Each species has specific adaptations to its environment, and since environmental parameters reveal geographic trends, it is reasonable to expect the existence of geographic trends in species characteristics as well. The aim of this study has been to evaluate the effect of environmental conditions on geographic variability of functional traits of European birds. I have analysed the effect of temperature, precipitation, productivity, altitude and habitat type (forest, open habitats, bush, settlements, wetlands) on clutch size, number of clutches per breeding season, egg size, incubation length, age of maturity, body mass, wing, tail, bill and tarsus length. I have used data from the European breeding bird atlas, so that I have calculated mean values of all the traits for quadrats 50x50 km, and then related them to environmental characteristics using OLS and GLS. Clutch size increases with temperature, whereas the number of clutches decreases with it, indicating possible trade-off between clutch size and the number of clutches, whose result is determined by the length of breeding season. Egg size decreases with temperature, possibly due to higher survival of large eggs (and consequently juveniles) in cold regions. Incubation length increases with both temperature and environmental productivity,...

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The human voice may convey meaningful information about socially and evolutionarily relevant characteristics of the vocalizer. In turn, listeners may readily evaluate personal characteristics, such as body size, on the basis of nonverbal voice features. Research investigating vocal communication of physical size in humans has focused on two salient and largely independent voice features, fundamental frequency and/or corresponding harmonics (perceived as voice pitch) and formant frequencies (resonance frequencies of the supralaryngeal vocal tract). However, the degree to which fundamental and formant frequencies reliably predict variation in body size controlling for sex and age, and their relative role in the perception or accurate estimation of body size, has to date been unclear. In the current thesis, using meta-analysis, I establish that formants reliably predict variation in men’s and women’s heights and weights. In contrast, fundamental frequency only weakly predicts men’s heights and women’s weights. These findings corroborate work on many other mammals whose vocal production, like humans, follows the source-filter model. Despite the lack of a robust physical relationship between fundamental frequency and size within sexes, I further demonstrate that listeners utilize voice pitch to accurately gauge men’s relative height. My research suggests that voice pitch indirectly facilitates accurate size assessment by providing a carrier signal (i.e., dense harmonics) for formants. This is the first evidence that pitch does not confound accurate size estimation. Finally, I demonstrate that voices with lowered pitch, but not raised pitch, are perceived as larger when projected from a low than high spatial location. These results suggest that strong cross-modal perceptual biases linking low pitch to low elevation and large size may, in some contexts, cause errors in size estimation. Taken together, this thesis provides a detailed account of human vocal communication of body size, which can play a meaningful role in sexual and social contexts. / Dissertation / Doctor of Philosophy (PhD)

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Fischgemeinschaften in Seen sind zunehmenden Umweltveränderungen unterworfen. Die Analyse des Einflusses abiotischer und biotischer Faktoren auf die Größenstruktur von Fischgemeinschaften ist entscheidend, um die Entwicklung von Seeökosystemen im Zuge von Umweltveränderungen vorhersagen zu können. Es ist jedoch schwierig Fische in Seen zu beproben, da Fanggeräte selektiv sind. Die Qualität und Vergleichbarkeit von Fischfängen durch Angler, Kiemennetze, Schleppnetze und Hydroakustik wurden untersucht. Anglerfänge eignen sich begrenzt für vergleichende Fischbestandsuntersuchungen. Vertikale Hydroakustik eignet sich für die Fischbiomassequantifizierung in geschichteten Seen. Der Beprobungsaufwand von Kiemennetzen kann reduziert werden, wenn zeitnah hydroakustische Untersuchungen statt finden. Schleppnetze eignen sich für die Beprobung von pelagischen Fischen in tiefen Seen. Die Größenstruktur von Fischgemeinschaften unterschied sich auf regionaler und überregionaler Ebene entlang abiotischer und biotischer Gradienten. Die regionale Studie zeigte, dass tiefe nährstoffarme Seen mit hohen Raubfischabundanzen durch mehr große Fische gekennzeichnet waren. Nährstoffreiche Flachseen mit wenigen Raubfischen zeigten höhere Anteile von mittleren Fischgrößen. Im überregionalen Vergleich führten Temperaturpräferenzen der Fischarten zu zwei Fischgemeinschaften, die sich in ihrer Größenzusammensetzung unterschieden. Ein hoher Anteil großer Salmoniden war typisch für hochgelegene Kaltwasserseen im Norden und Süden Europas. Flachlandseen mit Kühl- und Warmwasserfischen zeichneten sich durch eine Dominanz von kleinen Fischen aus. Das lässt vermuten, dass Fischgemeinschaften in Flachlandseen relativ robust gegenüber Umweltveränderungen reagieren. Kaltwasserfischgemeinschaften könnten im Zuge der Klimaerwärmung starken Änderungen unterliegen, da eine Veränderung in der Artenzusammensetzung von einer Verschiebung in der Größenstruktur hin zu mehr kleinen Fischen begleitet sein wird. / Lake fish assemblages are increasingly exposed to environmental change in their habitats. The investigation of abiotic and biotic factors and their effects on the size structure of lake fish is essential for predicting the response of fish assemblages to environmental change. However, sampling of lake fish is challenging because sampling gears are selective. The quality and correspondence of fish catches from angling, gillnetting, trawling and hydroacoustics were analysed. Non-standardised catch data from recreational angling were of limited quality for a comparison of fish stocks. Vertical hydroacoustics is an efficient method to quantify fish biomass in stratified lakes. Sampling effort of multi-mesh gillnets can be reduced when fish abundance estimates are derived from contemporarily operating hydroacoustics. Trawling is useful to sample pelagic fish assemblages in deep lakes. The size structures of fish assemblages differed on a small and a large geographical scale along gradients of abiotic and biotic lake descriptors and differences in fish assemblage composition. At a small geographical scale assemblages in deep and less nutrient-rich lakes with high predator abundances were characterised by a higher proportion of large fish. Shallow nutrient-rich lakes with few predators were characterised by more medium-sized fish. At a large geographical scale thermal optima of fish generated two types of assemblages with different size structures. A high proportion of large salmonids was observed in coldwater lakes at high elevation sites in northern and southern Europe. Lowland lakes with cool- and warmwater fish were dominated by small-sized individuals. The results suggest that lowland lake fish assemblages are relatively robust against environmental change. Coldwater fish assemblages instead may suffer dramatic consequences from global warming as expected species shifts are likely to be accompanied by shifts in the size structure towards smaller individuals.

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The purpose of this study was to assess the epidemiology of preventable risk factors for NCDs among the adult population in Tigray, Northern Ethiopia. A quantitative descriptive cross-sectional design was employed to describe the distribution of behavioural and biological risk factors for NCDs, assess the status of knowledge, perceptions, attitude and behaviour of the study participants for NCDs and their risk factors, and a matched case-control study to identify the determinants of hypertension. The data was collected using a structured questionnaire for the interview, physical measurements including weight and height scales, non-elastic measuring tape for waist and hip circumferences, Omron digital BP apparatus for blood pressure and heart rate; Accutrend Plus for measuring fasting blood glucose, cholesterol and triglycerides. For the descriptive cross-sectional study a total of 2347 participants were included, and for the matched case control study a total of 117 cases and 235 controls participated. Behavioural and biological risk factors were assessed. Only 0.8% of the study participants used optimal fruit serving per day. The prevalence of low level physical activity (<600 MET-minutes/week) was 44.8%. The magnitude of ever alcohol consumption was 66.8%. However, the magnitude of khat chewing and tobacco smoking among the study participants was not as high as the other risk factors i.e. 3.3% and 2.3% respectively. The magnitude of hypertension, central obesity, hyperglycaemia, hypercholesterolemia and hypertriglyceridemia was 9.9%, 22.2%, 3.5%, 30.3% and 32.2% respectively. Factors associated with the risks aforementioned were gender, age, place of residence, education, knowledge status on NCDs, mental stress and others. The status of knowledge on CVDs, breast and cervical cancers, diabetes and their potential risk factors was low and not comprehensive. Misconceptions on NCDs and body size and shape were pervasive. Risky behaviours underlying NCDs were rampant in the study population. Factors related to poor knowledge on NCDs were gender, age, place of residence, education and misconceptions on NCDs. The determinants of hypertension were physical inactivity, duration of alcohol intake, central obesity and mental stress. Awareness raising interventions on NCDs and their risk factors; improving socio-economic status and accessibility to health care settings have to be in place to curb these formidable problems. / Health Studies / D. Litt. et Phil. (Health Studies)

Die Körpergröße ist von großer Bedeutung für die Fitness vieler Tiere, weil sie positiv mit Überleben und Reproduktionserfolg korreliert ist. Große Rogner vieler Fischarten sind fruchtbarer und produzieren Nachkommen von höherer Qualität verglichen mit kleineren Weibchen. Auch für Milchner einiger Fischarten wurde ein Einfluss der Körpergröße auf frühe Lebensstadien nachgewiesen. Der größenabhängige paternale Effekt verglichen mit maternalen Effekten ist weniger gut untersucht. Das Verständnis der Variation im Reproduktionserfolg als Funktion der Körpergröße der Laicher ist wichtig, weil die Fischerei die Durchschnittsgröße des Laicherbestands reduziert. In vorliegender Dissertation wurden in Laborversuchen an Zebrafischen (Danio rerio) größenabhängige paternale und maternale Effekte auf den Reproduktionserfolg und die Auswirkungen größenselektiver Entnahme auf Körperlänge, Reifung und Reproduktionserfolg untersucht. Die Köperlänge und Kondition waren wichtige Determinanten der Reifung bei Zebrafischen. Größere Rogner zeigten höheren Reproduktionserfolg als kleinere Fische und ein signifikanter Einfluss der Milchnerkörperlänge auf die frühen Lebensstadien ihrer Nachkommen wurde dokumentiert. Längere Männchen wurden von Rognern auch bei der Paarung bevorzugt. Die größenabhängigen maternalen und paternalen Effekte waren ausschlaggebend für den erhöhten Reproduktionserfolg von Zebrafischlaichbeständen, die, verglichen mit kleinen Laichern, aus großen oder zufällig zusammengesetzten Individuen zusammengesetzt waren. Die größenselektive Entnahme führte zu phänotypischen und genetischen Veränderungen, die nach Einstellung der experimentellen Befischung persistierten. Das deutet an, dass die durch die Fischerei ausgelöste Evolution schwierig umkehrbar sein könnte. Die Köpergröße ist von überragender Bedeutung in der Reproduktionsbiologie des Zebrafisches und der Schutz großer Laichfische kann wichtig für den Erhalt der Reproduktionskapazität von befischten Beständen sein. / Body size is a fundamentally important trait for fitness in many animal species because it correlates positively with survival and reproductive success. In many fish species, large females exhibit higher fecundity and produce higher quality offspring compared to small females. Similarly, male body size can affect offspring quality and early life-history traits but the importance of these effects to the reproductive biology of fish is poorly studied. The extent to which variation in reproductive success is explained by parental body size is an important research topic because size-selective fishing usually reduces the average size of reproducing adults in a population. In my dissertation, I studied the parental size effects on reproductive success in a model species (zebrafish, Danio rerio). I also studied the effects of size-selective harvesting on body size, maturation and reproductive output. Body size and condition factor were important determinants of the initiation of maturation in zebrafish. Large females were found to have higher reproductive success compared to small females and a significant effect of male body size on early life-history traits was documented. I found that large males were also favored by the females resulting to differential allocation of reproductive resources toward large males. The maternal- and paternal-size effects ultimately led to elevated reproductive success of experimental spawning stocks consisting of large or random-sized individuals compared to spawning stocks consisting of small individuals. Size-selective harvesting induced rapid phenotypic and genetic changes, which persisted after selection was halted. This suggests that fishing-induced changes might be hard to reverse. My results emphasize the importance of body size to the reproductive biology of zebrafish and suggest that protecting large fish might be important to maintain the reproductive potential of exploited fish stocks.

Comprendre comment la biodiversité des prairies répond aux changements d’utilisation des terres constitue un enjeu majeur pour la conservation de la biodiversité et le maintien des services écosystémiques dans les paysages agricoles. Dans ce travail de thèse, nous avons cherché (i) à analyser la réponse simultanée d’un ensemble de taxons appartenant à différents niveaux trophiques (plantes, herbivores, pollinisateurs, prédateurs) aux changements d’utilisation des terres agissant à différentes échelles spatiales et temporelles ; (ii) à approcher les mécanismes impliqués dans cette réponse et notamment le rôle des interactions trophiques. Notre approche est basée sur l’utilisation de multiples traits fonctionnels liés à l’acquisition des ressources, la taille et la mobilité des organismes. Nous avons testé la réponse de ces traits à l’histoire des paysages, leur configuration et leur composition. Nous montrons qu’il existe une réponse générale de la diversité fonctionnelle multitrophique aux changements d’utilisation des terres. Nous mettons en évidence l’importance des effets historiques du changement d’utilisation des terres à l’échelle des paysages agricoles menaçant le maintien de communautés fonctionnellement diversifiées dans ces paysages. En considérant un set de traits multiples, notre travail a permis d’approcher certains mécanismes par lesquels les changements d’utilisation des terres présents et passés impactent différentes facettes de la biodiversité. Enfin, l’utilisation des traits fonctionnels a permis d’appréhender l’importance des interactions trophiques et leur implication dans la structuration des communautés animales dans les milieux agricoles. / Understanding how grassland biodiversity responds to land use intensification is crucial for both biodiversity conservation and the management of key ecosystem services in agricultural landscapes. My PhD aims at (i) identifying and generalising the effects of land use intensification operating at different spatial and temporal scales across multiple taxonomic groups and trophic levels (plants, herbivores, pollinators, predators and top-predators) ; (ii) investigating the underlying mechanisms of biodiversity response, and particularly the role of trophic interactions. We used multiple functional traits related to resource acquisition, the size of the organisms and their mobility. We tested how multitrophic functional trait diversity responded to landscape history, composition and heterogeneity. Considering multiple taxonomic groups simultaneously, our study brings out a clear response of overall biodiversity to land use intensification. We found that legacy effects of land use intensification operating at the landscape scale are major drivers of present-day multitrophic functional trait diversity in agricultural landscapes. By considering a core set of organismal traits reflecting similar functions across trophic levels, our approach reveals multiple dimensions by which land use intensification filters out biodiversity over time and allows us to generalise its effect across multiple trophic levels and trait-spectrum. Finally, trait-based approach allowed us to assess the importance of trophic interactions and their contribution in shaping animal communities in agricultural landscapes.