The energy exchange in the soil-snow-vegetation-atmospheresystem was studied to improve the quantitative knowledge of thegoverning processes. The lack of such knowledge contributes tothe uncertainty in the applicability of many existing modelsindependent of the temporal or spatial scale. The theoreticalbackground and available methods for measurements and numericalsimulations were reviewed. Numerical simulation models andavailable data sets representing open land and boreal forestwere evaluated in both diurnal and seasonal time-scales.Surface heat fluxes, snow depth, soil temperatures andmeteorological conditions were measured at an agriculturalfield in central Sweden over two winters, 1997-1999. Twoone-dimensional simulation models of different complexity wereused to simulate the heat and water transfer in thesoil-snow-atmosphere system and compared with the measurements.Comparison of simulated and observed heat fluxes showed thatparameter values governing the upper boundary condition weremore important than the formulation of the internal mass andheat balance of the snow cover. The models were useful toevaluate the lack of energy balance closure in the observedsurface heat fluxes, which underlined the importance ofimproved accuracy in eddy correlation measurements of latentflow during winter conditions. The representation of boreal forest in the land surfacescheme used within a weather forecast model was tested with athree-year data set from the NOPEX forest site in centralSweden. The formulation with separate energy balances forvegetation and the soil/snow beneath tree cover improvedsimulation of the seasonal and diurnal variations of latent andsensible heat flux compared with an older model version.Further improvements of simulated surface heat fluxes could beexpected if the variation of vegetation properties within andbetween years and a new formulation of the boundary conditionsfor heat flux into the soil is included. Keywords: Surface energy balance, Snow, Boreal forest,SVAT models, Eddy-correlation Measurements, Latent heat flux,Sensible heat flux, Net radiation, Soil temperature,Aerodynamic roughness, Surface resistance / QC 20100614

The aim of this thesis was to investigate the processes behind an unexpected runoff behaviour after a clear-cut in a boreal forest in northern Sweden (Balsjö). The risks of increased flooding, erosion, nutrient leakage and changes in the local ecosystems are some reasons why it is important to fully understand the effect of clear-cuts on the water balance. In northern boreal forests the snow is of great importance as it results in the main hydrological event of the year, the spring flood. In general, open areas accumulate more snow, have a lower evapotranspiration and therefore maintain a higher runoff than a forest. In a recent paired catchment study at Balsjö the expected pattern after a clear-cut was only shown in three out of five years (2007-2011). The expected increase in runoff did not occur in 2010 and 2011. Two hypothesized alternatives were year-to-year variation of ET or changes in soil water storage. In order to investigate this further the rainfall-runoff model HBV was used. First, the model was calibrated for the forest catchment (Ref) and the clear-cut catchment (CC), using observed data from Balsjö. To account for parameter uncertainty the calibration was performed using parameter optimization, resulting in 100 different parameter sets. Model results were evaluated using observed snow data from Balsjö and ET from Flakaliden, a nearby forest. Both the simulated snow and ET were quite consistent with the observed values. Finally the annual and the spring water balance were studied, using the simulated data. The simulated results did not detect the unexpected runoff behavior for the two years as clearly as the observations. The reason for this was that the model was calibrated for all five years, which meant that annual variations were not taken into account. The hypothesis, that higher ET could be the reason for the unexpected runoff behavior, could neither be dismissed nor confirmed by this thesis. This was because there were no observed data for the clear-cut area and limitations within the HBV model, which meant that sublimation and interception processes could not be analyzed separately. The model results indicated that the change in soil water storage was a more likely explanation for the unexpected runoff behavior. The simulation result showed that the meltwater was stored in the soil water storage. However, this theory does not seem likely since a clear-cut is normally wetter than a forest. The results of this thesis are consistent with other studies as they indicate that clear-cut effects should be studied seasonally as well as annually. The special feature of this thesis was the opportunity to study observed ET and investigate its influence on the water balance.

Koldioxidhalterna stiger i världen, vilket bidrar till global uppvärmning. Skogen har en viktig roll i klimatarbetet då stora mängder kol finns lagrad i skog och mark. Speciellt mycket kol finns i boreal skog, som bland annat finns i norra Sverige. Idag pågår en debatt huruvida det är mer fördelaktigt att behålla gammal skog som kolsänka, eller avverka den för att ge plats åt yngre träd. Syftet med litteraturstudien var att ge en klargörande bild över hur åldern på boreal skog påverkar ekosystemets kolbalans, genom att undersöka skogens kolupptagningshastighet samt möjlighet till kollagring. Studien visade att äldre skog har ett större totalt kollager, att medelåldrig skog har högst tillväxthastighet (nettoekosystemproduktion, NEP och nettoprimärproduktion, NPP) samt att NEP är negativ de första 10 åren. Studien visade på osäkerhet kring kolupptag i gamla skogar. Flera variabler, såsom substitution, störningar och biologisk mångfald är inte undersökta i studien och behövs tas i beaktning vid beslutsfattande. Trots detta, antyder litteraturstudien att det är fördelaktigt att låta gammal skog stå kvar som kolsänka, eftersom koldioxidutsläppen behöver minska snarast. Ytterligare ett syfte med studien var att kartlägga de missuppfattningar elever, i senare delen av skolan, har gällande kolets kretslopp. Studien visade på missuppfattningar gällande fotosyntes och respiration hos växter, nedbrytarnas roll samt sammanblandning av begrepp. För att åtgärda problemet kan uppgifter göras där atomer följs över olika nivåer, för att sätta processerna i sitt sammanhang. Stort fokus gällande missuppfattningarna ligger på fotosyntes och respiration och inga artiklar hittades gällande exempelvis det snabba, respektive långsamma, kretsloppet. Få studier är gjorda på svenska elever och då svensk kursplan lägger fokus på samband mellan olika nivåer kan missuppfattningarna vara i annorlunda i Sverige. Hur missuppfattningarna ser ut i svensk skola är föremål för vidare forskning. / The amount of carbon dioxide is rising in the atmosphere, which contributes, to global warming. Since large amounts of carbon are stored in forests, forests play an important part the efforts to improve the environment. Boreal forests, found in e.g. the northern part of Sweden, store especially large amounts of carbon. Today it is being debated whether it is more beneficial for the environment to keep old forests as a carbon sink, or to cut them down in order to make room for younger trees. The purpose of this literature study was to clarify how the age of the forest affects the carbon balance of the ecosystem, by examining the forest’s rate of carbon sequestration and capability of carbon storage. The literature study concluded that older forests, in total, store larger amounts of carbon, while middle-aged forests have he highest growth rate (net ecosystem production, NEP, and gross primary production, GPP) and that NEP is negative in the first ten years. The study showed some uncertainty concerning carbon uptake in old forests. Several variables, such as the effects of substitution, disturbances and biodiversity were not examined in this study and need to be further considered before making decisions. However, the literature study suggests that it is more beneficial to leave old forests as a carbon sink, since emissions of carbon dioxide urgently need to be reduced. Another purpose of the study was to examine the misconceptions that students, in the later school years, have regarding the carbon cycle. This study found misconceptions concerning photosynthesis and respiration in plants, the role of decomposers, as well as getting concepts mixed up with one another. To solve this problem, assignments could be designed where atoms are traced between different levels, in order to put the processes in context. A large part of the focus regarding these misconceptions has been on photosynthesis and respiration and no articles were found regarding e.g. the fast; and the slow; carbon cycle. Few studies of Swedish students have been made and since the Swedish curriculum focuses on the relationship between different levels, the misconceptions might be different in Sweden; what these misconceptions could be in Swedish schools is yet to be investigated.

A food web was constructed for a boreal-forest dominated national park in south-eastern Sweden, called Tyresta national park and its surrounding nature reserve. Due to increasing threats to mammalian species and their critical role in the food web mammals were used as the outset of the food web, and from there branching out to taxa trophically interacting with mammals directly and indirectly. The outset of the food web was constructed using available inventory data, focusing on mammals. From inventory data and 53 interaction sources, consisting of mostly primary literature a food web was created that includes 32 mammalian taxa, six bird families, five invertebrate taxa, fish, amphibians, fungi, reptiles, detritus and 9 different basal plant taxa. The results of the literature review was that the most consumed staple taxon of all were hexapods, with it being a staple food for 15 taxa, six of them being bats preying mainly on dipterans. There are a wide variety of ways that interspecific competition could come to affect taxa in the study area, for example through intraguild predation, increased dietary breadth or avoidance, or a combination of these. This method could serve as a complement to existing food webs or the construction of new ones in a cheaper, less intrusive and less time-consuming way, albeit missing local adaptations. The constructed food web structure and findings of interspecific competition in Tyresta can be used for further analyzing certain links or branching the web out even further, with increased depth or quantification of certain taxon.

A case study of a prehistoric site named Ängesviken, in eastern Jämtland, is presented in this thesis. Ängesviken is situated in a region that traditionally has been understood as peripheral and without a significant history prior to the Late Iron Age or even the mediaeval period. The site appears to have a continuous presence of human activities through a period of 3000 years, manifested through a horizontal stratigraphy. In order to study this site of abstruse character a multidisciplinary approach is used, where archaeology, paleoecology, and spatiality are combined. The conceptual framework of the study is that landscapes are the result of socio-ecological processes over time. To frame the data provided by chosen methods, theoretical frameworks of cultural niche construction theory and landscape patchiness applied, which provide insight of the socio-ecological systems present at Ängesviken during the last 3000 years. During the Iron Age, the site was used for pastoralism, combined with hunting. The archaeological and geographical context of Ängesviken indicate that the site might have been connected with other regions through networks of trade during this period. During the mediaeval period there is a reorganisation of the outlands, leading to a phase of regrowth, but the site could possibly still have been utilised as hunting grounds as there are mediaeval villages in the region. In the early modern period and modern period, the site is once again used for pastoralism. Today, the utilisation of the site has changed towards modern forestry and occasional hunting of elk. The continuous presence at Ängesviken could be explained by the resources the outlands provided. This case study indicates that the far-reaching networks of trade during the Iron Age led to a process of local modification of the ecosystem driven by an external market. This study shows that the anthropogenic modification of the boreal forest's ecosystem has a longer history in eastern Jämtland than traditionally has been thought. The investigation of Ängesviken highlights the importance of researching abstruse and previously uncertain sites from a multidisciplinary approach, as the different datatypes complement each other and results in a deeper knowledge of the site and the socio-ecological systems in a long-term perspective. / Uppsatsen består av en fallstudie av en arkeolgosik lokal vid namn Ängesviken i östra Jämtland. Platsen ligger i ett område som vid första anblick ter sig perifeiellt placerat i utmarkerna till medeltida byar och tidigmoderna fäbodar. Tidigare har en vikingatida byggnad sam en intilligande grav undersökts arkeologiskt. Arkeologiska undersökningar har visat att människor tycks ha använt platsen under en 3000-års period, men hur platsen använts eller påverkats av denna användning har arkeologin inte kunnat påvisa. För att undersöka mänskliga aktiviteter vid Ängesviken under de senaste 3000 åren, samt hur dessa aktiviteter påverkat landskapet, undersöks platsen från ett tvärvetenskapligt perspektiv. Arkeologi kombineras med paleoekologi och rumslig analys. Det konceptuella ramverket för undersökningen bygger på historisk ekologi där kulturell nische konstruktion kombineras med "landscape patchiness". "Landscape patchiness" har sitt ursprung ur ekologin och lägger fokus på lakala vegetationsstrukturer. De äldsta praktikerna på platsen är ännu inte fullt ut klargjorda men platsens läge i landskapet ter sig som en trolig orsak till de första aktiviteterna. Pollenanalysen visar att området används för djurhållning under järnåldern, en aktivitet som kom att förändra landskapets struktur och platsens ekologi. Järnålderns kulturella nische ter sig multifunktionell där djurhållning har kombinerats med andra nyttjanden av utmarksresurser. Pollenanalysen visar att platsen verkar överges under medeltid för att sedan åter brukas för djurhållning under tidigmodern och modern tid. Undersökningen visar på vikten av tvärvetenskapliga undersökningar av otydliga och tidigare svårtolkade arkeologiska lokaler och sammanhang inom det Skandinaviska inlandet.

<p>The energy exchange in the soil-snow-vegetation-atmospheresystem was studied to improve the quantitative knowledge of thegoverning processes. The lack of such knowledge contributes tothe uncertainty in the applicability of many existing modelsindependent of the temporal or spatial scale. The theoreticalbackground and available methods for measurements and numericalsimulations were reviewed. Numerical simulation models andavailable data sets representing open land and boreal forestwere evaluated in both diurnal and seasonal time-scales.Surface heat fluxes, snow depth, soil temperatures andmeteorological conditions were measured at an agriculturalfield in central Sweden over two winters, 1997-1999. Twoone-dimensional simulation models of different complexity wereused to simulate the heat and water transfer in thesoil-snow-atmosphere system and compared with the measurements.Comparison of simulated and observed heat fluxes showed thatparameter values governing the upper boundary condition weremore important than the formulation of the internal mass andheat balance of the snow cover. The models were useful toevaluate the lack of energy balance closure in the observedsurface heat fluxes, which underlined the importance ofimproved accuracy in eddy correlation measurements of latentflow during winter conditions.</p><p>The representation of boreal forest in the land surfacescheme used within a weather forecast model was tested with athree-year data set from the NOPEX forest site in centralSweden. The formulation with separate energy balances forvegetation and the soil/snow beneath tree cover improvedsimulation of the seasonal and diurnal variations of latent andsensible heat flux compared with an older model version.Further improvements of simulated surface heat fluxes could beexpected if the variation of vegetation properties within andbetween years and a new formulation of the boundary conditionsfor heat flux into the soil is included.</p><p><strong>Keywords</strong>: Surface energy balance, Snow, Boreal forest,SVAT models, Eddy-correlation Measurements, Latent heat flux,Sensible heat flux, Net radiation, Soil temperature,Aerodynamic roughness, Surface resistance</p> / QC 20100614

This research investigates soil respiration (Rs) in a boreal jack pine (Pinus banksiana Lamb.) fire scar chronosequence at Sharpsand Creek, Ontario, Canada. During two field campaigns in 2006 and 2007, Rs was measured in a chronosequence of fire scars in the range 0 to 59 years since fire. Mean Rs adjusted for soil temperature (Ts) and soil moisture (Ms) (Rs T,M) ranged from 0.56 μmol CO2/m2/s (32 years post fire) to 8.18 μmol CO2/m2/s (58 years post fire). Coefficient of variation (CV) of Rs adjusted for Ts and Ms ranged from 20% (16 years post fire) to 56% (58 years post fire). Across the field site, there was a significant exponential relationship between Rs adjusted for soil organic carbon (Cs) and Ts (P = 1.24*10-06; Q10 = 2.21) but no effect of Ms on Rs adjusted for Cs and Ts for the range 0.21 to 0.77 volumetric Ms (P = 0.702). Rs T,M significantly (P = 0.030) decreased after burning mature forest, though no significant (P > 0.1) difference could be detected between recently burned and unburned young forest. Rs was measured in recently burned boreal jack pine fire scar age categories that differed in their burn history and there was a significant difference in Rs T,M between previously 32 v 16 year old (P = 0.000) and previously 32 v 59 year old (P = 0.044) scars. There was a strong significant exponential increase in S R T,M with time since fire (r2 = 0.999; P = 0.006) for the chronosequence 0, 16 and 59 years post fire, and for all these age categories, Rs T,M was significantly different from one another (P < 0.05). The Joint UK Land Environment Simulator (JULES) was used to model vegetation re-growth over successional time at Sharpsand Creek, though it appeared to perform poorly in simulating leaf area index and canopy height. JULES probably over estimated heterotrophic Rs at Sharpsand Creek when Ts corrected simulated values were compared with measured Rs T,M. The results of this study contribute to a better quantitative understanding of Rs in boreal jack pine fire scars and will facilitate improvements in C cycle modelling. Further work is needed in quantifying autotrophic and heterotrophic contributions to soil respiration in jack pine systems, monitoring soil respiration for extended time periods after fire and improving the ability of JULES to simulate successional vegetation re-growth.

Afin de mieux comprendre les effets des changements climatiques sur le pergélisol, il s’avère essentiel d’obtenir une meilleure connaissance des facteurs physiques et biologiques l’influençant. Même si plusieurs études font référence à l’influence de la végétation sur le pergélisol à grande échelle, l’effet de la végétation sur la profondeur du front de dégel du pergélisol à l’échelle de mètres, tel qu’exploré ici, est peu connu. L’étude s’est effectuée dans une forêt boréale tourbeuse dans la zone à pergélisol discontinu au sud des Territoires du Nord-Ouest (N61°18’, O121°18’). Nous avons comparé la profondeur de dégel aux mesures du couvert végétal suivantes : densité arborescente, couvert arbustif, indice de surface foliaire et présence de cryptogames (lichens et bryophytes). Nous avons trouvé qu’une plus grande densité arborescente menait à une moins grande profondeur de dégel tandis que le couvert arbustif (<50cm de hauteur) n’avait aucune influence. De plus, la profondeur de dégel dépendait de l’espèce des cryptogames et des microformes. Cette recherche quantifie l’influence de la végétation par strate sur la dégradation du pergélisol. Ultimement, les résultats pourront être pris en considération dans la mise en place des modèles, afin de valider les paramètres concernant la végétation, la dégradation du pergélisol et le flux du carbone. / In order to better understand the impacts of climate change on permafrost degradation, it is important to understand the influence of abiotic and biotic factors on permafrost dynamics. While studies allude to the effect of broad vegetation groups on permafrost dynamics at landscape-scale, the role vegetation plays in affecting the spatial variability of active-layer development on the scale of metres, as explored here, is largely unknown. The study was carried out in a boreal forest-peatland landscape in the discontinuous permafrost zone in the southern Northwest Territories (N61°18’, W121°18’). We examined the influence of the following vegetation characteristics on the spatial variability of thaw depth: tree density, shrub cover, leaf area index, and cryptogam presence (lichen and bryophyte). We found that greater tree density was associated with shallower thaw depths while shrub cover (<50cm height) had a negligible influence. Furthermore, thaw depth depended on the cryptogam species cover and microform. This research quantifies the impact of vegetation by strata on thaw depth and may ultimately serve to refine vegetation parameters in ecosystem models and land surface schemes as part of climate models.

Submitted by Biblioteca de Pós-Graduação em Geoquímica BGQ (bgq@ndc.uff.br) on 2017-09-04T17:31:07Z No. of bitstreams: 1 TESE_Viviane Figueiredo Souza versão final.pdf: 1973892 bytes, checksum: 8b1ac31fbe25c8a6584e68f8e3b8ff71 (MD5) / Made available in DSpace on 2017-09-04T17:31:07Z (GMT). No. of bitstreams: 1 TESE_Viviane Figueiredo Souza versão final.pdf: 1973892 bytes, checksum: 8b1ac31fbe25c8a6584e68f8e3b8ff71 (MD5) / Conselho Nacional de Desenvolvimento Científico e Tecnológico / Universidade Federal Fluminense. Instituto de Química. Programa de Pós-Graduação em Geoquímica, Niterói, RJ / O nitrogênio (N) é um elemento imprescindível para todos os organismos do nosso planeta, entretanto o composto nitrogenado mais abundante, o gás dinitrogênio (N2), é assimilável apenas por poucos micro-organismos. Isso torna o N limitante, refletindo na sua disponibilização via mineralização de matéria orgânica (MO) e a nitrificação, que produzem amônio (NH4+) e nitrato (NO3-), respectivamente. Em solos, esses processos são regulados por fatores como conteúdo de MO e água no solo, pH e temperatura. Em ecossistemas florestais, tropicais e boreais, esses processos são muito relevantes e ainda pouco estudados, principalmente em áreas de floresta secundária sob influência de manejo. A mudança de uso do solo causa alterações na ciclagem e disponibilidade do N, nos fatores reguladores, e na emissão de óxido nitroso (N2O), um gás de efeito estufa. Padrões de recuperação florestal são bastante distintos entre diferentes florestas tropicais, como visto para floresta de Mata Atlântica e Amazônica, com taxas de mineralização em florestas jovens (10 anos) muito elevadas na Amazônia (20,9 μg N g-1 SWD d-1) e mais baixas na Mata Atlântica (3,2 μg N g-1 SWD d-1). Já a nitrificação teve um padrão semelhante, com baixas taxas em ambas as florestas jovens (0,6 6,8 μg N g-1 SWD d-1 na Mata Atlântica e Amazônia, respectivamente), o que indica uma ciclagem de N conservativa, evitando perdas via emissão de gás e lixiviação de NO3-. Entretanto, a emissão de N2O em área de restauração na Mata Atlântica foi maior do que na pristina (22 e 2,5 μg N2O-N g-1 SWD h-1, respectivamente) devido à maior temperatura do solo que estimula os processos microbianos produtores de N2O. Em termos de floresta pristina, a taxa de nitrificação foi bastante distinta entre a floresta de Mata Atlântica e Amazônica (0,08 e 15,9 μg N2O-N g-1 SWD h-1, respectivamente). Isso evidencia os diferentes fatores reguladores de cada região tropical, como regime de chuvas, composição vegetal, tipos de solo, etc. Em florestas boreais, foi visto que o conteúdo de MO e água do solo e o pH do solo são os principais reguladores, limitando a disponibilidade de N em florestas bem drenadas dominadas por coníferas, em comparação com florestas pouco drenadas dominadas por turfa. Dessa forma, verificamos que a mineralização e, principalmente, a nitrificação, são processos muito relevantes no controle de N, e mesmo em ecossistemas diferentes, os fatores reguladores muitas vezes são os mesmos. Isso evidencia a necessidade de mais estudos acerca da dinâmica do N nesses ambientes, principalmente em florestas em restauração / Nitrogen (N) is an essential element for all organisms; however, the most abundant nitrogen compound, dinitrogen (N2), is assimilable only by a few microorganisms. This makes N limiting, which reflects in its availability via organic matter (OM) mineralization and nitrification, which produce ammonium (NH4+) and nitrate (NO3-), respectively. In soils, these processes are regulated by factors such as OM and soil water content, pH and temperature. In forest ecosystems, tropical and boreal, these processes are very relevant and still poorly understood, especially in secondary forest previously deforested. The land use change causes alterations in cycling and availability of N, regulating factors, and emission of nitrous oxide (N2O), a greenhouse gas. Forest recovery patterns are quite distinct among different rainforests, as seen for Atlantic and Amazon rainforest, with very high levels of mineralization in young forests (10 years) in Amazonia (20.9 μg N2O-N g-1 SWD h-1) and low in the Atlantic Forest (3.2 μg N2O-N g-1 SWD h-1). Nitrification had a similar pattern between them, with low rates in both young forests (0.6 and 6.8 μg N2O-N g-1 SWD h-1 in the Atlantic and Amazon forest, respectively), indicating a conservative N cycling, avoiding losses through gas emission and NO3- leaching. However, the N2O emission in restoration area in the Atlantic Forest was higher than in pristine (22 and 2.5 μg N2O-N g-1 SWD h-1, respectively) due to the higher soil temperature, which stimulates microbial production of N2O. In terms of pristine forest, the nitrification rate was very different between the Atlantic forest and Amazonian forest (0.08 and 15.9 μg N2O-N g-1 SWD h-1, respectively), showing the different regulatory factors of each tropical region, such as rainfall regime, vegetal composition, soil types, etc. In boreal forests, OM, soil water content and soil pH were the main regulators, limiting the availability of N in conifer-dominated well-drained forests compared to peat-dominated poorly drained forests. In this way, we verified that mineralization and, mainly, nitrification are very relevant processes in the control of N, and even in extremely different ecosystems, the regulating factors are often the same. This evidences the need for more studies about N dynamics in these environments, especially in restoration forests

<p>The objective of this thesis is to evaluate the ecological consequences of buffer strip retention on riparian and terrestrial biodiversity. Earlier studies on forest buffer strips have evaluated their effectiveness in relation to water quality and aquatic biota. However, forests along streams are species rich habitats for many organism groups. Buffer strip management is assumed to be important also for protecting such species. Current approaches to biodiversity-oriented forest management practices need to be scientifically evaluated. In this thesis the effects on bryophytes and land snails have been evaluated.</p><p>A before-and-after experiment along 15 small streams in northern Sweden showed that buffer strips of 10 m on each side of the stream moderated the negative effects exhibited at the clear-cuts. The number of land snail species remained similar as to before logging and the number of vanished bryophyte species was lower in the buffer strips than in the clear-cuts. The ground moisture influenced the survival rate of land snails at the clear-cuts. At mesic sites many species vanished but at wet sites the snail fauna was unaffected by the logging.</p><p>Many bryophyte species, most of them liverworts, decreased or disappeared in the buffer strips. These were mostly growing on substrates elevated from the forest floor, such as logs, stumps and tree-bases. A number of nationally red-listed species, sensitive for changes in microclimate, were among those decreasing most. Thus, for the species in most need of protection the buffer strips were too narrow.</p><p>An experiment with bryophyte transplants followed over a season showed that wet ground moisture moderated the negative edge effects in narrow buffer strips. On the other hand, the growth in mesic and moist sites was almost as low as in comparable clear-cuts.</p><p>Microclimatic edge effects are stronger at south facing than north-facing edges of forest clear-cuts. This was shown in an experiment using bryophyte growth as an indicator of differences in microclimate. However, the depth of edge influence seemed to be similar between north- and south-facing forest edges, >30 m for one species. An explanation for this could be that wind penetrates deeper into edges than solar radiation and has a more variable direction.</p><p>In conclusion, narrow buffer strips consist entirely of edge habitat. For many species the environment in buffer strips is good enough for persistence. For others, most notably bryophyte species on convex substrates, wider buffer strips are needed to ensure long-term survival. </p>