The West Pacific Warm Pool (WPWP) plays an important role in the global heat budget and global hydrologic cycle, so knowledge about its past variability would improve our understanding of global climate. Variations in WPWP precipitation are most notable during El Niño-Southern Oscillation events, when climate changes in the tropical Pacific impact rainfall not only in the WPWP, but around the globe. The stalagmite records presented in this dissertation provide centennial-to-millennial-scale constraints of WPWP precipitation during three distinct climatic periods: the Last Glacial Maximum (LGM), the last deglaciation, and the Holocene. In Chapter 2, the methodologies associated with the generation of U/Th-based absolute ages for the stalagmites are presented. In the final age models for the stalagmites, dates younger than 11,000 years have absolute errors of ±400 years or less, and dates older than 11,000 years have a relative error of ±2%. Stalagmite-specific 230Th/232Th ratios, calculated using isochrons, are used to correct for the presence of unsupported 230Th in a stalagmite at the time of formation. Hiatuses in the record are identified using a combination of optical properties, high 232Th concentrations, and extrapolation from adjacent U/Th dates. In Chapter 3, stalagmite oxygen isotopic composition (d18O) records from N. Borneo are presented which reveal millennial-scale rainfall changes that occurred in response to changes in global climate boundary conditions, radiative forcing, and abrupt climate changes. The stalagmite d18O records detect little change in inferred precipitation between the LGM and the present, although significant uncertainties are associated with the impact of the Sunda Shelf on rainfall d18O during the LGM. A millennial-scale drying in N. Borneo, inferred from an increase in stalagmite d18O, peaks at ~16.5ka coeval with timing of Heinrich event 1, possibly related to a southward movement of the Intertropical Convergence Zone (ITCZ). An inferred precipitation maximum (stalagmite d18O minimum) during the mid-Holocene in N. Borneo supports La Niña-like conditions and/or a southward migration of the ITCZ over the course of the Holocene as likely mechanisms for the observed millennial-scale trends. In Chapter 4, stalagmite Mg/Ca, Sr/Ca, and d13C records reflect hydrologic changes in the overlying karst system that are linked to a combination of rainfall variability and cave micro-environmental effects. Dripwater and stalagmite geochemistry suggest that prior calcite precipitation is a mechanism which alters dripwater geochemistry in slow, stalagmite-forming drips in N. Borneo. Stalagmite Mg/Ca ratios and d13C records suggest that the LGM climate in N. Borneo was drier and that ecosystem carbon cycling may have responded to the drier conditions. Large amplitude decadal- to centennial-scale variability in stalagmite Mg/Ca, Sr/Ca and d13C during the deglaciation may be linked to deglacial abrupt climate change events.

O objetivo da presente tese é o de documentar, em escala milenar, as variações paleoclimáticas e paleoceanográficas superficiais do Atlântico Sul Ocidental desde o Último Máximo Glacial. Com esta finalidade, foram obtidas estimativas de paleotemperatura, paleossalinidade e paleoprodutividade baseadas na análise de fauna de foraminíferos planctônicos, isótopos estáveis de oxigênio e carbono em Globigerinoides ruber e nove datações de radiocarbono em amostras de um testemunho coletado na Bacia de Santos. Os dados foram complementados com a reanálise de outro testemunho previamente coletado em outro setor da mesma Bacia. Os resultados demonstram que as mudanças paleoceanográficas registradas no período analisado podem ser parcialmente explicadas por variações na exportação de calor e sal para o hemisfério norte relacionadas à atividade da célula de transporte meridional e à configuração das correntes superficiais associadas à circulação atmosférica. Outra parte significativa da variabilidade parece estar relacionada à intensidade do Sistema de Monção da América do Sul resultante de variações na insolação de acordo com o ciclo de precessão. Os principais desvios da tendência de variação da composição isotópica da água do mar esperada em função das mudanças de insolação coincidem com os grandes pulsos de degelo ocorridos em torno de 19, 14 e 8,2 mil anos AP. / The goal of this thesis is to document the paleoclimatic and paleoceanographic changes occurred at the millennial scale since the Last Glacial Maximum at the surface Western South Atlantic. Paleotemperature, paleosalinity and paleoproductivity estimates were obtained for this purpose based on faunal changes of foraminifera assemblages, carbon and oxygen stable isotopes in Globigerinoides ruber and nine radiocarbon datings in a core retrieved at Santos Basin. These data were supplemented by the reanalysis of a second core previously analyzed from a different sector of the same basin. The results demonstrate that paleoceanographic changes recorded in the analyzed period may be partially explained by changes in the heat and salt export to the northern hemisphere related to the meridional overturning cell and the surface currents related to atmospheric circulation. Another significant portion of the observed variability seems to be related to changes in the strength of the South American Monsoon System resulting from insolation changes according to the precessional cycle. The main departures from the expected trend of variation in the isotopic composition of seawater related to insolation changes coincide with great meltwater pulses occurred around 19, 14, and 8.2 kyr BP.

This research is associated to the FAPESP 2011/0095-7 thematic project, \"Interdisciplinary palaeoenviromental studies at Espirito Santo coast\" and to the FAPESP Research Program about Climate Global Change (PFPMCG-FAPESP). The objectives were: (1) to characterize the late Pleistocene connection between Atlantic and Amazon rainforest and possible routes of interaction (coast, continent or both); (2) to identify environmental characteristics that indicates the Northeastern Espírito Santo and Southern Bahia as a stable forest since the late Pleistocene; (3) reconstitute the vegetation dynamics and infer climate fluctuations. Three sedimentary cores were collected: at northern Espirito Santo (ES) state, southern Bahia (BA) and eastern Minas Gerais (MG). Carbon and nitrogen isotope analyses and bioindicators in sediments/ soils, chronologically determined by radiocarbon dating, contributed to the local palaeoenvironmental reconstitution. At ES, from ~33,460 to ~13,740 cal yr BP the vegetation was composed of arboreal/ shrub cold adapted species and the organic matter recorded the dominance of C3 plants (?13C ~-28?), suggesting a cold and humid forest domain. From ~13,740 to ~9500 cal yr BP vegetation changes, with the increase of grasses and decrease of arboreal types. Between ~9500 and to ~4000 cal yr BP herbs were abundant with a probable mixture of C3 and C4 plants (?13C ~-24?) and algae, suggesting a higher seasonality between humid and dry season. From ~7300 cal yr BP until present the peat layer initiates its deposition, and the pollen records represents the current vegetation. At BA, radiocarbon dating in high-resolution, C and N isotopes and palynology each ~150 years from ~6400 to ~3400 cal yr BP showed detailed vegetational dynamics. From ~6400 to ~5620 cal yr BP the pollen record indicates abundance of trees and shrubs (up to 96%). From ~5620 to ~4850 cal yr BP occurred a slightly increase of herbs (28%) but the ?13C of ~-28% indicate C3 plants domain. From 4850 to ~3400 cal yr BP trees and shrubs abundance increases up to 94%, indicating the forest domain. The results confirm the forest stability and high local humidity, allowing to recognize the area as a probable refugee since 6500 cal yr BP. Vegetation, climate and the landscape has changed at MG since ~10,000 cal yr BP. From 10,600 to ~9500 cal yr BP, pollen was not preserved, C/N values (2 to 11) reveal the algal contribution, the ?13C (~-20?) indicate the mixture of C3 and C4 plants and radiocarbon date inversions indicate that the material was reworked. From 9,500 to 7,500 cal yr BP the vegetation was predominantly herbaceous with mixture of C3 and C4 plant sources (?13C ~-17?). From 7,500 to 7,000 cal yr BP C3 plants (?13C ~-30?) as trees/shrubs domains the polen record. From ~7,000 to ~6500 cal yr BP the pollen records the herbaceous abundance, dominated by C4 plants (?13C ~-12 to 18?). Peat deposition starts at ~6500 cal yr BP composed by C3 plants, mainly dominated by trees. Pollen records since at least early Holocene and the floristics of current vegetation did not show any disjunct specie with Amazon / Este projeto está associado ao temático FAPESP 2011/00995-7, \"Estudos paleoambientais interdisciplinares na costa do Espírito Santo\", e ao Programa FAPESP de Pesquisas sobre Mudanças Climáticas Globais (PFPMCG-FAPESP). Os objetivos foram: (1) caracterizar a conexão da Mata Atlântica e Floresta Amazônica e possíveis rotas de interação (litorânea, continental ou ambas); (2) identificar padrões que caracterizem a região da costa norte do Espírito Santo e sul da Bahia como refúgios florestais; (3) reconstituir a dinâmica da vegetação e inferir flutuações climáticas desde o Pleistoceno tardio. Três testemunhos sedimentares foram coletados: ao norte do Espírito Santo (ES), ao sul da Bahia (BA) e ao leste de Minas Gerais (MG). Análises isotópicas de carbono e nitrogênio e bioindicadores em sedimento/solos, cronologicamente determinados por datação 14C, contribuíram para a reconstituição paleoambiental nestas regiões. No ES, de ~33.460 a ~13.740 anos cal AP, a vegetação era composta por espécies de árvores/arbustos típicas de regiões frias com o domínio de plantas C3 (?13C ~-28?), sugerindo a presença de uma floresta fria e úmida. De ~13.740 a ~9500 anos cal AP ocorreu o aumento das herbáceas e diminuição dos tipos arbóreos. Entre ~9500 e ~4000 anos cal AP, as herbáceas tornaram-se abundantes com mistura de plantas C3 e C4 (?13C~-24?) e algas, sugerindo maior sazonalidade entre as estações seca e úmida. De ~7300 anos cal AP até o presente, iniciou-se a deposição da turfa e pólen da vegetação atual. Na BA, a alta resolução de datação C-14, isótopos de C e N e palinologia a cada ~150 anos entre ~6400 e ~3400 anos cal AP revelaram a dinâmica da vegetação. Entre 6400 e 5620 anos cal AP arbóreas/arbustivas (96%) são abundantes. De ~5620 até ~4850 anos cal AP ocorreu um pequeno aumento de herbáceas (28%), e os valores de ?13C ~-28% indicam o domínio de plantas C3. De 4850 a ~3400 anos cal AP, a abundância de arbóreas/arbustivas (94%), indica domínio florestal. Os resultados confirmam a estabilidade florestal e a umidade local, permitindo reconhecer esta área como um possível refúgio florestal desde pelo menos 6500 anos cal AP. Em MG, desde ~10.000 anos cal AP ao presente, a vegetação, o clima e a paisagem modificaram consideravelmente. De 10.600 a ~9500 anos cal AP não houve preservação de pólen, os valores de C/N (~2 e 11) revelam a contribuição algal, o ?13C (~-20?) indica mistura de plantas C3 e C4 e inversões nas datações indicam o retrabalhamento do material. De 9500 a 7500 anos cal AP a vegetação era predominantemente herbácea, com mistura de C3 e C4 (?13C ~-17?). De 7500 a 700 anos cal AP, plantas C3 (?13C ~-30?) arbustivo-arbóreas dominam o registro polínico. De ~7000 a 6500 anos cal AP as herbáceasC4 (?13C ~-12?) são abundantes. A deposição de turfa iniciou em ~6500 anos cal AP, composta por plantas C3 arbóreas. Registros palinológicos desde o Holoceno inferior e a avaliação florística atual não revelaram a presença de espécies de distribuição disjunta com a Amazônia

La Béringie, un vaste territoire qui s’étend de la Sibérie orientale au Territoire du Yukon, est perçue comme le point d’entrée des populations humaines en Amérique. A la fin du Pléistocène, ce territoire déglacé aurait constitué un refuge aux premières populations préhistoriques se dispersant hors d’Asie. Selon les données génétiques et paléo-génétiques, la Béringie fut occupée au cours du Dernier Maximum Glaciaire (19 000-23 000 cal BP, années calibrées Before Present) par une population humaine qui demeura génétiquement isolée durant près de 8000 à 9000 ans, donnant ainsi naissance à la lignée des Natifs Américains qui allaient se disperser, plus tard, au sud des masses glaciaires nordaméricaines et jusqu’en Amérique du Sud. Cette « Beringian standstill hypothesis », toutefois, ne trouva aucun soutien dans le registre archéologique : en Sibérie orientale, le plus ancien site est daté à 32 000 cal BP, tandis qu’en Alaska et au Yukon, la présence humaine ne remonte pas au-delà de 14 000 cal BP. Dans les années 70-80’s, le site des Grottes du Poisson-Bleu (Yukon) livra des outils en pierre et des ossements supposés modifiés par les humains, enfouis dans un dépôt loessique pléistocène ; les découvertes encouragèrent les archéologues J. Cinq-Mars et R. Morlan à évoquer l’hypothèse d’une occupation humaine sporadique dans le nord du Yukon entre 11 000 et 30 000 cal BP environ. La nature anthropogénique des échantillons osseux soumis aux datations radiocarbones ainsi que l’intégrité de la stratigraphie furent toutefois remises en question par une majorité d’archéologues. La présente dissertation propose une analyse archéozoologique et taphonomique rigoureuse et systématique des assemblages fauniques de mammifères des Grottes I et II dans le but d’appréhender les facteurs responsables de l’accumulation et de la modification du matériel osseux. De nouvelles datations radiocarbones effectuées par le laboratoire Oxford Radiocarbon Accelerator Unit sur des ossements portant des traces indéniablement culturelles permettent une datation précise de l’occupation humaine du site. Les résultats illustrent plusieurs traces de découpe sur des os de cheval, caribou, wapiti et possiblement bison et mouflon, tandis que des ossements de mammouth pourraient avoir été collectés pour l’industrie osseuse. Les nouvelles datations AMS suggèrent que les Grottes du Poisson-Bleu étaient occupées de façon sporadique entre 12 000 et 24 000 cal BP, soit pendant et après le Dernier Maximum Glaciaire. Le site offre ainsi un soutien archéologique à l’hypothèse de l’isolation génétique des populations béringiennes à l’origine des premières dispersions en Amérique. L’histoire taphonomique des Grottes du Poisson-Bleu rejoint celle des sites karstiques béringiens qui illustrent des occupations interspécifiques alternées entre carnivores et des fréquentations humaines de courte durée pour des activités de chasse. En outre, les altérations anthropiques sur des os de cheval des Grottes I et II ravivent le débat sur les extinctions de la mégafaune à la fin du Pléistocène (ca. 14 000 cal BP). Le site souligne l’incomplétude du registre archéologique et invite à multiplier les efforts de recherche en Béringie si l’on veut être à même de comprendre la préhistoire du peuplement des Amériques. / Beringia, a vast landscape stretching from eastern Siberia to the Yukon Territory, is thought to be the initial entry point of humans into North and South America. At the end of the Pleistocene, this unglaciated region constituted a refugium for the first prehistoric populations dispersing out of Asia. According to genetic and palaeogenetic data, Beringia was occupied during the Last Glacial Maximum (19 000-23 000 cal BP, calibrated years before present) by a human population that remained genetically isolated for about 8000 to 9000 years, leading to the divergence of the Native American lineage that would eventually disperse south of the ice-sheets into North and South America. The « Beringian standstill hypothesis » is not well supported in the archaeological record, however: in eastern Siberia, the oldest archaeological site is dated to 32 000 cal BP while in Alaska and the Yukon, evidence for a human presence doesn’t exceed 14 000 cal BP. Excavated in the 70s-80s, the Bluefish Caves site (Yukon) yielded stone tools and bone remains thought to have been culturally modified, buried in a Pleistocene loess deposit; the discovery encouraged archaeologists J. Cinq-Mars and R. Morlan to propose that humans occupied the caves sporadically between about 11 000 and 30 000 cal BP. The anthropogenic nature of the bone samples submitted for radiocarbon analysis and the stratigraphic integrity of the site didn’t convince the scientific community, however. The current dissertation proposes a rigorous archaeozoological and taphonomic analysis of the mammal bone assemblages of Caves I and II in order to identify the agents responsible for the accumulation and modification of the bone material. The results show several cut marks on bone specimens belonging to horse, caribou, wapiti and possibly bison and Dall sheep, while mammoth skeletal remains may have been collected for bone industry. New radiocarbon dates obtained by an Oxford laboratory (Oxford Radiocarbon Accelerator Unit) on bone bearing indisputable evidence of cultural modification allow the precise dating of the human occupation at the site. The AMS dates suggest that the Bluefish Caves were occupied sporadically between 12 000 to 24 000 cal BP, i.e., during and after the Last Glacial Maximum. The site, therefore, offers archaeological support for the Beringian standstill hypothesis. The taphonomic history of the Bluefish Caves, as well as other Beringian karstic sites, shows use of the caves by various carnivores and short-term human occupations for hunting activities. Moreover, cultural modifications on horse bone from Caves I and II enhance the debate surrounding the megafaunal extinctions at the end of the Pleistocene (ca. 14 000 cal BP). The site underlines the incompleteness of the archaeological record and invites us to expand research efforts in Beringia if we are to understand the prehistory of the first people of the Americas.

Tandis que la variabilité climatique à long terme du Quaternaire terminal (oscillations glaciaire/interglaciaire) est relativement bien appréhendée aujourd’hui, l'origine et la modalité des variations climatiques haute-fréquence séculaires à millénaires, depuis la très haute fréquence des cycles de Dansgaard-Oeschger (D/O) jusqu'à la plus grande périodicité des événements d'Heinrich, restent toujours matière à débat. Parmi les éléments encore équivoques se trouvent notamment les modalités de transferts de chaleur méridiens et latitudinaux. A ce titre, la connexion hydrologique entre l’Atlantique et la Méditerranée apparaît comme un exemple d’étude particulièrement approprié vu qu’elle illustre le couplage de phénomènes jouant selon un double gradient Est-Ouest et Nord-Sud, reliant ainsi processus subtropicaux et nord atlantiques. Nous avons ainsi ciblé nos analyses sur des séquences prélevées autour du détroit de Gibraltar, avec pour objectif d’étendre les connaissances spatiales et temporelles de la variabilité climatique haute fréquence des derniers 50 000 ans dans le secteur de l’Atlantique Est subtropical et de la Méditerranée occidentale. La méthodologie de cette thèse est ainsi basée sur une comparaison multi-proxies qui inclut des analyses micropaléontologiques (dinokystes et foraminifères planctoniques) et géochimiques (isotopes stables et alkénones). Nous avons ainsi pu tester la cohérence des changements hydrologiques de surface inter- et intra-bassins et tenter de caractériser la migration des fronts hydrologiques associés. Grâce aux sites des marges marocaine et portugaise, notamment, nous avons pu vérifier l’impact des cycles de D/O sur la variabilité de l’intensité des cellules d’upwelling côtières dans ce secteur et sur la dynamique de la veine d’eau méditerranéenne profonde (MOW), couplant signaux des masses d’eau superficielles (température, salinité et productivité) aux paléo-intensités de la MOW. / While the long-term climatic variability of the Quaternary is relatively well understood today, the causes and processes at the origin of the rapid and brutal climatic variability that characterized the last glacial period (Dansgaard-Oeschger cycles), including meridional and latitudinal heat transfer mechanisms, are still subject to debate. As such, studying the hydrological connection between the Atlantic and the Mediterranean Sea appears particularly appropriate since it illustrates the coupling acting on a double East-West and North-South gradient. It furthermore links subtropical processes and North Atlantic ones. We focused our analysis on sedimentary sequences retrieved around the Strait of Gibraltar, in order to extend the spatial and temporal knowledges about the impact of the high frequency climatic variability of the last 50 000 years in the sector of the eastern subtropical Atlantic and of the western Mediterranean Sea. The methodology of this thesis is based on a multi-proxy compilation coupling micropaleontological (dinocysts and planktonic foraminifera) and geochemical (stable isotopes and alkenones) approaches. We have tested the consistency of the sea-surface paleohydrological changes at inter-and intra-basins scales and tried to characterize the associated migration of the hydrological fronts. Sites of the Portuguese and Moroccan margins, in particular, allowed us to document the impact of the D/O cycles on the intensity of coastal upwelling cells as well as on the dynamics of the MOW, by coupling sea surface signals (temperature, salinity and productivity) to paleointensity of the MOW.

O objetivo do presente estudo é avaliar o uso associado dos proxies ?18O e Mg/Ca obtidos em testas de foraminíferos. Para alcançar tal objetivo foram realizadas análises de ?18O e Elemento/Ca em amostras de foraminíferos planctônicos (G. ruber e G. sacculifer) e bentônicos (Cibicidoides spp., C. corpulentus, Uvigerina spp. e H. elegans) de dois testemunhos do Atlântico Sul. Os resultados demonstraram que apesar dos proxies ?18O e Mg/Ca apresentarem perfis distintos em relação às espécies os mesmos se correlacionam de forma apropriada, sendo possível estimar a temperatura e os sinais de ?18OSW e ?18OLocal a partir dos mesmos. A partir dos dados de temperatura estimados com base em Mg/Ca foi possível identificar uma anomalia negativa das águas superficias entre o LGM e o HL de -1,5 ± 0,2 °C, enquanto que nas águas profundas observou-se uma anomali positiva de 1,7 ± 0,4 °C para o mesmo período. Os dados de ?18OSW estimados com base nos dados de temperatura e ?18O indicaram uma oscilação de 1,0 ± 0,1 ? e 2,0 ± 0,2 ? para as espécies planctônicas e bentônicas durante o LGM e o HL. Segundo o valor esperado para a variação do volume de gelo (~1,2 ?) há uma oscilação muito baixa da salinidade nas águas superficias da região de estudo, em contraste com uma oscilação bem marcada da salinidade nas águas de fundo. As estimativas de ?18OLocal indicaram uma oscilação entre o LGM e o HL muito pequena da salinidade (~0,1 ?) nas águas superficiais, com uma oscilação maior nas águas de fundo (1,0 ± 0,3 ?). O aumento da salinidade das águas de fundo durante o LGM corrobora a anomalia positiva da temperatura observada neste estudo, visto que, para a ocorrência da mesma, é necessário que a densidade das águas de fundo seja suficiente para manter a estratificação da coluna de água. / The goal of this study is to evaluate the associated use of the proxies ?18O and Mg/Ca, both analyzed in foraminifera tests. ?18O and trace metals analyses were performed in order to achieve this purpose on samples of planktonic (G. ruber and G. sacculifer) and benthic (Cibicidoides spp., C. corpulentus, Uvigerina spp. and H. elegans) species of foraminifera from two cores from South Atlantic. Despite of the fact that the proxies used on this study present distinct fits within the species they correlate very well, being possible to estimate the values of temperature, ?18OSW and ?18OLocal. Based on the temperature estimated by Mg/Ca it was possible to identify a negative anomaly of -1,5 ± 0,2 °C between the LGM and HL, meanwhile a positive anomaly of 1,7 ± 0,4 °C was observed in the deep waters for the same period. The ?18OSW data estimated by temperature and ?18O indicate an oscilation of 1,0 ± 0,1 ? and 2,0 ± 0,2 ? for planktonic and benthic species between the LGM and the HL. Based on the literature value for the ice volume signal (~1,2 ?) the superficial waters of the study area indicated a very low salinity oscilation, opposite to the high salinity oscillation in the deep waters. The ?18OLocal estimatives exhibited a very weak salinity oscillation between LGM and HL in the superficial waters (~0,1 ?), providing a much more strong oscilation in the deep waters (1,0 ± 0,3 ?). The salinity increase during the LGM in the deep waters establish the validity of the positive temperature anomaly observed in this study, since the density of the deep water needs to be adequate to maintain the water column stratification.

O objetivo do presente estudo é avaliar o uso associado dos proxies ?18O e Mg/Ca obtidos em testas de foraminíferos. Para alcançar tal objetivo foram realizadas análises de ?18O e Elemento/Ca em amostras de foraminíferos planctônicos (G. ruber e G. sacculifer) e bentônicos (Cibicidoides spp., C. corpulentus, Uvigerina spp. e H. elegans) de dois testemunhos do Atlântico Sul. Os resultados demonstraram que apesar dos proxies ?18O e Mg/Ca apresentarem perfis distintos em relação às espécies os mesmos se correlacionam de forma apropriada, sendo possível estimar a temperatura e os sinais de ?18OSW e ?18OLocal a partir dos mesmos. A partir dos dados de temperatura estimados com base em Mg/Ca foi possível identificar uma anomalia negativa das águas superficias entre o LGM e o HL de -1,5 ± 0,2 °C, enquanto que nas águas profundas observou-se uma anomali positiva de 1,7 ± 0,4 °C para o mesmo período. Os dados de ?18OSW estimados com base nos dados de temperatura e ?18O indicaram uma oscilação de 1,0 ± 0,1 ? e 2,0 ± 0,2 ? para as espécies planctônicas e bentônicas durante o LGM e o HL. Segundo o valor esperado para a variação do volume de gelo (~1,2 ?) há uma oscilação muito baixa da salinidade nas águas superficias da região de estudo, em contraste com uma oscilação bem marcada da salinidade nas águas de fundo. As estimativas de ?18OLocal indicaram uma oscilação entre o LGM e o HL muito pequena da salinidade (~0,1 ?) nas águas superficiais, com uma oscilação maior nas águas de fundo (1,0 ± 0,3 ?). O aumento da salinidade das águas de fundo durante o LGM corrobora a anomalia positiva da temperatura observada neste estudo, visto que, para a ocorrência da mesma, é necessário que a densidade das águas de fundo seja suficiente para manter a estratificação da coluna de água. / The goal of this study is to evaluate the associated use of the proxies ?18O and Mg/Ca, both analyzed in foraminifera tests. ?18O and trace metals analyses were performed in order to achieve this purpose on samples of planktonic (G. ruber and G. sacculifer) and benthic (Cibicidoides spp., C. corpulentus, Uvigerina spp. and H. elegans) species of foraminifera from two cores from South Atlantic. Despite of the fact that the proxies used on this study present distinct fits within the species they correlate very well, being possible to estimate the values of temperature, ?18OSW and ?18OLocal. Based on the temperature estimated by Mg/Ca it was possible to identify a negative anomaly of -1,5 ± 0,2 °C between the LGM and HL, meanwhile a positive anomaly of 1,7 ± 0,4 °C was observed in the deep waters for the same period. The ?18OSW data estimated by temperature and ?18O indicate an oscilation of 1,0 ± 0,1 ? and 2,0 ± 0,2 ? for planktonic and benthic species between the LGM and the HL. Based on the literature value for the ice volume signal (~1,2 ?) the superficial waters of the study area indicated a very low salinity oscilation, opposite to the high salinity oscillation in the deep waters. The ?18OLocal estimatives exhibited a very weak salinity oscillation between LGM and HL in the superficial waters (~0,1 ?), providing a much more strong oscilation in the deep waters (1,0 ± 0,3 ?). The salinity increase during the LGM in the deep waters establish the validity of the positive temperature anomaly observed in this study, since the density of the deep water needs to be adequate to maintain the water column stratification.

L’introduction de petites faunes au sein du régime alimentaire humain dans le passé constitue un sujet de recherche essentiel dans la compréhension des modes de subsistance des chasseurs-cueilleurs, notamment au Paléolithique supérieur. Durant cette période, et notamment au Dernier Maximum Glaciaire, le lapin de garenne (Oryctolagus cuniculus) domine les assemblages fauniques de nombreux sites archéologiques de la Péninsule Ibérique. Plusieurs hypothèses ont été avancées afin de comprendre le rôle que ce petit gibier a pu jouer dans la subsistance des groupes humains. Néanmoins, la multitude et la variété de ces hypothèses reflètent, d’un côté, le manque de consensus quant à la portée de l’exploitation du lapin dans la subsistance durant cette période dans la Péninsule Ibérique, et elles soulignent d’un autre côté que l’appréhension des implications économiques, technologiques, environnementales et sociales de son exploitation est une tâche complexe. A partir de ce double constat, j’ai dès lors établi comme problématique d’explorer les potentielles motivations liées à la chasse du lapin et d'appréhender le rapport entre les humains et cette espèce de petite taille, et ce dans l’objectif d’obtenir une meilleure compréhension des modes de subsistance adoptés pendant cette période en Ibérie. L’approche développée dans cette thèse repose sur la proposition que l’explication du pourquoi les humains exploitaient le lapin doit être traitée par une compréhension du comment les humains chassaient ce petit gibier. Si nous voulons analyser le régime alimentaire humain dans le passé, nous devons effectivement comprendre comment les humains prenaient leurs décisions cynégétiques. A cette fin, un des cadres théoriques principalement utilisé en archéologie, la Optimal Foraging Theory, a été employé, en complément de la modélisation à base d’agents. La problématique a été répartie en trois questions de recherche distinctes et complémentaires, chacune faisant l’objet d’une publication. Dans le premier article, l’application du Diet Breadth Model issu de la Optimal Foraging Theory a permis d’observer que l’exploitation d’une garenne est optimale lorsque plusieurs individus capturent un minimum de 7 lapins à l’aide de filets au cours du Dernier Maximum Glaciaire dans la Péninsule Ibérique. Dans le second article, nous avons mené des expériences de simulation à travers le développement d’un modèle à base d’agents pour évaluer les implications de ce type d’exploitation dans un cadre plus réaliste au sein duquel les humains interagissent, avant de l'intégrer, dans le troisième article, dans différents contextes socio-économiques qui lui sont associés et observés dans le registre ethno-historique. Nos résultats montrent que la composition du régime alimentaire humain et, par-là, le retour énergétique quotidien, sont influencés par l'organisation sociale entre les membres d'un groupe, la stratégie de chasse et la technologie associée afin d’exploiter une garenne. Particulièrement, la rentabilité de l’exploitation des ressources est encore plus grande si les femmes, les enfants et les ainés sont impliqués dans l’exploitation des garennes à travers des battues et/ou l’aide de filets. Par conséquent, cette thèse expose que le développement de formations socio-culturelles optimales basées sur une division des tâches de subsistance selon l’âge et/ou le sexe (et de la technologie qui leur est associée) a pu permettre aux groupes humains de s'adapter aux environnements de la Péninsule Ibérique au Dernier Maximum Glaciaire. Finalement, le lapin ayant probablement été un complément alimentaire important pendant cette période, cette thèse souligne la nécessité de considérer son exploitation (et celle de petites faunes en général) lors de l’interprétation du registre zooarchéologique car des changements d’un point de vue économique, technologique mais aussi socio-culturel dans les modes de subsistance adoptés dans le passé peuvent être ainsi appréhendés. / The introduction of small fauna into the human diet in the past is an essential research topic in the understanding of hunter-gatherer subsistence patterns, particularly in the Upper Paleolithic. During this period, and particularly during the Last Glacial Maximum, the European rabbit (Oryctolagus cuniculus) dominates the faunal assemblages of many archaeological sites in the Iberian Peninsula. Several hypotheses have been proposed to understand the role played by this small game in the subsistence of past human groups. Nevertheless, the number and diversity of these hypotheses first reflect a lack of agreement as to the extent to which rabbits played a role in subsistence during this period in the Iberian Peninsula. Secondly, they underline the complexity of understanding the economic, technological, environmental, and social implications of rabbit exploitation. From these observations, I established as the problematic to explore the potential motivations behind rabbit hunting, and to apprehend the relationship between humans and this small game, with the aim of providing a better understanding of the subsistence patterns adopted during this period in Iberia. The approach developed in this thesis is based on that the explanation of why humans exploited rabbits must be addressed by an understanding of how humans hunted them. If we are to analyze human diet in the past, we must indeed understand how humans made their hunting decisions. To this end, one of the theoretical frameworks mainly used in archaeology, the Optimal Foraging Theory, has been adopted, complemented by Agent-Based Modeling. I divided the problematic into three distinct and complementary research questions, each the subject of a publication. In the first article, the application of the Diet Breadth Model derived from the Optimal Foraging Theory enabled us to observe that the exploitation of a warren is optimal when several individuals capture a minimum of 7 rabbits using nets during the Las Glacial Maximum in the Iberian Peninsula. In the second paper, we carried out simulation experiments through the development of an Agent-Based Model to assess the implications of this type of exploitation in a more realistic environment in which humans interact, before integrating it, in the third paper, into different socio-economic contexts related to it and observed in the ethno-historical record. Our results show that the composition of the human diet and, consequently, the daily energetic return are influenced by the social organization between group members, the hunting strategy and the associated technology employed for exploiting a warren. In particular, the energetic efficiency of resource exploitation is even greater if women, children, and elders are involved in exploiting warrens through net hunting and/or drives. Consequently, this thesis argues that the development of optimal socio-cultural group configurations based on an age and/or gender-based division of labor (and their associated technology) may have enabled human groups to successfully adapt to the environments of the Iberian Peninsula during the Last Glacial Maximum. Finally, as rabbit was probably an important energetic source during this period, this thesis highlights the need to consider its exploitation (and that of small game in general) when interpreting the zooarchaeological record, as not only economic and technological, but also socio-cultural changes in subsistence patterns adopted in the past can be apprehended.

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Boreal forests are particularly vulnerable to climate change, experiencing a much more drastic increase in temperatures and having a limited amount of more northern refugia. The trees making up these vast and important ecosystems already had to adapt previously to environmental pressures brought about by the repeated glaciations during past ice ages. Studying the patterns of adaption of these trees can thus provide valuable insights on how to mitigate future damage. This thesis presents and analyses population structure, demo- graphic history and the distribution of fitness effects (DFE) of the diploid Betula pendula and tetraploid B. pubescens across Scandinavia. Birches–being widespread in boreal forests as well as having great economical importance–constitute superb model species. The analyses of this work confirm the expectations on postglacial population expansion and diploid-tetraploid introgression. They furthermore ascertain the presence of two genetic clusters and a remarkably similar DFE for the species. This work also contributes with a transparent, reproducible and reusable pipeline which facilitates running similar analyses for related species.