Thesis (M.S. in civil engineering)--Washington State University, May 2007. / Includes bibliographical references.

Orientador: Adriano Wagner Ballarin / Banca: Mario Tomazello Filho / Banca: Edson Luiz Furtado / Resumo: O látex é o principal produto extraído da seringueira (Hevea brasiliensis). No Brasil, a madeira de seringueira, ao final do ciclo produtivo de látex, é utilizada tradicionalmente para fins energéticos, mas trabalhos internacionais relatam práticas consolidadas de agregação de valor a ela. Diversos pesquisadores têm observado crescimento e agravamento do problema da seca do painel de sangria nos plantios nacionais, doença que compromete a produção de látex. O objetivo principal deste trabalho foi avaliar a qualidade da madeira de árvores dos clones comerciais GT 1 e RRIM 600 em três condições distintas de sanidade: árvores com produção normal de látex (sadias); árvores com presença de seca do painel de sangria por origem abiótica/fisiológica e árvores com presença da seca do painel de sangria por origem biótica/patológica. Foram amostradas seis árvores de cada condição de sanidade dos dois clones de plantios na região de São José do Rio Preto-SP com idade igual ou superior a 20 anos. Os parâmetros anatômicos e físico-mecânicos avaliados foram o comprimento das fibras (presença e delimitação de madeira juvenil e adulta), densidade básica e aparente da madeira, retratibilidade e índice de anisotropia, resistência à compressão paralela às fibras, resistência e rigidez à flexão e dureza Janka. Os resultados mostraram que a madeira juvenil ocorre desde a medula até cerca de 40% e 32% do raio para os clones GT 1 e RRIM 600, respectivamente, podendo-se associar a idade do plantio de 6 a 8 anos como transição entre madeira juvenil e adulta. A densidade básica média da madeira de seringueira dos dois clones... (Resumo completo, clicar acesso eletrônico abaixo) / Abstract: Latex is the main product extracted from rubber tree (Hevea brasiliensis). In Brazil, at the end of the production cycle of latex, rubber wood is traditionally used for energy purposes but international studies have reported consolidated practice that result in added-value products. Several researchers have observed raised and worsening of tapping panel dryness, a disease that impairs the production of latex. The main objective of this study was to evaluate the quality of rubber wood of commercial clones GT 1 and RRIM 600 in three distinct health conditions: trees producing latex (healthy), trees with tapping panel dryness of abiotic/physiological origin and trees with tapping panel dryness of biotic/pathological origin. Six trees of each health condition of the two clones from plantations in the São José do Rio Preto region - Sao Paulo State, Brazil - aged 20 years or more were sampled. Anatomical and physical-mechanical parameters evaluated were fibers length (occurrence and delimitation juvenile-mature wood), specific gravity and apparent density, shrinkage and anisotropy index, compression parallel to grain strength, bending strength and rigidity and Janka hardness. Results showed that juvenile wood occurs from pith up to 40% and 32% of ray for GT 1 and RRIM 600 clones respectively, allowing the association of the age of 6 to 8 years of plantation as transition between juvenile and mature wood. The mean specific gravity of rubber wood from both clones varied from 580 kg/m³ to 595 kg/m³ and there wasn't statistical difference of this parameter among health conditions and between clones. Rubber wood... (Complete abstract click electronic access below) / Mestre

Made available in DSpace on 2014-06-11T19:30:20Z (GMT). No. of bitstreams: 0 Previous issue date: 2011-08-02Bitstream added on 2014-06-13T21:00:54Z : No. of bitstreams: 1 leonello_ec_me_botfca.pdf: 1914995 bytes, checksum: 57b29ba4eed421b275526c5dee3e7bc3 (MD5) / Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP) / O látex é o principal produto extraído da seringueira (Hevea brasiliensis). No Brasil, a madeira de seringueira, ao final do ciclo produtivo de látex, é utilizada tradicionalmente para fins energéticos, mas trabalhos internacionais relatam práticas consolidadas de agregação de valor a ela. Diversos pesquisadores têm observado crescimento e agravamento do problema da seca do painel de sangria nos plantios nacionais, doença que compromete a produção de látex. O objetivo principal deste trabalho foi avaliar a qualidade da madeira de árvores dos clones comerciais GT 1 e RRIM 600 em três condições distintas de sanidade: árvores com produção normal de látex (sadias); árvores com presença de seca do painel de sangria por origem abiótica/fisiológica e árvores com presença da seca do painel de sangria por origem biótica/patológica. Foram amostradas seis árvores de cada condição de sanidade dos dois clones de plantios na região de São José do Rio Preto-SP com idade igual ou superior a 20 anos. Os parâmetros anatômicos e físico-mecânicos avaliados foram o comprimento das fibras (presença e delimitação de madeira juvenil e adulta), densidade básica e aparente da madeira, retratibilidade e índice de anisotropia, resistência à compressão paralela às fibras, resistência e rigidez à flexão e dureza Janka. Os resultados mostraram que a madeira juvenil ocorre desde a medula até cerca de 40% e 32% do raio para os clones GT 1 e RRIM 600, respectivamente, podendo-se associar a idade do plantio de 6 a 8 anos como transição entre madeira juvenil e adulta. A densidade básica média da madeira de seringueira dos dois clones... / Latex is the main product extracted from rubber tree (Hevea brasiliensis). In Brazil, at the end of the production cycle of latex, rubber wood is traditionally used for energy purposes but international studies have reported consolidated practice that result in added-value products. Several researchers have observed raised and worsening of tapping panel dryness, a disease that impairs the production of latex. The main objective of this study was to evaluate the quality of rubber wood of commercial clones GT 1 and RRIM 600 in three distinct health conditions: trees producing latex (healthy), trees with tapping panel dryness of abiotic/physiological origin and trees with tapping panel dryness of biotic/pathological origin. Six trees of each health condition of the two clones from plantations in the São José do Rio Preto region – Sao Paulo State, Brazil - aged 20 years or more were sampled. Anatomical and physical-mechanical parameters evaluated were fibers length (occurrence and delimitation juvenile–mature wood), specific gravity and apparent density, shrinkage and anisotropy index, compression parallel to grain strength, bending strength and rigidity and Janka hardness. Results showed that juvenile wood occurs from pith up to 40% and 32% of ray for GT 1 and RRIM 600 clones respectively, allowing the association of the age of 6 to 8 years of plantation as transition between juvenile and mature wood. The mean specific gravity of rubber wood from both clones varied from 580 kg/m³ to 595 kg/m³ and there wasn’t statistical difference of this parameter among health conditions and between clones. Rubber wood... (Complete abstract click electronic access below)

Diss. (sammanfattning) Uppsala : Sveriges lantbruksuniv. / Härtill 4 uppsatser.

Acetylation of wood is a modification technique that chemically alters the wood substance and enhances several properties of wood. The basic principle is to impregnate wood with acetic anhydride to react and replace OH-groups with acetyl groups in the wood cell wall. In this way, the hygroscopicity of the modified wood is significantly reduced resulting in increased dimensional stability and durability compared with unmodified wood.The objective of this work was to study finger-jointing of acetylated Scots pine (Pinus sylvestris L.) using a conventional melamine urea formaldehyde (MUF) adhesive. Two different types of acetylated pine specimens were investigated, acetylated pine sapwood (APS) and acetylated juvenile pine (AJP), the latter originating from young forest thinning trees (ca 20-30 years). The goal was to evaluate the bending strength, i.e. modulus of rupture (MOR), of such finger-jointed samples, in particular when the acetylated wood was combined with unmodified wood, in this case, Norway spruce (Picea Abies L. Karst) (US). The finger-jointing were performed at Moelven Töreboda by applying their existing industrial procedures. In total, five different of finger jointed sample groups were prepared combining the different specimens: APS-APS, AJP-AJP, US-US, APS-US, and AJP-US. Standardized procedures were used to determine the MOR of the finger-jointed samples, both unexposed at the factory condition state and after a water-soaking-drying cycle. In addition, the experiments also included determination of the moisture content (MC), density, and modulus of elasticity (MOE) (in bending along the grain) of the individual specimens.At the unexposed state, the APS-APS samples showed the highest MOR of 63,1 MPa, while those of the AJP-AJP showed the lowest value of 42,4 MPa. The corresponding values for the US-US, AJP-US and APS-US samples was 56,7, 47,5 and 46,9 MPa, respectively. In contrast to a typical wood failure for the US-US samples, a low amount of wood failure was observed in all cases involving the acetylated wood, indicating a low adhesive anchoring in the wood substrate at the finger-joint, although a surprisingly high strength was obtained for the APS-APS samples. A significantly lower MC content of 4,9 % and a remarkably low value of 1,7 %, was found for the APS and AJP, respectively, compared with 9,2% for the US. The significantly lower MC combined with an assumed increased hydrophobicity of the acetylated wood possible causes a less effective MUF-wood bonding, or adhesion, compared with that of the unmodified wood. Possible, so-called over penetration of the MUF resin in the acetylated wood could also be an explanation for the poor wood-adhesive anchoring. The MOE of the individual APS, AJP and US specimens was 12,6, 8,3 and 11,4 GPa, respectively, indicating a significantly lower mechanical performance of AJP, and hence also of finger-joints of AJP, despite its very low MC, possible due to a higher microfibril angle in the cell walls in juvenile wood compared with mature wood. No clear correlation was found between the MOR and density of the acetylated samples.For the samples exposed to a water-soak-drying cycle, the highest MOR, and lowest reduction of 14 % compared with the unexposed state, was obtained for the US-US samples, whereas all samples involving the acetylated wood showed a distinctly higher reduction. The MOR of the AJP-AJP and AJP-US samples were reduced with 47 % and 50 %, respectively, while the MOR of the APS-APS and APS-US samples were reduced with 43 % and 23 %, respectively. It should be emphasized, however, that after the standard drying-time, which was the same for all samples, the acetylated samples, compared with the untreated ones, did not dry out to the same level as for the dry unexposed state, i.e. the acetylated samples had a high MC of ca 30-40% in these MOR tests. This high MC level could be the main reason for the dramatic strength losses. Furthermore, a less efficient wood-MUF adhesion as well as the drying under acidic conditions may also be possible causes for the reduced bending strength of the finger-jointed samples with acetylated wood.

Submitted by Cássia Santos (cassia.bcufg@gmail.com) on 2017-07-25T12:46:48Z No. of bitstreams: 2 Dissertação - Mariana Dianese Alves de Moraes - 2017.pdf: 2047536 bytes, checksum: 5d64fdcced984e8d8b288a0799a3e6e0 (MD5) license_rdf: 0 bytes, checksum: d41d8cd98f00b204e9800998ecf8427e (MD5) / Approved for entry into archive by Luciana Ferreira (lucgeral@gmail.com) on 2017-07-26T11:50:30Z (GMT) No. of bitstreams: 2 Dissertação - Mariana Dianese Alves de Moraes - 2017.pdf: 2047536 bytes, checksum: 5d64fdcced984e8d8b288a0799a3e6e0 (MD5) license_rdf: 0 bytes, checksum: d41d8cd98f00b204e9800998ecf8427e (MD5) / Made available in DSpace on 2017-07-26T11:50:30Z (GMT). No. of bitstreams: 2 Dissertação - Mariana Dianese Alves de Moraes - 2017.pdf: 2047536 bytes, checksum: 5d64fdcced984e8d8b288a0799a3e6e0 (MD5) license_rdf: 0 bytes, checksum: d41d8cd98f00b204e9800998ecf8427e (MD5) Previous issue date: 2017-02-06 / Conselho Nacional de Pesquisa e Desenvolvimento Científico e Tecnológico - CNPq / Knowledge of the quality and energy potential of african mahogany wood is essential for its proper use. The objective of this work was to evaluate the anatomical characteristics, basic density and energy potential of the wood of Khaya ivorensis (A. Chev) obtained from the thinning of a commercial plantation at the age of seven years. Five medium trees cut during thinning were selected and two wooden discs were cut in different longitudinal positions (0, DAP, 25, 50, 75 and 100% of the commercial height; minimum diameter of 6 cm): (i) one disc of each (Density, immediate chemical, higher calorific value and energy density), and the density and density of the vessels, (Density, energetic strength and energy density) were used to compose a sample of all longitudinal positions for the characterization of biomass, Durability, volumetric expansion and tensile strength by diametrical compression). For the production of charcoal, wedges of wood of discs of the height of the DAP (1.3 meters) were used, being carbonized in muffle furnace for the later characterization (basic and energetic density, immediate chemical, higher calorific value and gravimetric yield). The evaluations of the basic density and the anatomical characteristics of the wood of K. ivorensis at the age of 7, from thinning, showed that the exchange meristem is forming until this period the so-called juvenile wood, not being indicated for the solid uses such as furniture and floors. The models of the longitudinal profiles of the basic density of the wood were similar for all the trees, characterized by the decrease of the base to the intermediate portion, with increase to the apex of the trunk. For the length of the fibers the values did not present a model of common longitudinal variation along the trunk of the trees. The width, lumen diameter and wall thickness of the fibers presented a variation model characterized by lower values at the base of the trunk, followed by decreasing-stabilization of the values up to the top of the trunk. The characteristics of biomass and charcoal indicated lower quality parameters for energy use. Biomass carbonization promoted the reduction of basic density and volatile materials and the increase of fixed carbon, ash, higher calorific value and energy density. The compaction of the biomass improved its energetic and physical characteristics, being the indicated form for the energetic use of the juvenile wood of K. ivorensis, obtained in the thinning of the commercial plantation. / O conhecimento da qualidade e do potencial energético da madeira do mogno africano é fundamental para o seu uso adequado. O presente trabalho teve como objetivo avaliar as características anatômicas, densidade básica e potencial energético da madeira de Khaya ivorensis (A. Chev), obtida do desbaste de um plantio comercial, aos sete anos de idade. Foram selecionadas cinco árvores médias cortadas durante o desbaste e seccionados dois discos de madeira em diferentes posições longitudinais (0, DAP, 25, 50, 75 e 100% da altura comercial; diâmetro mínimo de 6 cm): (i) um disco de cada posição foi utilizado para a determinação da densidade básica, dimensões das fibras (comprimento, espessura da parede, diâmetro do lume e largura total) e dos vasos (diâmetro tangencial, frequência e área ocupada) e (ii) um disco utilizado para o corte de cunhas que foram, posteriormente, trituradas e moídas para compor uma amostra composta de todas as posições longitudinais para a caracterização da biomassa (densidade à granel, química imediata, poder calorífico superior e densidade energética) e produção e caracterização de briquetes (densidade aparente, energética, durabilidade, expansão volumétrica e resistência à tração por compressão diametral). Para a produção de carvão vegetal foram utilizadas cunhas de madeira de discos da altura do DAP (1,3 metros), sendo carbonizadas em forno mufla para a posterior caracterização (densidade básica e energética, química imediata, poder calorífico superior e rendimento gravimétrico). As avaliações da densidade básica e das características anatômicas da madeira de K. ivorensis com 7 anos de idade, proveniente do desbaste, apontaram que o meristema cambial está formando, até este período, o lenho denominado juvenil, não sendo indicada a sua utilização para os usos sólidos como movelaria e pisos. Os modelos dos perfis longitudinais da densidade básica do lenho foram similares para todas as árvores, caracterizados pelo decréscimo da base até a porção intermediária, com aumento até o ápice do tronco. Para o comprimento das fibras os valores não apresentaram um modelo de variação longitudinal comum ao longo do tronco das árvores. A largura, diâmetro do lume e espessura da parede das fibras apresentaram um modelo de variação caracterizado por menores valores na base do tronco, seguido de decréscimo-estabilização dos valores até o topo do tronco. As características da biomassa e do carvão vegetal indicaram parâmetros de qualidade inferiores para o uso energético. A carbonização da biomassa promoveu a redução da densidade básica e materiais voláteis e o aumento do carbono fixo, cinzas, poder calorífico superior e densidade energética. A compactação da biomassa na forma de briquetes melhorou as características energéticas e físicas, sendo a forma mais indicada para o uso energético da madeira juvenil, obtida no desbaste do plantio comercial, aos sete anos de idade.

Afin de valoriser au mieux le bois de feuillu de qualité secondaire, il est important de comprendre ses propriétés et d'optimiser ses propriétés mécaniques. La variabilité de certaines propriétés structurelles, physiques et mécaniques du bois de hêtres et de chênes issus d’éclaircis a été étudiée. Des billons tirés de ces arbres ont été déroulés en trois épaisseurs (2.1, 3, and 4.2 mm). La qualité du placage a été évaluée en mesurant la proportion du nœud et les fissurations. Les propriétés mécaniques du LVL (lamibois) réalisé à partir de ces placages ont été mesurées par méthode destructive et non destructive. Chez les deux espèces, le bois juvénile est concentré dans une petite zone près de la moelle, de sorte qu'il y aura peu ou pas de bois juvénile déroulé. Les propriétés mécaniques et physiques du bois des deux espèces sont fortement influencées par sa densité. Le placage de 3 mm d'épaisseur a fourni les propriétés mécaniques optimales pour le LVL pour les deux espèces. Ces propriétés mécaniques sont comparables à celles de LVL réalisés à partir d'autres essences de feuillus. Le placage interne a fourni du LVL avec une densité plus élevée, mais des propriétés mécaniques plus faibles en raison de sa proportion de nœuds plus élevée. / To improve the value of secondary quality hardwood, it is important to understand its wood properties and optimize its mechanical properties. The variability of some structural, physical and mechanical properties of wood from beech and oak trees issued from thinning was studied. Logs from these trees were rotary peeled in three different veneer thicknesses (2.1, 3, and 4.2 mm). Veneer quality was assessed by measuring the veneer knot proportion and checking properties. The mechanical properties of LVL (laminated veneer lumber) made from these veneers were measured using the static and dynamic method. Juvenile wood in both species is concentrated in a very small area near the pith thus there will be no or low part juvenile wood peeled. The wood mechanical and physical properties of both species were strongly influenced by the wood density. The 3 mm thick veneer provided the optimal mechanical properties for LVL for both species. Such mechanical properties are comparable to LVL made with other hardwood species. The internal veneer provided LVL with higher density, but lower mechanical properties as a result of its higher proportion of knots.

Dans le contexte du développement durable qui a vu l’introduction de la directive produits biocides BPD 98/8/CE, l’étude des méthodes innovantes de préservation du bois comme le traitement thermique revêt une importance prépondérante. Le traitement thermique du bois permet d’améliorer ses propriétés de résistance biologique, de stabilité dimensionnelle ainsi que son aspect esthétique, sans ajout de produit chimique. Les études actuelles sur la problématique de la qualité du bois traité thermiquement se focalisent sur les caractéristiques finales du bois déjà traité, l’influence des conditions de traitement ou encore l’effet essence. Les propriétés intrinsèques du bois avant le traitement ne sont pas encore prises en compte. Les propriétés du bois telles que la densité ou la composition chimique étant variables principalement sous l’effet de l’activité humaine comme la sylviculture, l’objectif de cette thèse était d’évaluer l’impact de cette variabilité chez le chêne sessile (Quercus petraea Liebll.) et le sapin (Abies alba Mill) sur leur modification par voie thermique. Un scanner et un micro-densitomètre à rayons X ont été utilisés pour caractériser la variation de la densité des échantillons de planches et des cernes de croissance provenant des arbres étudiés. Des traitements thermiques ont été réalisés dans un four pilote à conduction sous vide de type macro-thermobalance et un analyseur thermogravimétrique (ATG). Des analyses chimiques ont été également réalisées. Les résultats montrent qu’en prenant la perte de masse due à la dégradation thermique du bois comme réponse, les types de tissus du bois et la composition chimique influencent sa thermo-dégradation. Que ce soit chez Quercus petraea ou chez Abies alba, le bois de printemps était plus sensible au traitement thermique que son voisin de bois d’été. De plus, les portions radiales du tronc, du bois juvénile à l’aubier en passant par le bois mature, se dégradaient suivant des cinétiques différentes. En conclusion, la variation de la microstructure et la composition chimique de ces bois influencent leur cinétique globale de thermo-dégradation. La sylviculture impacte cette différence intraspécifique de cinétique de dégradation à l’échelle intra- et interarbre. En effet, dans le cas du sapin pectiné, une gestion très dynamique des forêts dans le but de stimuler la croissance rapide des arbres qui produisent de gros bois contenant des cernes très larges, est source de variation dans la structure anatomique et la composition chimique à l’intérieur des arbres en comparaison aux petits bois à croissance lente plus homogènes. Toutes ces analyses ont pour objectif final de comprendre le lien entre les propriétés initiales du bois et les modifications thermiques intervenant au cours du traitement afin d’apporter une information utile aux industriels lors du choix des pièces de bois destinées au traitement thermique en vue d’une amélioration de la qualité du bois traité thermiquement / In the context of sustainable development which has seen the introduction of the biocides directive BPD 98/8/CE in the EU, innovative wood preservation practices such as Heat Treatment (HT) become relevant. Wood HT, also termed wood thermal modification, is a physical modification technology by which wood is heated at around 200 °C in an inert atmosphere. The main purpose of the treatment is to improve the biological durability and dimensional stability of wood. Current studies on thermally modified wood (TMW) quality are focusing on treated material, on treatment conditions or on species effect on the end-product characteristics. Relatively little is known about the effect of intrinsic wood properties on its thermal modification. As wood properties vary especially under the influence of human activities through sylviculture, this thesis studied the effect of European oak and silver fir wood density and chemical composition on their thermal modification kinetic. An X rays computed tomography (CT) and densitometer were used to characterize wood samples. Boards were heat-treated by conduction under vacuum using a pilot furnace, whereas sawdust samples underwent thermo-gravimetric analysis under nitrogen. The analysis allowed finding intra- and interspecific variations, especially within growth rings and along radial direction (from pith to bark). Forest management impacted heat modification kinetic of the studied samples, especially in silver fir where fast grown wood was more sensitive to treatment. The finding will be used as additional information to the wood industry which will account for homogeneity of loadings destined to heat treatment

Holz aus Kurzumtriebsplantagen (KUP) mit Pappeln (Populus spp.) kann eine bedeutende Rolle im Mix der Bioenergieressourcen in Deutschland spielen. Trotz eines günstigen Energieinput-Energieoutput-Verhältnisses ist das Erreichen wirtschaftlich zufriedenstellender Ergebnisse mit KUP jedoch auf den meisten potenziellen Anbauflächen durch hohe Ansprüche der Pappeln an die Wasserversorgung erschwert. Hohe Produktivitätsraten der Pappeln sind an einen hohen Wasserverbrauch gebunden und viele Trockenheitsanpassungen führen zu deutlichen Ertragsrückgängen. In der vorgestellten Arbeit wurde eine sechsjährige Vollgeschwister-F1-Kartierungspopulation der Europäischen Zitter Pappel (Aspe, Populus tremula L., Wuchsperiode 1998-2003) physiologisch und genetisch untersucht, um mögliche Wege zu einer züchterischen Verbesserung der Trockenheitstoleranz von Pappeln diskutieren zu können. Dabei wurde das Zuchtziel der Trockenheitstoleranz als Minimierung der Ertragsrückgänge unter trockenen Bedingungen definiert. Neben wuchsleistungsbezogenen Größen (Biomassegesamtleistung (BM, oberirdische Dendromasse), Biomassezuwachs (iBM), Radialzuwachs (ir), Baumhöhe (h)) wurden physiologisch holzanatomische Eigenschaften untersucht, die retrospektiv anhand der Jahrringe messbar sind und zur nicht direkt messbaren Eigenschaft der Trockenheitstoleranz in einer Beziehung stehen. Diese waren die Kohlenstoff- und Sauerstoffisotopsignatur (δ13C, δ18O), die Faser- und Gefäßgliedlänge (FL, GL), die Gefäßlumenquerschnittsfläche (AG), die Gefäßdichte (GD), der potenziell Saftstrom leitende Querschnittsflächenanteil (LQ), der hydraulisch gewichtete Gefäßlumendurchmesser (Dh) und die röntgendensitometrische Holzdichte (RD). Um trockenheitsbedingte physiologische Reaktionen vom Einfluss der Juvenilität der Kartierungspopulation unterscheiden zu können, wurden regionaltypische Juvenilitätstrends der RD, FL, GL, und des ir anhand eines zweiten, für das östliche Deutschland repräsentativen Aspen-Kollektivs aus natürlicher Sukzession ermittelt. Bedingt durch Trockenheitsanpassungen bzw. eine bevorzugte Wurzelentwicklung nach dem Verpflanzen zeigte die Kartierungspopulation in den ersten drei Jahrringen Abweichungen von den regionaltypischen Juvenilitätstrends. In den Trendverläufen der Kartierungspopulation heben sich die beiden Trockenvegetationsperioden 2000 und 2003 ab, wobei bis zum Sommer 2003 infolge des Starkniederschlages 2002 (Flut) ein außergewöhnlich gutes Grundwasserangebot herrschte. Alle untersuchten phänotypischen Eigenschaften zeigten 2000 starke trockenheitsbedingte Abweichungen. Im Jahrring 2003 wichen nur die GL und die RD von ihren Juvenilitätstrends ab. Außerdem konnte anhand der δ13C und δ18O Werte eine signifikante Abnahme der Wassernutzungseffizienz bzw. eine Zunahme der Transpiration im Jahr 2003 gezeigt werden. Die übrigen Größen folgten ihren Juvenilitätstrends und stiegen an. Die Jahrringdatensätze 2000 und 2003 der RD waren nicht signifikant mit der BM korreliert, dagegen zeigten die δ13C Datensätze 2002 und 2003 schwach positive Korrelationen mit der BM. Der trockenheitstoleranteste Genotyp verband seine überdurchschnittliche BM mit einer hohen Wassernutzungseffizienz (angezeigt durch überdurchschnittliche δ13C Werte), mit einer überdurchschnittlichen AG und mit einer nicht unterdurchschnittlichen RD in Höhe des Populationsmittels. Aufgrund des Fehlens negativer Korrelationen zwischen BM und δ13C bzw. BM und RD in der vorliegenden Arbeit können δ13C und RD als nützliche Weiser für die Unterscheidung der Trockenheitstoleranz verschiedener Aspen zu züchterischen Zwecken vorgeschlagen werden. Außer der BM unter trockenen Bedingungen kann keine der untersuchten Eigenschaften als alleiniger Trockenheitstoleranzweiser empfohlen werden. Zu einer Trockenheitstoleranzbewertung sollten Merkmalspaare verwendet werden, von denen ein Merkmal positiv mit dem Ertrag korreliert ist und das andere eine Trockenheitsanpassung verkörpert. Dadurch werden sowohl das primäre Zuchtziel eines höchstmöglichen Ertrages als auch eine bessere Trockenheitsangepasstheit berücksichtigt. Zwei verschiedene Trockenheitstoleranzindizierungen wurden angewendet, um die Kartierungsnachkommen entsprechend ihrer Trockenheitstoleranz einem Ranking zu unterziehen. Dabei wurden in beiden Fällen Bäume mit einem höheren Ertrag besser platziert. Der zweite Schwerpunkt der Arbeit lag auf der genetischen Kartierung von Quantitative Trait Loci (QTL) für die untersuchten phänotypischen Jahrringeigenschaften mit Bezug zur Trockenheitstoleranz. Als Basis für das QTL Mapping wurden, der Pseudo-Testcross-Mapping-Strategie folgend, genetische Kopplungskarten für die Elternbäume der Kartierungspopulation konstruiert. Die maternale Karte (P. tremula, „Schandau 4“) deckte mit 157 Markern (144 AFLP, 13 SSR) in 30 Kopplungsgruppen 1.369 cM ab, die 21 paternalen Kopplungsgruppen mit 148 Markern (132 AFLP, 16 SSR) überspannten 1.079 cM des Genoms (P. tremula, „Lichtenhain 1“). Die im Vergleich zur haploiden Chromosomenzahl der Pappeln (19) hohen Zahlen an Kopplungsgruppen sowie die hohen Zahlen an Doublets und unkartierten Markern zeigten eine geringe Genomabdeckung an. So konnte nur eine begrenzte Zahl, höchstens zwei QTL für die untersuchten phänotypischen Jahrringeigenschaften mit Bezug zur Trockenheitstoleranz, kartiert werden. Ein QTL Bereich mit pleiotropem Effekt auf mehrere wachstumsbezogene Größen wurde auf der maternalen Kopplungsgruppe 1 (dem Populus Chromosom I zuzuordnen) detektiert. Die Signifikanz der Effekte dieses QTL auf den Radialzuwachs entwickelte sich steigend mit zunehmendem Baumalter. / Wood production in short rotation coppices (SRC) with poplar (Populus spp.) can contribute significantly to the future bio energy supply mix in Germany. Although the energy-input to energy-output ratio is rather good, SRC often do not meet cost effectiveness due to high water demand of poplar species. High biomass productivity depends on optimal water supply. Also, numerous adaptations to water deficits result in an undesirable decrease of yield. Combined physiological and genetic investigations were conducted within a six-year old F1-full-sib crossbred population of European aspen (Populus tremula L., growing period 1998-2003). Possible implications on selection, breeding or improvement of poplar cultivars showing a high tolerance to water deficits are discussed. For the work presented here, the breeding goal of higher water deficit tolerance was defined as the minimisation of yield losses under dry conditions. Beyond growth related traits (aggregate yield (BM), aboveground woody biomass), biomass increment (iBM) and radial increment (ir), physiological and wood anatomical traits were included; these are related to reactions to water deficit and are measurable on tree rings retrospectively. These traits were the Carbon- and Oxygen isotope ratios (δ13C, δ18O), the fibre length and vessel element length (FL, GL), the vessel lumen cross sectional area (AG), the vessel density (GD), the cumulative vessel lumen area to cross sectional area ratio (LQ), the hydraulically weighted mean vessel lumen diameter (Dh) and wood density assessed by X-ray densitometry (RD). To distinguish the drought induced physiologic reactions from juvenile developmental patterns of the mapping population, juvenile trends of RD, FL, GL, and ir, which are representative of habitats in south-eastern Germany, were investigated in a second aspen collective that was selected from natural succession. During the first three years, the mapping population showed deviations from the juvenile trends due to water deficit adaptations or preferential root development, respectively. Due to drought in the growing seasons of 2000 and 2003, the juvenile trends show outstanding values for both years, although ground water supply in 2003 was exceptionally good following the intense rain event of 2002 (Elbe flood 2002). The tree ring traits of both years stand out from the juvenile trends due to drought adaptations. In 2000, all phenotypic traits showed a significant deviation from their respective trends. In 2003, only GL and RD showed an adaptation to drought as observable by a deviation from their juvenile trends. A significant decrease in water use efficiency (WUE) and an increase in net transpiration, respectively, were shown for 2003 by means of δ13C or δ18O values. All other traits showed an increase following their juvenile trends. RD data for 2000 and 2003 were not significantly correlated with BM, but δ13C data (and therefore WUE) for 2002 and 2003 revealed a weakly positive correlation with BM. The genotype that was most tolerant to water deficits showed a combination of a superior growth with a superior WUE (by means of δ13C), a superior AG, and an RD close to but not less than the population average. Due to the lack of negative correlation between BM and δ13C or BM and RD in the present work, δ13C and RD can be valuable proxies for the determination of drought tolerance of aspen trees for tree improvement purposes. Aside from BM under dry conditions, no other traits that were investigated can be recommended as a stand-alone proxy for water deficit tolerance. For a water deficit tolerance evaluation, pairs of traits should be used, of which one trait is positively correlated with yield and the other represents a water deficit adaptation. Both the primary breeding goal of the highest possible yield as well as a better water deficit tolerance should always be considered in this context. Two different drought tolerance indices were used to rank the individuals of the mapping population according to their water deficit tolerance. In both cases, trees with higher BM were ranked better. The second focus of the present work is on genetic mapping of Quantitative Trait Loci (QTL) for the investigated tree-ring traits that refer to water deficit reaction. As a basis for the QTL-mapping approach, genetic linkage maps were constructed for each parental tree of the F1-full-sib crossbred mapping-population following the two-way pseudo-testcross mapping strategy. The maternal map (P. tremula, “Schandau 4”) consisted of 157 markers (144 AFLP, 13 SSR) in 30 linkage groups and covered 1,369 cM. The 21 linkage groups of the paternal map (P. tremula, “Lichtenhain 1”) covered 1,079 cM of the genome (144 AFLP, 13 SSR). Compared with the haploid chromosome number (19) of the Populus genome, the high number of linkage groups, doublets and unlinked markers indicated low genome coverage. Only a low number of QTL was detected, maximal two per in¬vesti¬gated phenotypic trait with a relation to water deficit tolerance. One QTL having a pleiotropic effect on several growth related traits was detected on the maternal linkage group 1 (corresponding to the Populus Chromosome I). The significance of the QTL effects seemed to increase with tree age.

Holz aus Kurzumtriebsplantagen (KUP) mit Pappeln (Populus spp.) kann eine bedeutende Rolle im Mix der Bioenergieressourcen in Deutschland spielen. Trotz eines günstigen Energieinput-Energieoutput-Verhältnisses ist das Erreichen wirtschaftlich zufriedenstellender Ergebnisse mit KUP jedoch auf den meisten potenziellen Anbauflächen durch hohe Ansprüche der Pappeln an die Wasserversorgung erschwert. Hohe Produktivitätsraten der Pappeln sind an einen hohen Wasserverbrauch gebunden und viele Trockenheitsanpassungen führen zu deutlichen Ertragsrückgängen. In der vorgestellten Arbeit wurde eine sechsjährige Vollgeschwister-F1-Kartierungspopulation der Europäischen Zitter Pappel (Aspe, Populus tremula L., Wuchsperiode 1998-2003) physiologisch und genetisch untersucht, um mögliche Wege zu einer züchterischen Verbesserung der Trockenheitstoleranz von Pappeln diskutieren zu können. Dabei wurde das Zuchtziel der Trockenheitstoleranz als Minimierung der Ertragsrückgänge unter trockenen Bedingungen definiert. Neben wuchsleistungsbezogenen Größen (Biomassegesamtleistung (BM, oberirdische Dendromasse), Biomassezuwachs (iBM), Radialzuwachs (ir), Baumhöhe (h)) wurden physiologisch holzanatomische Eigenschaften untersucht, die retrospektiv anhand der Jahrringe messbar sind und zur nicht direkt messbaren Eigenschaft der Trockenheitstoleranz in einer Beziehung stehen. Diese waren die Kohlenstoff- und Sauerstoffisotopsignatur (δ13C, δ18O), die Faser- und Gefäßgliedlänge (FL, GL), die Gefäßlumenquerschnittsfläche (AG), die Gefäßdichte (GD), der potenziell Saftstrom leitende Querschnittsflächenanteil (LQ), der hydraulisch gewichtete Gefäßlumendurchmesser (Dh) und die röntgendensitometrische Holzdichte (RD). Um trockenheitsbedingte physiologische Reaktionen vom Einfluss der Juvenilität der Kartierungspopulation unterscheiden zu können, wurden regionaltypische Juvenilitätstrends der RD, FL, GL, und des ir anhand eines zweiten, für das östliche Deutschland repräsentativen Aspen-Kollektivs aus natürlicher Sukzession ermittelt. Bedingt durch Trockenheitsanpassungen bzw. eine bevorzugte Wurzelentwicklung nach dem Verpflanzen zeigte die Kartierungspopulation in den ersten drei Jahrringen Abweichungen von den regionaltypischen Juvenilitätstrends. In den Trendverläufen der Kartierungspopulation heben sich die beiden Trockenvegetationsperioden 2000 und 2003 ab, wobei bis zum Sommer 2003 infolge des Starkniederschlages 2002 (Flut) ein außergewöhnlich gutes Grundwasserangebot herrschte. Alle untersuchten phänotypischen Eigenschaften zeigten 2000 starke trockenheitsbedingte Abweichungen. Im Jahrring 2003 wichen nur die GL und die RD von ihren Juvenilitätstrends ab. Außerdem konnte anhand der δ13C und δ18O Werte eine signifikante Abnahme der Wassernutzungseffizienz bzw. eine Zunahme der Transpiration im Jahr 2003 gezeigt werden. Die übrigen Größen folgten ihren Juvenilitätstrends und stiegen an. Die Jahrringdatensätze 2000 und 2003 der RD waren nicht signifikant mit der BM korreliert, dagegen zeigten die δ13C Datensätze 2002 und 2003 schwach positive Korrelationen mit der BM. Der trockenheitstoleranteste Genotyp verband seine überdurchschnittliche BM mit einer hohen Wassernutzungseffizienz (angezeigt durch überdurchschnittliche δ13C Werte), mit einer überdurchschnittlichen AG und mit einer nicht unterdurchschnittlichen RD in Höhe des Populationsmittels. Aufgrund des Fehlens negativer Korrelationen zwischen BM und δ13C bzw. BM und RD in der vorliegenden Arbeit können δ13C und RD als nützliche Weiser für die Unterscheidung der Trockenheitstoleranz verschiedener Aspen zu züchterischen Zwecken vorgeschlagen werden. Außer der BM unter trockenen Bedingungen kann keine der untersuchten Eigenschaften als alleiniger Trockenheitstoleranzweiser empfohlen werden. Zu einer Trockenheitstoleranzbewertung sollten Merkmalspaare verwendet werden, von denen ein Merkmal positiv mit dem Ertrag korreliert ist und das andere eine Trockenheitsanpassung verkörpert. Dadurch werden sowohl das primäre Zuchtziel eines höchstmöglichen Ertrages als auch eine bessere Trockenheitsangepasstheit berücksichtigt. Zwei verschiedene Trockenheitstoleranzindizierungen wurden angewendet, um die Kartierungsnachkommen entsprechend ihrer Trockenheitstoleranz einem Ranking zu unterziehen. Dabei wurden in beiden Fällen Bäume mit einem höheren Ertrag besser platziert. Der zweite Schwerpunkt der Arbeit lag auf der genetischen Kartierung von Quantitative Trait Loci (QTL) für die untersuchten phänotypischen Jahrringeigenschaften mit Bezug zur Trockenheitstoleranz. Als Basis für das QTL Mapping wurden, der Pseudo-Testcross-Mapping-Strategie folgend, genetische Kopplungskarten für die Elternbäume der Kartierungspopulation konstruiert. Die maternale Karte (P. tremula, „Schandau 4“) deckte mit 157 Markern (144 AFLP, 13 SSR) in 30 Kopplungsgruppen 1.369 cM ab, die 21 paternalen Kopplungsgruppen mit 148 Markern (132 AFLP, 16 SSR) überspannten 1.079 cM des Genoms (P. tremula, „Lichtenhain 1“). Die im Vergleich zur haploiden Chromosomenzahl der Pappeln (19) hohen Zahlen an Kopplungsgruppen sowie die hohen Zahlen an Doublets und unkartierten Markern zeigten eine geringe Genomabdeckung an. So konnte nur eine begrenzte Zahl, höchstens zwei QTL für die untersuchten phänotypischen Jahrringeigenschaften mit Bezug zur Trockenheitstoleranz, kartiert werden. Ein QTL Bereich mit pleiotropem Effekt auf mehrere wachstumsbezogene Größen wurde auf der maternalen Kopplungsgruppe 1 (dem Populus Chromosom I zuzuordnen) detektiert. Die Signifikanz der Effekte dieses QTL auf den Radialzuwachs entwickelte sich steigend mit zunehmendem Baumalter. / Wood production in short rotation coppices (SRC) with poplar (Populus spp.) can contribute significantly to the future bio energy supply mix in Germany. Although the energy-input to energy-output ratio is rather good, SRC often do not meet cost effectiveness due to high water demand of poplar species. High biomass productivity depends on optimal water supply. Also, numerous adaptations to water deficits result in an undesirable decrease of yield. Combined physiological and genetic investigations were conducted within a six-year old F1-full-sib crossbred population of European aspen (Populus tremula L., growing period 1998-2003). Possible implications on selection, breeding or improvement of poplar cultivars showing a high tolerance to water deficits are discussed. For the work presented here, the breeding goal of higher water deficit tolerance was defined as the minimisation of yield losses under dry conditions. Beyond growth related traits (aggregate yield (BM), aboveground woody biomass), biomass increment (iBM) and radial increment (ir), physiological and wood anatomical traits were included; these are related to reactions to water deficit and are measurable on tree rings retrospectively. These traits were the Carbon- and Oxygen isotope ratios (δ13C, δ18O), the fibre length and vessel element length (FL, GL), the vessel lumen cross sectional area (AG), the vessel density (GD), the cumulative vessel lumen area to cross sectional area ratio (LQ), the hydraulically weighted mean vessel lumen diameter (Dh) and wood density assessed by X-ray densitometry (RD). To distinguish the drought induced physiologic reactions from juvenile developmental patterns of the mapping population, juvenile trends of RD, FL, GL, and ir, which are representative of habitats in south-eastern Germany, were investigated in a second aspen collective that was selected from natural succession. During the first three years, the mapping population showed deviations from the juvenile trends due to water deficit adaptations or preferential root development, respectively. Due to drought in the growing seasons of 2000 and 2003, the juvenile trends show outstanding values for both years, although ground water supply in 2003 was exceptionally good following the intense rain event of 2002 (Elbe flood 2002). The tree ring traits of both years stand out from the juvenile trends due to drought adaptations. In 2000, all phenotypic traits showed a significant deviation from their respective trends. In 2003, only GL and RD showed an adaptation to drought as observable by a deviation from their juvenile trends. A significant decrease in water use efficiency (WUE) and an increase in net transpiration, respectively, were shown for 2003 by means of δ13C or δ18O values. All other traits showed an increase following their juvenile trends. RD data for 2000 and 2003 were not significantly correlated with BM, but δ13C data (and therefore WUE) for 2002 and 2003 revealed a weakly positive correlation with BM. The genotype that was most tolerant to water deficits showed a combination of a superior growth with a superior WUE (by means of δ13C), a superior AG, and an RD close to but not less than the population average. Due to the lack of negative correlation between BM and δ13C or BM and RD in the present work, δ13C and RD can be valuable proxies for the determination of drought tolerance of aspen trees for tree improvement purposes. Aside from BM under dry conditions, no other traits that were investigated can be recommended as a stand-alone proxy for water deficit tolerance. For a water deficit tolerance evaluation, pairs of traits should be used, of which one trait is positively correlated with yield and the other represents a water deficit adaptation. Both the primary breeding goal of the highest possible yield as well as a better water deficit tolerance should always be considered in this context. Two different drought tolerance indices were used to rank the individuals of the mapping population according to their water deficit tolerance. In both cases, trees with higher BM were ranked better. The second focus of the present work is on genetic mapping of Quantitative Trait Loci (QTL) for the investigated tree-ring traits that refer to water deficit reaction. As a basis for the QTL-mapping approach, genetic linkage maps were constructed for each parental tree of the F1-full-sib crossbred mapping-population following the two-way pseudo-testcross mapping strategy. The maternal map (P. tremula, “Schandau 4”) consisted of 157 markers (144 AFLP, 13 SSR) in 30 linkage groups and covered 1,369 cM. The 21 linkage groups of the paternal map (P. tremula, “Lichtenhain 1”) covered 1,079 cM of the genome (144 AFLP, 13 SSR). Compared with the haploid chromosome number (19) of the Populus genome, the high number of linkage groups, doublets and unlinked markers indicated low genome coverage. Only a low number of QTL was detected, maximal two per in¬vesti¬gated phenotypic trait with a relation to water deficit tolerance. One QTL having a pleiotropic effect on several growth related traits was detected on the maternal linkage group 1 (corresponding to the Populus Chromosome I). The significance of the QTL effects seemed to increase with tree age.