augavietė, medynų amžiaus vidurkis, skalsumas, medynų tūris. Darbo rezultatai – nustatyta esama sumedėjusios augalijos rūšinė sudėtis, medynų įvairovė, ekologinė būklė. Punios šilo gamtinių objektų būklė yra gera. Pagal medynų charakteristiką Punios šile pušynai užima didžiausią plotą, eglynai per puse mažesnį, o ąžuolynai ir beržynai beveik vienodą plotą. / Post–graduate work examined the Nemunas loops regional park the forest of Punia woody vegetation. Object of the work - forest of Punia with nature reserve and botanical - zoological preserve that are in it. Now forest of Punia - one of the most important part of the Nemunas Loops Regional Park. Aim of the work - the ecological assessment of Nemunas loops regional park forest of Punia woody vegetation. Method of the work - analysis of literary material and on the basis of the Nemunas Loops Regional Park located in forest of Punia for the woody vegetation studies, the observations set out in the distribution of tree species, their distribution areas. It was found the ecological status of the woods. Studies were conducted in all 48 blocks of forest of Punia. For different block was made separate table, which is reflected in the total area in hectares occupied by block, parcel number, stand composition, characterized place of growth, average age of stands, the volume of stands. Results of the work - the existing woody vegetation species composition, stand diversity, the ecological status was set. Forest of Punia natural objects are in good condition. Under the stands testimonial pine forests occupy the largest area of the forest of Punia, fir - a half lower, and the oak woods, and birch forest takes near the same area.

A diversidade beta pode ser definida como a mudança na composição de espécies entre locais em uma determinada área geográfica. Quantificar a contribuição relativa dos diferentes fatores que a afetam é essencial para entender como é mantida a diversidade das comunidades. Nosso estudo tem foco na vegetação lenhosa da região do Alto Camaquã, sul do Brasil (ca.30-31° S, 52-54° W), que está inserida no bioma Pampa e é caracterizada por ampla heterogeneidade ambiental com mosaicos de floresta e campo naturais. Com o objetivo de responder quais fatores tem mais influência na estruturação da comunidade de plantas lenhosas dessa região, distribuímos sistematicamente 60 unidades amostrais onde foram levantadas todas as espécies de plantas lenhosas. Primeiramente particionamos a diversidade beta através de RDA parcial e verificamos que as variáveis ambientais – que incluem variáveis climáticas, topográficas e edáficas – explicaram 28,4% da variação na composição de espécies, a distância geográfica explicou 16,6%, 14,7% foi compartilhado entre os dois componentes e 40,3% permaneceu não explicado. Com isso ficou claro que fatores determinísticos são mais importantes na estruturação das comunidades lenhosas. Em uma posterior análise de árvore de regressão multivariada, as variáveis climáticas foram selecionadas como as mais influentes. Além disso, a região de estudo é uma das mais bem conservadas do Estado. Portanto, em um segundo momento, investigamos a influência de fatores adicionais na diversidade beta das comunidades lenhosas, considerando o histórico de 300 anos de presença de manejo com pecuária familiar do Alto Camaquã. A partir de dados de um zoneamento agroecológico, exploramos a influência das diferentes tipologias de pecuária sobre dois estratos das comunidades de plantas lenhosas, plântulas e adultos, controlando o efeito da variação climática através de correlações matriciais. Encontramos que o manejo não está relacionado com o turnover de espécies de plântulas, contudo explica 12% da variação da diversidade beta de arbóreas. Concluímos, portanto, que a heterogeneidade climática da região gera heterogeneidade ambiental, sendo esta a principal determinante da diversidade das comunidades, apesar de processos neutros também influenciarem, em menor proporção. A presença do manejo não afeta de forma equitativa a comunidade de plantas lenhosas, contribuindo para a heterogeneidade florística da região. / Beta diversity can be defined as the shift in species composition among sites in a geographical area of interest. Quantifying the relative contributions of different processes that affect beta diversity is essential for understanding how diversity is maintained in communities. Our study focuses on the woody vegetation of the Alto Camaquã region, southernmost Brazil (c. 30-31° S, 52-54° W), which is within the domain of the Pampa biome and presents wide environmental heterogeneity showing natural forest-grassland patches. Aiming to answer which factors are most influential in the structuring of the woody plant communities, we systematically placed 60 sampling plots throughout the study area for vegetation survey. We partitioned beta diversity through partial RDA and verified that the environmental variables - which include climatic, topographic and edaphic variables - explained 28.4% of the variation in species composition, geographic distance accounted for 16.6%, 14.7% was shared between the two components and 40.3% of the variation remained unexplained. The deterministic processes are clearly the most important in structuring the woody communities. Further analysis using multivariate regression tree selected the climatic variables as the most influential. The study region shows a well conserved physiognomy, regardless of its 300-years history of land use for family cattleraising; hence in a second moment we investigated additional factors affecting diversity patterns of the plant communities surveyed, starting from available agroecological zoning data for land management. We explored the effect of the different typologies of family cattle-raising over two strata of the woody vegetation community, seedling and adult plants, controlling the effect of climate heterogeneity using matrix correlations (partial Mantel tests). We found that land management is not correlated with the species turnover of seedlings, yet explains 12% of the variation in the adult plants beta diversity. We concluded, hence, that climatic heterogeneity creates habitat heterogeneity, being the main determinant of community diversity, although neutral processes are also influent. Land management does not affect the woody plant community evenly, thus contributing to the floristic heterogeneity of the region.

A diversidade beta pode ser definida como a mudança na composição de espécies entre locais em uma determinada área geográfica. Quantificar a contribuição relativa dos diferentes fatores que a afetam é essencial para entender como é mantida a diversidade das comunidades. Nosso estudo tem foco na vegetação lenhosa da região do Alto Camaquã, sul do Brasil (ca.30-31° S, 52-54° W), que está inserida no bioma Pampa e é caracterizada por ampla heterogeneidade ambiental com mosaicos de floresta e campo naturais. Com o objetivo de responder quais fatores tem mais influência na estruturação da comunidade de plantas lenhosas dessa região, distribuímos sistematicamente 60 unidades amostrais onde foram levantadas todas as espécies de plantas lenhosas. Primeiramente particionamos a diversidade beta através de RDA parcial e verificamos que as variáveis ambientais – que incluem variáveis climáticas, topográficas e edáficas – explicaram 28,4% da variação na composição de espécies, a distância geográfica explicou 16,6%, 14,7% foi compartilhado entre os dois componentes e 40,3% permaneceu não explicado. Com isso ficou claro que fatores determinísticos são mais importantes na estruturação das comunidades lenhosas. Em uma posterior análise de árvore de regressão multivariada, as variáveis climáticas foram selecionadas como as mais influentes. Além disso, a região de estudo é uma das mais bem conservadas do Estado. Portanto, em um segundo momento, investigamos a influência de fatores adicionais na diversidade beta das comunidades lenhosas, considerando o histórico de 300 anos de presença de manejo com pecuária familiar do Alto Camaquã. A partir de dados de um zoneamento agroecológico, exploramos a influência das diferentes tipologias de pecuária sobre dois estratos das comunidades de plantas lenhosas, plântulas e adultos, controlando o efeito da variação climática através de correlações matriciais. Encontramos que o manejo não está relacionado com o turnover de espécies de plântulas, contudo explica 12% da variação da diversidade beta de arbóreas. Concluímos, portanto, que a heterogeneidade climática da região gera heterogeneidade ambiental, sendo esta a principal determinante da diversidade das comunidades, apesar de processos neutros também influenciarem, em menor proporção. A presença do manejo não afeta de forma equitativa a comunidade de plantas lenhosas, contribuindo para a heterogeneidade florística da região. / Beta diversity can be defined as the shift in species composition among sites in a geographical area of interest. Quantifying the relative contributions of different processes that affect beta diversity is essential for understanding how diversity is maintained in communities. Our study focuses on the woody vegetation of the Alto Camaquã region, southernmost Brazil (c. 30-31° S, 52-54° W), which is within the domain of the Pampa biome and presents wide environmental heterogeneity showing natural forest-grassland patches. Aiming to answer which factors are most influential in the structuring of the woody plant communities, we systematically placed 60 sampling plots throughout the study area for vegetation survey. We partitioned beta diversity through partial RDA and verified that the environmental variables - which include climatic, topographic and edaphic variables - explained 28.4% of the variation in species composition, geographic distance accounted for 16.6%, 14.7% was shared between the two components and 40.3% of the variation remained unexplained. The deterministic processes are clearly the most important in structuring the woody communities. Further analysis using multivariate regression tree selected the climatic variables as the most influential. The study region shows a well conserved physiognomy, regardless of its 300-years history of land use for family cattleraising; hence in a second moment we investigated additional factors affecting diversity patterns of the plant communities surveyed, starting from available agroecological zoning data for land management. We explored the effect of the different typologies of family cattle-raising over two strata of the woody vegetation community, seedling and adult plants, controlling the effect of climate heterogeneity using matrix correlations (partial Mantel tests). We found that land management is not correlated with the species turnover of seedlings, yet explains 12% of the variation in the adult plants beta diversity. We concluded, hence, that climatic heterogeneity creates habitat heterogeneity, being the main determinant of community diversity, although neutral processes are also influent. Land management does not affect the woody plant community evenly, thus contributing to the floristic heterogeneity of the region.

A diversidade beta pode ser definida como a mudança na composição de espécies entre locais em uma determinada área geográfica. Quantificar a contribuição relativa dos diferentes fatores que a afetam é essencial para entender como é mantida a diversidade das comunidades. Nosso estudo tem foco na vegetação lenhosa da região do Alto Camaquã, sul do Brasil (ca.30-31° S, 52-54° W), que está inserida no bioma Pampa e é caracterizada por ampla heterogeneidade ambiental com mosaicos de floresta e campo naturais. Com o objetivo de responder quais fatores tem mais influência na estruturação da comunidade de plantas lenhosas dessa região, distribuímos sistematicamente 60 unidades amostrais onde foram levantadas todas as espécies de plantas lenhosas. Primeiramente particionamos a diversidade beta através de RDA parcial e verificamos que as variáveis ambientais – que incluem variáveis climáticas, topográficas e edáficas – explicaram 28,4% da variação na composição de espécies, a distância geográfica explicou 16,6%, 14,7% foi compartilhado entre os dois componentes e 40,3% permaneceu não explicado. Com isso ficou claro que fatores determinísticos são mais importantes na estruturação das comunidades lenhosas. Em uma posterior análise de árvore de regressão multivariada, as variáveis climáticas foram selecionadas como as mais influentes. Além disso, a região de estudo é uma das mais bem conservadas do Estado. Portanto, em um segundo momento, investigamos a influência de fatores adicionais na diversidade beta das comunidades lenhosas, considerando o histórico de 300 anos de presença de manejo com pecuária familiar do Alto Camaquã. A partir de dados de um zoneamento agroecológico, exploramos a influência das diferentes tipologias de pecuária sobre dois estratos das comunidades de plantas lenhosas, plântulas e adultos, controlando o efeito da variação climática através de correlações matriciais. Encontramos que o manejo não está relacionado com o turnover de espécies de plântulas, contudo explica 12% da variação da diversidade beta de arbóreas. Concluímos, portanto, que a heterogeneidade climática da região gera heterogeneidade ambiental, sendo esta a principal determinante da diversidade das comunidades, apesar de processos neutros também influenciarem, em menor proporção. A presença do manejo não afeta de forma equitativa a comunidade de plantas lenhosas, contribuindo para a heterogeneidade florística da região. / Beta diversity can be defined as the shift in species composition among sites in a geographical area of interest. Quantifying the relative contributions of different processes that affect beta diversity is essential for understanding how diversity is maintained in communities. Our study focuses on the woody vegetation of the Alto Camaquã region, southernmost Brazil (c. 30-31° S, 52-54° W), which is within the domain of the Pampa biome and presents wide environmental heterogeneity showing natural forest-grassland patches. Aiming to answer which factors are most influential in the structuring of the woody plant communities, we systematically placed 60 sampling plots throughout the study area for vegetation survey. We partitioned beta diversity through partial RDA and verified that the environmental variables - which include climatic, topographic and edaphic variables - explained 28.4% of the variation in species composition, geographic distance accounted for 16.6%, 14.7% was shared between the two components and 40.3% of the variation remained unexplained. The deterministic processes are clearly the most important in structuring the woody communities. Further analysis using multivariate regression tree selected the climatic variables as the most influential. The study region shows a well conserved physiognomy, regardless of its 300-years history of land use for family cattleraising; hence in a second moment we investigated additional factors affecting diversity patterns of the plant communities surveyed, starting from available agroecological zoning data for land management. We explored the effect of the different typologies of family cattle-raising over two strata of the woody vegetation community, seedling and adult plants, controlling the effect of climate heterogeneity using matrix correlations (partial Mantel tests). We found that land management is not correlated with the species turnover of seedlings, yet explains 12% of the variation in the adult plants beta diversity. We concluded, hence, that climatic heterogeneity creates habitat heterogeneity, being the main determinant of community diversity, although neutral processes are also influent. Land management does not affect the woody plant community evenly, thus contributing to the floristic heterogeneity of the region.

L’évaluation de la performance des ouvrages hydrauliques en remblai est un enjeu majeur pour les gestionnaires en charge de leur sécurité. Qu'il s'agisse de digues ou de barrages, la présence de végétation arborescente représente un facteur de fragilité important. Deux types de mécanismes de détérioration peuvent ainsi être initiés ou aggravés : l’érosion interne et l’érosion externe. Or, ce facteur n’a été pris en compte que partiellement dans les modèles d’évaluation existants. De plus, les petits barrages en remblai peuvent avoir des dispositifs d’étanchéité différents et ne sont pas toujours équipés de drains et d’instruments d’auscultation: il est alors nécessaire d’adapter les modèles à chaque cas. L’objectif de la thèse est de répondre à ces verrous scientifiques. Les résultats de nos recherches concernent l’établissement de modèles d’évaluation de la performance basés sur des approches à base de connaissances.La démarche a été appliquée sur des ouvrages réels : 3 tronçons de digues de protection contre les inondations ainsi que 7 barrages en remblai ce qui nous a permis de montrer sa pertinence et donc son potentiel d’utilisation opérationnelle. / The performance assessment of embankment hydraulic works is a major challenge for managers in charge of their security. For both dikes and dams, the presence of trees represents an important fragility factor. Two types of deterioration mechanism can thus be initiated or aggravated: internal erosion or scouring. However, this factor has been taken only partially into account in existing assessment models. In addition, small embankment dams can have different sealing devices and are not always equipped with drains and monitoring instruments: it is thus necessary to customize the models in each case. The aim of the thesis was to remove these scientific obstacles. The results of our research concern the design of performance assessment models relying on knowledge-based approaches.The approach was implemented for real works: three sections of river dikes and seven small embankment dams which allowed us to show its relevance and therefore its operational potential.

Student Number : 9008538J - MSc Dissertation - School of Biology - Faculty of Science / O’Regan SP, 2005. The long-term effects of fire frequency and season on the woody vegetation in the Pretoriuskop sourveld of the Kruger National Park. MSc Dissertation, University of the Witwatersrand, Johannesburg. The role of fire in the management of conservation areas has historically been a contentious issue in which traditional agricultural principles and ever-changing conservation principles tend to collide. The Kruger National Park (KNP) in the early 1950s was no exception where the appropriate use of fire and its ecosystem consequences were hotly debated. The controversy surrounding the management of fire in the KNP highlighted the significant lack of understanding of fire and its role in the ecosystem and because of this controversy, the Experimental Burn Plot (EBP) experiment was established in 1954. The EBP experiment comprised 12 treatments, and a pseudo-randomised block design was used in which the 12 fire treatments were replicated four times each in four of the six major vegetation zones identified at the time. The EBP experiment originally comprised 192 experimental plots approximately 7 Ha in size each and covered approximately 12 km2 in the KNP. The twelve fire treatments were an annual burn in August, biennial and triennial burns in February, April, August, October, and December, and a control on which fire was excluded. Despite having been plagued with negative assessments from internal and external researchers from its inception, the EBP experiment was meticulously maintained, and it has now become a valuable research asset in the KNP. Four replicates of twelve plots each were located in the Pretoriuskop sourveld landscape of the KNP. These replicates were named Fayi, Kambeni, Numbi, and Shabeni after nearby landmarks. The Pretoriuskop region is a moist infertile mesic-savanna, which experiences on average 744mm of rain annually. The dominant tree species in Pretoriuskop are Dichrostachys cinerea and Terminalia sericea and the dominant grass species is Hyperthelia dissoluta. A baseline survey of the woody vegetation was done on all the Pretoriuskop plots in 1954 by HP Van Der Schijff. A second survey of the woody vegetation on all the Pretoriuskop plots was done in 1996 by SP O’Regan. This provided a 42-year period of treatment application over which the effects of fire frequency and season on the woody vegetation of the Pretoriuskop region were studied. The aim of this study was to investigate the long-term effects of the twelve fire treatments on the density, structure, and species composition of the woody vegetation in Pretoriuskop. The objectives of this study were: 1. To carry out a complete re-survey of the trees and shrubs on the Pretoriuskop EBPs using similar methods as those used in the baseline survey in 1954. 2. To capture into a digital format pertinent woody vegetation survey data from surveys that had been conducted on the Pretoriuskop EBPs between 1954 and 1996. 3. To compare the density, structure, and composition of the woody vegetation on the Pretoriuskop EBPs between 1954 and 1996, to determine the effects of fire on the woody vegetation of Pretoriuskop. 4. To investigate the history of the Kruger National Park Experimental Burn Plots experiment. The four replicates in the Pretoriuskop region were found generally to have very similar woody vegetation traits (density, species composition, and structural composition). However, the EBPs were established and surveyed in two distinct phases, the first phase comprised the control, August Annual, and the Biennial plots, and the second phase comprised the Triennial plots. The baseline structural composition of the plots established in the first phase was different from the structural composition of the plots in the second phase. Furthermore, the Pretoriuskop EBPs are located in two distinct vegetation types, namely the open and the closed Terminalia sericea \ Combretum woodlands of the Pretoriuskop region. The Numbi and Shabeni replicates are in the open Terminalia sericea \ Combretum woodlands, and the Kambeni and Fayi replicates are in the closed Terminalia sericea \ Combretum woodlands. It was found that the species composition of the plots was influenced by the location of the plots in the different vegetation types. The exclusion of fire in the Pretoriuskop sourveld results in an increase in the density of the overstorey and understorey woody vegetation, and an increase in the number of species, species diversity, and species evenness. This is because fire sensitive and fire intolerant woody species become more abundant as the period between fires increases. In Pretoriuskop, there is no evidence of relay floristic succession, because fire sensitive and fire intolerant woody species do not replace fire tolerant species. Instead, the floristic succession is accumulative and fire tolerant, fire sensitive, and fire intolerant woody species coexist as the period between fires increases. Woody species tolerant of frequent fires in Pretoriuskop are Albizia versicolor, Catunaregam spinosa, Lonchocarpus capassa, Pavetta schumanniana, Senna petersiana, Strychnos madagascariensis, and Turraea nilotica. Woody species that are sensitive or intolerant of fire in Pretoriuskop are Acacia swazica, Bauhinia galpinii, Combretum mossambicense, Commiphora neglecta, Croton gratissimus, Dalbergia melanoxylon, Diospyros lycioides, Diospyros whyteana, Euclea natalensis, Hyperacanthus amoenus, Kraussia floribunda, Ochna natalitia, Olea europaea, Psydrax locuples, Putterlickia pyracantha, Tarenna supra-axillaris, and Zanthoxylum capense. Dichrostachys cinerea and Terminalia sericea were found to dominate in areas that had been burnt frequently as well as areas where fire has been excluded. The change in the density of the woody vegetation as the inter-fire period increases is not linear but rather J shaped with an initial decrease in the density as the inter-fire period increases from 1 year to 3 years. This initial decrease in density is the result of a loss of very short (<1m tall) woody individuals. In contrast, there is no initial decrease in the number of tree equivalents (phytomass) of the woody vegetation as the inter-fire period increases. After the initial decrease in the density of the woody vegetation, the density increases as the inter-fire period increases beyond 3 years. Generally in Pretoriuskop, post fire age of the vegetation was found to be an important factor affecting the structure of the woody vegetation, and as the inter-fire period increases the number of structural groups, the structural diversity, and the structural evenness of the woody vegetation increases. As the inter-fire period increases the number of single-stem individuals relative to the number of multi-stem individuals increases, and the average height of the woody vegetation increases. The findings regarding the effects of fire frequency on the Pretoriuskop EBPs were similar to the findings on other fire experiments in mesic African savannas. The finding on the Pretoriuskop EBPs differed from the findings in other fire trials that were in arid savannas in Africa. Generally, the exclusion of fire in moist savannas (> 600 mm of rain annually) results in the woody vegetation becoming denser, while the exclusion of fire in arid to semi-arid savannas (< 600mm of rain annually) does not result in the woody vegetation becoming denser. In Pretoriuskop, fires occurring in summer between December and February have a different impact on the density, species composition, and structure of the woody vegetation than fires occurring in winter between August and October. Furthermore, fires occurring in April have a different impact on the density, species composition, and structure of the woody vegetation in Pretoriuskop. Woody vegetation burnt by summer fires is denser than woody vegetation burnt by winter fires. The number of species and species diversity of the woody vegetation is also higher in vegetation burnt by summer fires in comparison with vegetation burnt by winter fires. The density and species composition of woody vegetation in areas that have been burnt in summer fires is more similar to areas where fire has been excluded than to areas that have been burnt in winter fires. The woody species associated with vegetation burnt in summer fires and where fire has been excluded are Euclea natalensis, Antidesma venosum, Diospyros lycioides, Phyllanthus reticulatus, Grewia flavescens, Grewia monticola, Ochna natalitia, Peltophorum africanum, Rhus pyroides, Diospyros mespiliformis, Rhus transvaalensis, Securinega virosa, Putterlickia pyracantha, Rhus pentheri, Commiphora neglecta, Heteropyxis natalensis, and Olea europaea. Structurally the average height of the woody vegetation is taller in areas burnt by winter fires than in areas burnt by summer fires. The woody vegetation in areas burnt in summer fires have more single-stem individuals relative to multi-stem individuals than in areas burnt in winter fires. The structural composition of areas burnt in summer fires is more similar to areas where fire has been excluded than with areas burnt in winter fires. The structure of the woody vegetation in areas burnt in winter fires is generally dominated by multi-stem individuals that are 0-1m tall or 3-5m tall. The structure of the woody vegetation in areas burnt in summer fires or where fire has been excluded is dominated by both single-stem and multi-stem individuals of all heights and basal diameters. Findings regarding the effect of early dry season fires (April) in comparison with late dry season fire (August) on the woody vegetation are consistent with the findings on other fire trails in Africa. However, a comparison of all the fire-timing treatments between the Pretoriuskop and Satara EBPs in the KNP reveals that the timing of fires affects the woody vegetation differently in different areas even when the affects at certain times appear similar. The data collected on the Pretoriuskop EBPs reveals that there have been significant changes in the woody vegetation in Pretoriuskop between 1954 and 1996. The density of the woody vegetation increased between 1954 and 1996 by almost 200%. The number of species and the species diversity of the woody vegetation also increased between 1954 and 1996. In 1954, there were approximately equal numbers of single-stem and multi-stem individuals, while in 1996 there were more multi-stem individuals than single-stem individuals. The increase in atmospheric CO2 levels between 1954 and 1996 is believed to have been a factor that has driven the changes in the woody vegetation of Pretoriuskop between 1954 and 1996.