Em ambientes costeiros, a variabilidade da biomassa de fitoplâncton é dirigida por forçantes físicas locais que apresentam distintas escalas temporais em ocorrência, intensidade e durabilidade. No canal de São Sebastião, ambiente advectivo, parte-se do pressuposto de que qualquer adição da biomassa de fitoplâncton não se deve ao crescimento local, mas ao transporte de células fitoplanctônicas movidas pelas correntes guiadas a ventos e marés. Dessa maneira, o objetivo central deste trabalho é analisar a variabilidade em alta frequência das variações da biomassa fitoplanctônica e investigar suas relações temporais com a intensidade e direção dos ventos e a amplitude das marés. Em média, o acúmulo de clorofila é estável no tempo, com curtos episódios de lento aumento e diminuições bruscas da biomassa fitoplanctônica, sendo os valores encontrados relativamente baixos, característicos de ambientes oligomesotróficos. Análises espectrais da série temporal da concentração de clorofila-a e da altura da maré revelaram alta correlação entre as duas séries durante as marés de sizígia, com periodicidades de aproximadamente 12 e 24 horas. Em relação à direção e intensidade do vento, o espectro cruzado de ondaletas apresentou áreas de forte correlação com a clorofila-a e periodicidades dominantes de três dias. Além disso, observou-se que ventos vindos do quadrante norte/nordeste carregam águas com maiores concentrações de materiais particulados ou detritos. / In coastal environments, the dynamics of phytoplankton biomass is driven by local physical forcings that have different time scales in occurrence, intensity and durability. For being an advective environment, it is normally assumed that any positive change in phytoplankton biomass in the São Sebastião Channel is not due to local growth, but to the transport of phytoplankton cells moved by currents associated with winds and tides. Thus, the main objective of this project is to analyze the variability in high-frequency variations of the phytoplankton biomass and investigate their temporal relationships with intensity and direction of winds and high tides. On average, the chlorophyll accumulation is stable over time, with short episodes of slow rise and sudden decreases of phytoplankton biomass and relatively low values, characteristic of oligo-mesotrophic environments. Spectral analysis of the time series of chlorophyll a concentration and tide height showed a high correlation between the two series during the spring tides, and dominant periodicities of about 12 to 24 hours. Regarding direction and intensity of the wind, the cross wavelet spectrum presented strong correlation areas with chlorophyll a and dominant periodicities of 3 days. In addition, it was observed that winds from the north/northeast quadrant carry water with higher concentrations of particulate matter or debris.

Fundação Carlos Chagas Filho de Amparo a Pesquisa do Estado do Rio de Janeiro / The Amazon river, located in northernBrazil, discharges between 80,000 and 250,000 m3s-1 of water onto the adjacent shelf, creating a plume of brackish water that extends hundreds of kilometers away from the river mouth. This river also carries a large amount of fine sediments to the ocean where fluid mud has been found in the topset and upper foreset layers of the subaqueous delta formed on the mid-shelf. One of the main goals of this dissertation is to describe how turbulence and suspended sediment concentration vary along the Northern Channel of the Amazon river. Water column measurements were carried out in October 2008 at six anchor stations (P1, P3, P5, P6, P8 e P9) located seaward of the river mouth; P1 and P9 were 125 km apart. Each station was occupied during 13 hours during which current speed and direction were continuously sampled with a 600 kHz Teledyne-RDI ADCP; hourly profiles of temperature, salinity, turbidity and depth were also obtained. Water samples were collected for determination of Suspended Particulate Matter (SPM) concentration and calibration of the turbidity sensor. Current speed reached values above 1.5 m s1 in the along-channel direction (NE-SW); a remarkable ebb-flood asymmetry was observed and flows were strongly ebb-dominated. Throughout the water column, SPM concentration at stations P1 and P3 varied between 100 and 300 mg L1 in association with the presence of freshwater. In contrast, a strong salinity gradient was observed between stations P6 and P9, coinciding with the occurrence of concentrations of SPM above 10 g L-1 (fluid mud). At stations P3, P5 and P6, interface between freshwater from the Amazon river and salt water from the continental shelf, shear stresses wereestimated through four diferents methods: Reynolds, Turbulent Kinetic Energy (TKE), modified TKE and Quadratic Law; in the nearbed region (3 mab) the computed values varied between 0 and 3 Pa. At the three stations (P3, P5 and P6) the lowest and the highest shear stress values were obtained through, respectively, the Reynolds and the TKE methods. Over the whole water column turbulence intensity was estimated through the standard deviation of the turbulent component of the along-channel current velocity (root-mean square of u); from these values, it was estimated the turbulent dissipation of energy (G), whose values at 3 mab varied between zero and 20 s1.

Em ambientes costeiros, a variabilidade da biomassa de fitoplâncton é dirigida por forçantes físicas locais que apresentam distintas escalas temporais em ocorrência, intensidade e durabilidade. No canal de São Sebastião, ambiente advectivo, parte-se do pressuposto de que qualquer adição da biomassa de fitoplâncton não se deve ao crescimento local, mas ao transporte de células fitoplanctônicas movidas pelas correntes guiadas a ventos e marés. Dessa maneira, o objetivo central deste trabalho é analisar a variabilidade em alta frequência das variações da biomassa fitoplanctônica e investigar suas relações temporais com a intensidade e direção dos ventos e a amplitude das marés. Em média, o acúmulo de clorofila é estável no tempo, com curtos episódios de lento aumento e diminuições bruscas da biomassa fitoplanctônica, sendo os valores encontrados relativamente baixos, característicos de ambientes oligomesotróficos. Análises espectrais da série temporal da concentração de clorofila-a e da altura da maré revelaram alta correlação entre as duas séries durante as marés de sizígia, com periodicidades de aproximadamente 12 e 24 horas. Em relação à direção e intensidade do vento, o espectro cruzado de ondaletas apresentou áreas de forte correlação com a clorofila-a e periodicidades dominantes de três dias. Além disso, observou-se que ventos vindos do quadrante norte/nordeste carregam águas com maiores concentrações de materiais particulados ou detritos. / In coastal environments, the dynamics of phytoplankton biomass is driven by local physical forcings that have different time scales in occurrence, intensity and durability. For being an advective environment, it is normally assumed that any positive change in phytoplankton biomass in the São Sebastião Channel is not due to local growth, but to the transport of phytoplankton cells moved by currents associated with winds and tides. Thus, the main objective of this project is to analyze the variability in high-frequency variations of the phytoplankton biomass and investigate their temporal relationships with intensity and direction of winds and high tides. On average, the chlorophyll accumulation is stable over time, with short episodes of slow rise and sudden decreases of phytoplankton biomass and relatively low values, characteristic of oligo-mesotrophic environments. Spectral analysis of the time series of chlorophyll a concentration and tide height showed a high correlation between the two series during the spring tides, and dominant periodicities of about 12 to 24 hours. Regarding direction and intensity of the wind, the cross wavelet spectrum presented strong correlation areas with chlorophyll a and dominant periodicities of 3 days. In addition, it was observed that winds from the north/northeast quadrant carry water with higher concentrations of particulate matter or debris.

As the demands for energy increased with the global increase in population, there is a need to create and invest in more clean and renewable energy sources. Energy derived from the movement of the tides is an ancient concept that is currently being harnessed in a handful of large tidal range locations. However, the need to move from fossil fuel driven energy sources to those that are clean and non-polluting is a priority for a sustainable future. Globally, hydropower potential is estimated to be more than 16,400-Terawatt hours annually. Given that the electricity consumption worldwide was at 15,068-Terawatt hours in 2016, if properly utilized, hydropower could supply a substantial percentage of current demand. Most of the current hydropower supply is drawn from well-established dams and tidal barrage systems. However, tidal power plants that harness the change in water height and flow along the coast (i.e. using tidal energy) have the potential to push these figures even higher. Although there is no exact number for lengths of global coastlines, there are estimates that put that number between 220,000 and 880,000 miles of coasts. These opportunities in tidal energy technologies that harness energy from the sea may one day be the key to solving our energy crises. This research explored in detail a closed, convergent system for optimal extraction of head-driven tidal energy with minimal adverse environmental effects. The long-term goal of this project is to create a system that is viable in low tidal range locations traditionally not considered for locations of tidal energy systems, therefore increasing the overall global tidal energy portfolio. By implementing a closed system of ‘bladders’ and convergent nozzles to optimize the flow rate of the contained fluid, the proposed system can 1) derive tidal energy in low tidal range geographies 5 2) avoid typical hazards like system biofouling, marine life propeller impacts, and 3) allow for ease of installation, operation, and maintenance.

Fundação Carlos Chagas Filho de Amparo a Pesquisa do Estado do Rio de Janeiro / The Amazon river, located in northernBrazil, discharges between 80,000 and 250,000 m3s-1 of water onto the adjacent shelf, creating a plume of brackish water that extends hundreds of kilometers away from the river mouth. This river also carries a large amount of fine sediments to the ocean where fluid mud has been found in the topset and upper foreset layers of the subaqueous delta formed on the mid-shelf. One of the main goals of this dissertation is to describe how turbulence and suspended sediment concentration vary along the Northern Channel of the Amazon river. Water column measurements were carried out in October 2008 at six anchor stations (P1, P3, P5, P6, P8 e P9) located seaward of the river mouth; P1 and P9 were 125 km apart. Each station was occupied during 13 hours during which current speed and direction were continuously sampled with a 600 kHz Teledyne-RDI ADCP; hourly profiles of temperature, salinity, turbidity and depth were also obtained. Water samples were collected for determination of Suspended Particulate Matter (SPM) concentration and calibration of the turbidity sensor. Current speed reached values above 1.5 m s1 in the along-channel direction (NE-SW); a remarkable ebb-flood asymmetry was observed and flows were strongly ebb-dominated. Throughout the water column, SPM concentration at stations P1 and P3 varied between 100 and 300 mg L1 in association with the presence of freshwater. In contrast, a strong salinity gradient was observed between stations P6 and P9, coinciding with the occurrence of concentrations of SPM above 10 g L-1 (fluid mud). At stations P3, P5 and P6, interface between freshwater from the Amazon river and salt water from the continental shelf, shear stresses wereestimated through four diferents methods: Reynolds, Turbulent Kinetic Energy (TKE), modified TKE and Quadratic Law; in the nearbed region (3 mab) the computed values varied between 0 and 3 Pa. At the three stations (P3, P5 and P6) the lowest and the highest shear stress values were obtained through, respectively, the Reynolds and the TKE methods. Over the whole water column turbulence intensity was estimated through the standard deviation of the turbulent component of the along-channel current velocity (root-mean square of u); from these values, it was estimated the turbulent dissipation of energy (G), whose values at 3 mab varied between zero and 20 s1.

La détection et la caractérisation d'exoplanètes apparaissent clairement comme des thèmes centraux de l'observation astronomique pour les années à venir. Les projets spatiaux ou au sol sont nombreux (HARPS, CoRoT, Kepler, JWST, SPHERE...), mais les études théoriques visant à l'analyse et à la compréhension des données recueillies et à venir sont nécessaires. Durant cette thèse j'ai étudié divers processus physiques affectant la structure interne et l'évolution des planètes géantes, aussi bien au sein, qu'à l'extérieur de notre système solaire. J'ai notamment modélisé en détail: -L'impact de l'irradiation intense émise par l'étoile sur l'atmosphère d'une planète à faible distance orbitale, et l'effet induit sur l'évolution interne de cette planète. -Le couplage par dissipation de marée de l'évolution orbitale et thermique d'une planète interagissant avec sa proche étoile parente. -L'effet de la déformation due aux marées sur les paramètres observables d'une planète en transit grâce au suivi photométrique de son passage devant l'étoile. -L'incidence sur la structure et l'évolution d'une diminution de l'efficacité du transport de chaleur par convection due à un gradient d'éléments lourd dans l'enveloppe gazeuse d'une planète géante, conduisant au phénomène de convection double-diffusive. A travers l'étude des ces divers processus, j'ai développé différents modèles analytiques et codes numériques qui sont à la fois flexibles et robustes, et qui permettent maintenant d'étudier certaines propriétés des nouveaux objets substellaires détectés à mesure qu'ils sont découverts. / The detection and characterization of extrasolar planets clearly appears as one of the main goals of observational astronomy for the coming years. Space and ground project are numerous, but theoretical studies aimed at analyzing and understanding available and future data are needed. During this thesis, I study various physical processes affecting the internal structure and evolution of both solar, and extrasolar giant planets. In particular I investigate : -the impact of the intense stellar irradiation received by a close in planet on its subsequent internal evolution. This allows me to quantify the radius anomaly of bloated Hot Jupiters and to constrain their internal composition. -the tidal and centrifugal distortion of a fluid planet. By using both analytical and numerical models, I show how non-sphericity of the planet affects transit measurements, yielding an underestimation of its radius. -how the presence of double-diffusive convection caused by a heavy elements gradient in the gaseous envelope of a planet can decrease the efficiency of its internal heat transport, and affect its structure and evolution. -the coupling between the orbital and the thermal evolution of a planet arising from the strong star-planet tidal interaction. Subsequently, I find that tidal heating alone is not a viable explanation for the observed radius anomaly of transiting planets. Through these different studies, I developed various analytical models and numerical codes that are both flexible and robust, and which now allow one to study the properties of new extrasolar planets and brown dwarfs as they are discovered.

Les géomètres de la section Survey de l’Organisation Européenne pour la Recherche Nucléaire (CERN) utilisent le nivellement hydrostatique HLS pour effectuer des alignements verticaux précis. Le HLS atteint des précisions micrométriques, ce qui lui permet d’être utilisé pour les expériences à but de physique fondamentale comme le Large Hadron Collider (LHC). HLS mesure certes des déformations qui ont pour conséquence de désaligner tout accélérateur de particules lié au sol, mais il mesure également d’autres phénomènes aux caractéristiques bien particulières. Parmi ces phénomènes mesurés, les marées terrestres représentent une part très largement majoritaire du signal. Leur effet sur les mesures HLS est périodique et engendre une inclinaison longue base qui n’aboutit pas au désalignement relatif des aimants constitutifs d’un accélérateur. Les objectifs de ce doctorat sont de pouvoir prédire les effets ne perturbant pas l’alignement relatif d’un accélérateur de particules et ainsi corriger les mesures HLS de ces signaux. En effet, les tolérances planimétrique et altimétrique à respecter dans le domaine des accélérateurs de particules sont de plus en plus serrées. Par exemple, le Compact Linear Collider (CLIC), aujourd’hui à l’étude de faisabilité, nécessite une précision d’alignement à 3σ de 10 μm dans une fenêtre glissante de 200 m selon les directions transversale et verticale. Le HLS est candidat pour assurer cet alignement vertical mais l’amplitude de marée est d’environ +/-20 μm à 200 m, rendant nécessaire la prise en compte de ce phénomène longue base pour que l’instrumentation réponde aux besoins du CLIC. Ce doctorat est inspiré des travaux déjà réalisés sur les inclinomètres longue base et décrit les effets mesurés par HLS afin de classer ces phénomènes selon qu’ils désalignent ou non un accélérateur de particules. Enfin, les outils et modèles pour prédire les effets maitrisables sont utilisés pour anticiper les différents signaux mesurés par les HLS installés au CERN. / The surveyors of the Large Scale Metrology section of the European Organization for Nuclear Research (CERN) use hydrostatic levelling systems (HLS) to perform precise vertical alignment measurements. The HLS achieves micrometer accuracy, which allows it to be used for the fundamental physics experiments such as the Large Hadron Collider (LHC). An HLS measures the deformations that lead to the misalignment of any particle accelerator linked to the ground, but it also measures other phenomena with very particular characteristics. Among these measured phenomena Earth tides form the main part of the signal. Their effect on HLS measurements is periodic and produces a long baseline tilt that does not lead to a relative misalignment of the magnets that constitute the accelerator.The objectives of this doctoral research are to be able to predict the effects which do not disturb the relative alignment of a particle accelerator and to remove these signals from the HLS measurements. Indeed, the horizontal and vertical positioning tolerances to be respected in the realm of particle accelerators are becoming tighter and tighter. For example, the Compact Linear Collider (CLIC), currently the object of a feasibility study, requires a 3σ alignment accuracy of 10 μm in a sliding window of 200 m, in both the transverse and vertical directions. The HLS is a candidate for measuring the vertical alignment but the amplitude of the tidal effect is approximately +/-20 μm over200 m, making it necessary to take into account this long baseline phenomenon for the instrument to meet the CLIC requirements.This doctoral thesis is inspired by previous work on the long baseline tiltmeters and describes the effects measured by HLS in order to classify the measured phenomena according to whether they could produce a misalignment of a particle accelerator or not. Finally, the tools and models to predict those effects that are well understood are used to anticipate the different signals measured by HLS installed at CERN.

In this thesis we examine intra-seasonal oscillations (ISO) in the aqua-planet setup of the Community Atmospheric Model (CAM) version 5.1, mainly based on July and January climatological sea surface temperature (SST). We investigate mainly two questions -what should be the SST distribution for the existence of (a) northward moving ISO in summer, and (b) eastward moving MJO-like modes in winter. In the first part of the thesis we discuss the northward propagation. A series of experiments were performed with zonally symmetric and asymmetric SST distributions. The basic lower boundary condition is specified from zonally averaged observed July and January SST. The zonally symmetric July SST experiment produced an inter tropical convergence zone (ITCZ) on both sides of the equator. Poleward movement is not clear, and it is confined to the region between the double ITCZ. In July, the Bay of Bengal (BOB) and West Pacific SST is high compared to the rest of the northern tropics. When we impose a zonally asymmetric SST structure with warm SST spanning about 80 of longitude, the model shows a monsoon-like circulation, and some northward propagating convective events. Analysis of these events shows that two adjacent cells with cyclonic and anticyclonic vorticity are created over the warm SST anomaly and to the west. The propagation occurs due to the convective region drawn north in the convergence zone between these vortices. Zonally propagating Madden-Julian oscillations (MJO) are discussed in the second part of the thesis. All the experiments in this part are based on the zonally symmetric SST. The zonally symmetric January SST configuration gives an MJO-like mode, with zonal wave number 1 and a period of 40-90 days. The SST structure has a nearly meridionally symmetric structure, with local SST maxima on either side of the equator, and a small dip in the equatorial region. If we replace this dip with an SST maximum, the time-scale of MJO becomes significantly smaller (20-40 days). The implication is that an SST maximum in the equatorial region reduces the strength of MJO, and a flat SST profile in the equatorial region is required for more energetic of MJO. This result was tested and found to be valid in a series of further experiments.

Nous avons développé, et évalué en détail, un nouveau prototype d'inclinomètre longue base à niveaux hydrostatiques appelé " wth2o ". Ce système, aux principes particulièrement simples grâce à l'absence de pièces mécaniques mobiles, présente une grande fiabilité et une excellente stabilité (dérive linéaire de 0.05 µrad par mois). Sa haute résolution jusque dans la gamme des ondes sismiques longues périodes (où, par exemple, la résolution est meilleure que 0.001 masec, soit 5. 10-12 rad), et son niveau de bruit très bas, ont permis d'obtenir des résultats d'analyses harmoniques de marées terrestres (5 ans d'enregistrements) en excellent accord avec les modèles, et dont les écarts quadratiques moyens sont les plus bas de toutes les mesures inclinométriques réalisées au Laboratoire de Géodynamique de Walferdange (Luxembourg). A titre d'exemple, l'amplitude de l'onde M2 est déterminée avec une incertitude de seulement 0.003 masec, tandis que sa phase est déterminée avec un EQM de 0.028°, ce qui correspond à une incertitude de seulement 3.3 secondes. Cet instrument permit également d'observer des phénomènes rarement mesurés avec ce type d'appareil, tel les modes sphéroïdaux et toroïdaux les plus graves des oscillations libres de la Terre excitées par le séisme du Dénali (Mw 7.9) en novembre 2002, ou les passages successifs des ondes de Love jusqu'à G7, correspondant à 3 révolutions de ces ondes de surface autour du globe. Il fut également possible de séparer pour la première fois dans une analyse harmonique de marée terrestre inclinométrique, les constituants des groupes ter- et quater-diurnes. Ces très petites ondes enregistrées sont vraisemblablement liées aux ondes de marées océaniques propres aux eaux peu profondes présentes en Mer du Nord. Les modèles théoriques détaillés de cet appareil (comprenant des composantes relatives à l'amortissement produit par l'écoulement des fluides entre les électrodes des capteurs capacitifs) ont permis de dériver les solutions des équations du mouvement en composante inclinométrique et accélérométrique. De ces solutions furent tirées les formes théoriques des fonctions de transfert de l'appareil. Ces fonctions de transferts furent comparées avec succès aux mesures expérimentales de la réponse en fréquence. Grâce aux formes analytiques des fonctions de transfert en composante inclinométrique et accélérométrique qui sont données dans le présent travail, il est possible de calculer les caractéristiques géométriques optimales pour la construction d'un prototype devant répondre aux besoins particuliers d'une nouvelle application. Une étude originale des effets de ménisques (déformations de l'interface des fluides au contact de la paroi solide des pots) a montré que, s'ils n'ont pas d'influence sur les mesures en mode différentiel, ils peuvent par contre introduire des erreurs de plusieurs pour-cent sur l'estimation de la sensibilité par déplacement d'une quantité de liquide supposée connue. Cette erreur, ne dépendant que des propriétés physico-chimiques des fluides et matériaux en contacts, provient des variations de volumes de liquide contenu dans les ménisques lors des mouvements des interfaces. Si ces mouvements sont effectués lors des étalonnages sans que la ligne de contact ne se déplace, ces erreurs [en %] seront indépendantes du volume déplacé. Ces erreurs ne sont pas particulières à notre instrument mais peuvent affecter les étalonnages réalisés de la sorte pour tout inclinomètre à niveaux hydrostatiques, quel que soit le principe de mesure de l'instrument (mesures différentielles de pression, positionnement de flotteurs etc…). / We developed and evaluated a new prototype of long base water-tube tiltmeter named "wth2o". This system, particularly simple by the absence of moving parts, showed a great reliability and a fairly high stability (linear drift rate of 0.05 µrad/month). We analyzed a 5 years-long data set of Earth tides measurements performed in the Underground Laboratory for Geodynamics in Walferdange (Luxemburg). Its high resolution up to the long-period seismic band (where for instance the resolution is better than 0.001 masec, or 5. 10-12 rad) and its very low noise rate enabled us to obtain results in excellent accordance with the models, with the lowest root mean squares among all the results obtained with other tiltmeters in Walferdange. For instance, the amplitude of the M2 wave is estimated with a RMS as small as 0.003 masec and its phase is determined with an uncertainty below 0.028°, which represents 3.3 seconds only. With the "wth2o" we have also observed some events rarely measured with a water-tube, such as the gravest toroidal and spheroidal modes of the Earth free oscillations excited by the Mw 7.9 Denali earthquake or the successive passages of Love waves (up to G7) circling the globe. For the first time in an analysis of tilt Earth tide measurements, it was possible to separate the small constituents of the ter- and quater-diurnal band. The presence of these very small waves is most likely due to the effects of the shallow-water tides known to be remarkable in the North Sea. Theoretical models of this instrument (taking into account the damping produced by the liquids' flow between the plates of the capacitive sensors) allowed to obtain the solutions of the equations of motion for tilt as well as for acceleration. From these solutions we were able to produce very accurate theoretical transfer functions, as confirmed by the successful comparison with observed frequency responses. Thanks to the analytical tilt and acceleration transfer functions given in the present document it is possible to calculate the best geometrical characteristics for the construction of a new prototype having to respond to specific requirements of a new application. A new extended study of the menisci effects (deformations of the interface of the fluids in contact with the solid wall of the end vessels) showed that they do not influence the differential measurements but they could introduce errors of a few percents in the calibration factor when this factor is evaluated by the displacement of a "known" volume of liquid. This error depends only on the physical and chemical properties of the fluids and solids put in contact. It is due to the variation of volume of liquid trapped in the menisci while the interface is moving up- or downwards. If these interface movements are made without displacing the contact line, the errors [in %] will remain the same whatever the displaced volume of liquid could be. These errors can not only affect our results but can also be found in the calibration procedure of any kind of water-tube (central differential pressure, float positioned etc…).

Les plages de sable sont des écosystèmes dynamiques couvrant 70% des côtes mondiales. Ces systèmes abritent un cortège spécifique unique et assurent des fonctions essentielles de nourricerie, de nurserie et d’épuration des eaux. Or à proximité des côtes, la population humaine est en développement constant ce qui accroît les multiples pressions d’origine anthropique sur les écosystèmes côtiers. L’apport en excès de nutriments constitue une menace majeure qui peut se traduire par le développement d’importantes biomasses de macroalgues opportunistes (eutrophisation). La fréquence et l’intensité de ces blooms, communément formés de chlorophycées à court cycle de vie (ulves) et appelés marées vertes (MV), s'amplifient sur les côtes françaises et dans le Monde menaçant le fonctionnement de systèmes prépondérants et uniques.La plupart des études visant à déterminer les effets des MV sur la structure et le fonctionnement d’écosystèmes sédimentaires ont été conduites dans des environnements abrités, micro- ou atidaux. Cette problématique est restée presqu’inexplorée dans des écosystèmes plus dynamiques (systèmes ouverts et macrotidaux) en raison des difficultés de mise en place d’échantillonnage et de détection des effets de stress d’origine anthropique, inhérentes à la variabilité de ces systèmes. Cette thèse a donc eu pour objectif principal de combler ce manque et produire des connaissances en étudiant les réponses in situ des communautés benthiques de plages de sable fin macrotidales en présence ou non de MV. Quatre études ont été menées à différentes échelles spatio-temporelles et en considérant différents compartiments biologiques pour répondre à cette question générale.Ce travail de thèse montre qu’à l’échelle de la région Bretagne (variabilité intégrée sur 2700km de côtes et 7 ans) les communautés benthiques d’écosystèmes dynamiques sont modifiées significativement et de manière conservative par la présence de MV. Ce travail démontre aussi que les marées vertes impactent différemment la faune benthique en fonction du type d’habitat (plages semi-exposées vs. exposées), de la profondeur (mediolittoral vs. infralittoral), et du compartiment biologique (macrofaune benthique vs. juvéniles de poissons plats). Ces comparaisons ont permis d’identifier la faune benthique de médiolittoral inférieur des plages exposées comme étant le système le plus affecté par les MV. L’étude des variations à fine échelle spatio-temporelle de ce dernier montre que les caractéristiques faunistiques (uni- et multi-variées) sont modifiées le long d’un gradient de couverture d’algues vertes. Par exemple, la β-diversité décroît significativement le long de ce gradient. Afin d’explorer les processus pouvant expliquer ces modifications, et déterminer si ces changements ont des répercussions sur le fonctionnement de l’écosystème « plage de sable », les effets de l’accumulation d’ulves sur le réseau trophique à différent(e)s niveaux/échelles ont été mesurés. Les résultats de cette étude montrent qu’une importante biomasse d’ulves induit un changement de la structure entière du réseau trophique et une modification importante du fonctionnement trophique des plages. Les expérimentations menées au cours de cette thèse montrent que les changements observés sont induits par des effets directs (consommation de débris d’ulves) et indirects (modifications d’autres sources de nourriture) de la présence des MV.Cette thèse propose un cadre de travail visant à mieux détecter les effets de stress anthropiques sur la structure et le fonctionnement d’écosystèmes dynamiques. Dans un contexte de changement global forçant les écosystèmes à faire face à de multiples stress, cette approche pourrait se révéler particulièrement utile pour démêler, comprendre et prédire les effets de perturbations induites par les activités humaines sur le fonctionnement des écosystèmes et constituer une aide à la gestion de ces environnements particuliers. / Highly dynamic systems, often considered as resilient systems, are characterised by abiotic and biotic processes under continuous and strong changes in space and time. Because of this variability, the detection of overlapping anthropogenic stress is challenging. Coastal areas harbour dynamic ecosystems in the for of open sandy beaches, which cover the vast majority of the world’s ice-free coastline. These ecosystems are currently threatened by increasing human-induced pressure, among which mass-development of opportunistic macroalgae (mainly composed of Chlorophyta, so called green tides), resulting from the eutrophication of coastal waters. The ecological impact of opportunistic macroalgal blooms (green tides, and blooms formed by other opportunistic taxa), has long been evaluated within sheltered and non-tidal ecosystems. Little is known, however, on how more dynamic ecosystems, such as open macrotidal sandy beaches, respond to such stress. This thesis assesses the effects of anthropogenic stress on the structure and the functioning of highly dynamic ecosystems using sandy beaches impacted by green tides as a study case. The thesis is based on four field studies, which analyse natural sandy sediment benthic community dynamics over several temporal (from month to multi-year) and spatial (from local to regional) scales. In this thesis, I report long-lasting responses of sandy beach benthic invertebrate communities to green tides, across thousands of kilometres and over seven years; and highlight more pronounced responses of zoobenthos living in exposed sandy beaches compared to semi-exposed sands. Within exposed sandy sediments, and across a vertical scale (from inshore to nearshore sandy habitats), I also demonstrate that the effects of the presence of algal mats on intertidal benthic invertebrate communities is more pronounced than that on subtidal benthic invertebrate assemblages, but also than on flatfish communities. Focussing on small-scale variations in the most affected faunal group (i.e. benthic invertebrates living at low shore), this thesis reveals a decrease in overall beta-diversity along a eutrophication-gradient manifested in the form of green tides, as well as the increasing importance of biological variables in explaining ecological variability of sandy beach macrobenthic assemblages along the same gradient. To illustrate the processes associated with the structural shifts observed where green tides occurred, I investigated the effects of high biomasses of opportunistic macroalgae (Ulva spp.) on the trophic structure and functioning of sandy beaches. This work reveals a progressive simplification of sandy beach food web structure and a modification of energy pathways over time, through direct and indirect effects of Ulva mats on several trophic levels. Through this thesis I demonstrate that highly dynamic systems respond differently (e.g. shift in δ13C, not in δ15N) and more subtly (e.g. no mass-mortality in benthos was found) to anthropogenic stress compared to what has been previously shown within more sheltered and non-tidal systems. Obtaining these results would not have been possible without the approach used through this work; I thus present a framework coupling field investigations with analytical approaches to describe shifts in highly variable ecosystems under human-induced stress.